Lycopytes and Sphenophytes ● Lycophytes – club mosses ● Sphenophytes – horsetails ● Appeared more than 400 million years ago. ● Most ancient of the vascular plants. Ancient Forests ● The Earth’s first forests were composed of these plants. ● The fossilized remains of these plants were transformed into coal beds. Ferns ● More successful and more complex than the club mosses and horsetails. ● Abundant in wet or seasonally wet habitats throughout the world. ● Grow best in wet tropical areas. Structures ● True leaves, roots and stems. ● Rhizomes: creeping, underground stems. ● Fronds: large leaves. Reproductive Structures ● Sporangia: structures found under the fronds which produce spores. ● Sori (sing: sorus): clusters of sporangia. ● Prothallium: green, heart shaped gametophyte. ● Antheridium: produces sperm. Grows on prothallium. ● Archegonium: produces eggs. Grows on prothallium. Life Cycle ● Alternation of generations. ● Sporophyte is more obvious, dominant stage. ● Spores are produced in sporangia. ● Spores are dispersed by the wind and grow into the prothallium (gametophyte). ● The gametophyte is small and lives for a short time. It lacks vascular tissue and only grows in moist areas. ● Archegonia (egg production) and antheridia (sperm production) grow on the gametophyte. ● Fertilization occurs when flagellated sperm SWIM to the eggs. ● Diploid sporophyte grows on gameophtye. ** Water is still required for reproction.