Plants General Biology II Lecture Land Plants – monophyletic group, “embryophytes” o o o Nonvascular Hepatophyta – liverworts Anthocerophyta – hornworts Bryophyte – mosses Vascular, seedless Lycopodiophyta – club mosses Monilophyta – horsetails, whisk ferns, ferns Vascular, seed o Gymnosperms Cycadophyta Ginkgophyta Gnetophyta Coniferophyta Angiosperms Monocot Eudicot Know facts about each group, look at evolutionary history Life Cycle o Alternation of Generations In nonvascular plants, gametophyte generation is dominant; in vascular plants, sporophyte generation is dominant o o Nonvascular plants Flagellated sperm rely on water source Female gametophyte – archegonium, produces egg Male gametophyte – antheridium, produces many sperm Vascular Plants Megaspores (produced in SMALL numbers in megasporangia) develop into female megagametophyte Micropores (produced in LARGE numbers in microsporangia) develop into microgametophyte (pollen grain) Seed Plants Seed – well protected resting stage o Tissues from 3 generations Seed coat and megasporangium from sporophyte parent Haploid female gametophyte – nutrients Embryo – new diploid sporophyte Gymnosperms “naked seeds” o Megastrobilus – female seed-bearing cone; modified stem o Microstrobilus – male, pollen bearing cone; modified leaf o Fertilization Pollen distributed by wind 2 sperm travel through pollen tube – one fertilizes, other degenerates Angiosperms o Double Fertilization Microgametophyte has 2 male gametes o 1 fuses with egg 1 fuses with 2 haploid nuclei of female to make triploid endosperm Megagametophyte is only 7 cells Can be monoeicius or dioecius o Most dioecius plants have methods that prevent self-fertilization Flower Structure Pistil (can be one carpel or several carpels fused together), bear megasporangium o Stigma Style Ovary – develops into fruit Ovule – develops into seed Stamen, bear microsporangia Anther Filament Flowers can be perfect or imperfect 3 patterns of flowering: Annuals – complete life cycle in 1 year and have little or no secondary growth Biennials – take 2 years, store carbs underground during 1st year and then produce flowers 2nd year o Perennials – live 3 or more years Fruits – develop from ovary of carpel Simple – develop from one carpel, cherries Aggregate – develop from several carpels on same flower, raspberries Multiple – develop from cluster of flowers, pinapple Accessory – fruits develop from parts other than carpels Plant Body of Angiosperms o o o o 3 vegetative organs organized into root and shoot systems: Stems Roots Leaves Root Taproot Fibrous roots (most of the monocots) 3 Zones: Zone of maturation, zone of elongation, zone of cell division; root cap Monocot vs. eudicot root Stems Support leaves and flowers Have buds Axillary buds – can form new branches Terminal buds – at tips, can form new branches upward, can form flowers Can be modified – potatoes, runners on strawberries, barrel on cactus to store water Monocot vs. eudicot stem Leaves Consists of blade and petiole that attaches it to stem Can be modified – cacti spines, onion o Root and shoot systems consist of apical meristems at tips (clusters of undifferentiated cells) o 3 tissue types 1. Dermal Forms the epidermis 2. Ground – makes up most of the plant body 3 cell types: o Palisade mesophyll Spongy mesophyll o Collenchyma – provides support o Sclerenchyma – fibers and schlereids, rigid support 3. Vascular Xylem – transports water and nutrients from roots o Cells called tracheary elements (die before assuming role) o Water moves through xylem by transpiration-tension-cohesion mechanism Phloem – transports products of photosynthesis o o Parenchyma – sites for photosynthesis and storage Cells called sieve tube elements, still living Primary vs Secondary Growth Primary – extension of root and shoots Apical meristems All monocots, some eudicots Secondary - increases diameter, production of wood Eudicots Lateral meristems o Vascular cambium – provides secondary xylem toward inside of stem and phloem toward outside of stem o Cork cambium Cork, cork cambium, and phelloderm make up the periderm Periderm + Secondary Phloem (everything external to vascular cambium) becomes bark Hormones Plant development dependent on hormone regulation Gibberellins – stem elongation Auxins – phototropism and gravitropism; inhibit abscission (detachment of leaf from stem) Effects of ethylene gas