Chapter 20 Phylogenetic Trees Figure 20.5 Branch point: where lineages diverge Taxon A 3 Taxon B 4 Taxon C Sister taxa 2 Taxon D ANCESTRAL LINEAGE 1 5 Taxon E Taxon F Taxon G This branch point represents the common ancestor of taxa A−G. This branch point forms a polytomy: an unresolved pattern of divergence. Basal taxon • Which animal represents the basal taxon? – Camels • Which animal is least like the others? – Camels • Are pigs and peccaries sister taxa? – Yes, they share an immediate common ancestor. • Did baleen whales evolve from toothed whales? – No! Do not assume that a taxon evolved from the taxon next to it. • Which are more closely related, hippos and toothed whales or hippos and pecorans? – Hippos and toothed whales • Which are more closely related, hippos and toothed whales or hippos and baleen whales? – You can’t tell from the information given Leopard (common name) What level of classification do humans share with leopards? Species: Panthera pardus Genus: Panthera Family: Felidae Order: Carnivora Domain Kingdom Phylum Class Class: Mammalia Phylum: Chordata Domain: Bacteria Kingdom: Animalia Domain: Eukarya Domain: Archaea Leopard (common name) What genus is the leopard in? Species: Panthera pardus Panthera Domain: Bacteria Kingdom: Animalia Domain: Eukarya Domain: Archaea Concept 20.2: Phylogenies are inferred from morphological and molecular data • The similarities used to infer phylogenies must result from shared ancestry. • Bat and bird wings are homologous as forelimbs, but analogous as functional wings. Evaluating Molecular Homologies • Analogous structures or molecular sequences are also called homoplasies. • Molecular homologies are determined based on the degree of similarity in nucleotide sequence between taxa. • What happens in a frameshift mutation? • Systematists use computer programs when analyzing comparable DNA segments from different organisms. Figure 20.8-4 1 CCAT CAGA GTCC 2 CCAT CAGA GTCC Deletion 1 CCAT CAGA GTCC 2 CCAT CAGA GTCC G T A Insertion 1 CCAT CAAGTCC 2 CCAT GTA CAG AGTCC 1 CCAT CA AGTCC 2 CCAT GTA CAG AGTCC • Shared bases in nucleotide sequences that are otherwise very dissimilar are called molecular homoplasies. A C GG A T A G T C C A C T A G G C A C T A T C A C C GA C A G G T C T T T G A C T A G • About how much DNA would you expect two species to share based on chance alone?