Chapter 2 Neuromuscular Fundamentals 2-1 Manual of Structural Kinesiology Skeletal Muscles Responsible for movement of body and all of its joints Muscle contraction produces force that causes joint movement Muscles also provide protection posture and support produce a major portion of total body heat 2-2 Muscle Nomenclature Muscles are usually named due to visual appearance anatomical location function Shape – deltoid, rhomboid Size – gluteus maximus, teres minor Number of divisions – triceps brachii Direction of its fibers – external abdominal oblique 2-3 Shape of Muscles & Fiber Arrangement Cross section diameter factor in muscle’s ability to exert force greater cross section diameter = greater force exertion Muscle’s ability to shorten longer muscles can shorten through a greater range more effective in moving joints through large ranges of motion Neuromuscular Fundamentals 2-4 Manual of Structural Kinesiology Muscle Terminology Origin Structurally, the proximal attachment of a muscle or the part that attaches closest to the midline or center of the body Functionally & historically, the least movable part or attachment of the muscle Insertion Structurally, the distal attachment or the part that attaches farthest from the midline or center of the body Functionally & historically, the most movable part is generally considered the insertion 2-5 Types of muscle contraction Muscle contractions can be used to cause, control, or prevent joint movement or to initiate or accelerate movement of a body segment to slow down or decelerate movement of a body segment to prevent movement of a body segment by external forces All muscle contractions are either isometric or isotonic Neuromuscular Fundamentals 2-6 Manual of Structural Kinesiology Types of muscle contraction Muscle Contraction (under tension) Isometric Isotonic Preventing Motion Concentric Causing Neuromuscular Fundamentals Motion 2-7 Eccentric Controlling Motion Manual of Structural Kinesiology Types of muscle contraction Isometric contraction active tension is developed within muscle but joint angles remain constant static contractions significant amount of tension may be developed in muscle to maintain joint angle in relatively static or stable position may be used to prevent a body segment from being moved by external forces preventing motion Neuromuscular Fundamentals 2-8 Manual of Structural Kinesiology Types of muscle contraction Concentric contractions involve muscle developing active tension as it shortens Causing motion Eccentric contractions involve the muscle lengthening under active tension Controlling motion 2-9 Role of Muscles Agonist muscles cause joint motion through a specified plane of motion when contracting concentrically known as primary or prime movers Some agonist muscles, because of their relative location, size, length, or force generation capacity, are able to contribute significantly more to the joint movement than other agonists Assisters or assistant movers Agonist muscles that contribute significantly less to the joint motion 2-10 Role of Muscles Antagonist muscles located on opposite side of a joint from the agonist have the opposite concentric action known as contralateral muscles work in cooperation with agonist muscles by relaxing & allowing movement when contracting concentrically perform the opposite joint motion of agonist Ex. quadriceps muscles are antagonists to hamstrings in knee flexion 2-11 Role of Muscles Stabilizers surround joint or body part contract to fixate or stabilize the area to enable another limb or body segment to exert force & move known as fixators essential in establishing a relatively firm base for the more distal joints to work from when carrying out movements Ex. biceps curl muscles of scapula & glenohumeral joint must contract in order to maintain shoulder complex & humerus in a relatively static position so that the biceps brachii can more effectively perform curls Proximal stability is needed for distal mobility. 2-12 Role of Muscles Synergist assist in action of agonists not necessarily prime movers for the action known as guiding muscles assist in refined movement & rule out undesired motions 2-13 Role of Muscles Force Couples Force couples occur when two or more forces are pulling in different directions on an object, causing the object to rotate about its axis Coupling of muscular forces together in the body can result in a more efficient movement 2-14 Lines of Pull Consider the following 1. Exact locations of bony landmarks to which muscles attach proximally & distally and their relationship to joints 2. Planes of motion through which a joint is capable of moving 3. Muscle’s relationship or line of pull relative to the joint’s axes of rotation 4. As a joint moves the line of pull may change & result in muscle having a different or opposite action than in the original position 5. Effect of a muscle’s relative length on its ability to generate force 2-15 Factors affecting muscle tension development All or None Principle - regardless of number, individual muscle fibers within a given motor unit will either fire & contract maximally or not at all Difference between lifting a minimal vs. maximal resistance is the number of muscle fibers recruited The number of muscle fibers recruited may be increased by activating those motor units containing a greater number of muscle fibers activating more motor units increasing the frequency of motor unit activation 2-16 Motor Control A motor neuron and the muscle fibers it innervates. Fine control small motor units contain as few as 20 muscle fibers per nerve fiber eye muscles Allow for greater dexterity because of a lower neuron to muscle fiber ratio. Gross control gastrocnemius muscle has 1000 fibers per nerve fiber One motor neuron controls many muscle fibers which allows for strength production. Recruitment and Stimulus Intensity Strength of muscle contraction is dependant of # of motor units recruited Multiple motor unit summation The harder the activity is the more motor units will be recruited. Lifting 1 lb vs. 100 lbs How do you explain the rapid strength gains when you first start training? All or None Principle Typical muscle contraction The number of motor units responding (and number of muscle fibers contracting) within the muscle may vary significantly from relatively few to virtually all depending on the number of muscle fibers within each activated motor unit & the number of motor units activated 2-19 Metabolism and Skeletal Muscle Fibers Types There are 3 different types skeletal muscle fibers based histological differences, duration of a twitch and the method of ATP production slow oxidative fibers fast oxidative fibers fast glycolytic fibers Proportions genetically determined Fast Glycolytic, Fast-Twitch Fibers Fast glycolytic, fast-twitch fibers: rich in enzymes for phosphagen and glycogen-lactic acid systems Limited # of mitochondria and high concentration of glycogen stores makes it adapted for anaerobic metabolism a lack of myoglobin in glycolytic fibers results in a white color sarcoplasmic reticulum releases calcium quickly so contractions are quicker which are required for movements that produce speed and power. extraocular eye muscles, gastrocnemius and biceps brachii Slow- Twitch Fibers Slow oxidative, slow-twitch fibers Oxidative fibers contain greater amounts of mitochondria and myoglobin which binds oxygen. Rich blood supply and high concentration of myoglobin these fibers appear red in color. adapted for endurance (resistant to fatigue) Soleus and postural muscles of the back are predominantly this type. Fast Oxidative Fibers Fast oxidative fibers: characteristics of both fast and slow fibers. have a fast twitch (use ATP quickly) Increased mitochondria make it moderately resistant to fatigue Usually make up 10% of fibers. Training will make these fibers adapt to become functionally more fast or slow. Cellular Adaptations to Physical Demands Strength training: high intensity training stresses anaerobic pathways. Increased # and size of glycolytic associated enzymes and substrates ATP, creatine phosphate and glycogen Endurance training: Enhance the aerobic pathways. increased # and size of mitochondrial membranes and associated enzymes. This will increase O2 uptake ( VO2 the formation of lactic acid. max) which will delay Factors affecting muscle tension development Number of muscle fibers per motor unit varies significantly From less than 10 in muscles requiring precise and detailed such as muscles of the eye To as many as a few thousand in large muscles that perform less complex activities such as the quadriceps Neuromuscular Fundamentals 2-25 Manual of Structural Kinesiology Muscle Length - Tension Relationship Generally, depending upon muscle involved Greatest amount of tension can be developed when a muscle is stretched between 100% to 130% of its resting length Stretch beyond 100% to 130% of resting length significantly decreases the amount of force production A proportional decrease in ability to develop tension occurs as a muscle is shortened When shortened to around 50% to 60% of resting length ability to develop contractile tension is essentially reduced to zero Muscle Length - Tension Relationship Maximal ability of a muscle to develop tension & exert force varies depending upon the length of the muscle during contraction Neuromuscular Fundamentals 2-27 Manual of Structural Kinesiology Muscle Length - Tension Relationship Ex. 1 Increasing ability to exert force squat slightly to stretch the gastroc, hamstrings, & quadriceps before contracting same muscles concentrically to jump Ex. 2. Reducing ability to exert force isolate the gluteus maximus by maximally shortening the hamstrings with knee flexion Neuromuscular Fundamentals 2-28 Manual of Structural Kinesiology Reciprocal Inhibition or Innervation Antagonist muscles groups must relax & lengthen when the agonist muscle group contracts This reciprocal innervation effect occurs through reciprocal inhibition of the antagonists Activation of the motor units of the agonists causes a reciprocal neural inhibition of the motor units of the antagonists This reduction in neural activity of the antagonists allows them to subsequently lengthen under less tension Biceps contracts triceps must relax Active & Passive Insufficiency As muscle shortens its ability to exert force diminishes Active insufficiency is reached when the muscle becomes shortened to the point that it can not generate or maintain active tension Passively insufficiency is reached when the opposing muscle becomes stretched to the point where it can no longer lengthen & allow movement Active Insufficiency When a multi-joint muscle contracts simultaneously over 2 joints the muscle is unable to generate a maximum muscle contraction. Ex. Rectus femorus contracts concentrically to both flex the hip & extend the knee Hamstrings: contracts both knee flexion and hip extension Gastrocnemius: Knee flexion and plantar flexion Biceps brachii: shoulder flexion and elbow flexion 2-31 Manual of Structural Kinesiology Passive Insufficiency Agonist muscle unable to reach full range of motion because of inadequate length of antagonistic two joint muscle. Hamstrings can’t stretch enough to allow both maximal hip flexion & maximal knee extension. Knee flexion cannot be fully achieved if the hip is fully extended. Rectus femoris length limits motion. Finger flexion is limited if wrist flexion occurs simultaneously. Proprioception Subconscious mechanism by which body is able to regulate posture & movement by responding to stimuli originating in proprioceptors of the joints, tendons, muscles, & inner ear Proprioceptors - internal receptors located in skin, joints, muscles, & tendons which provide feedback relative to: tension, length, & contraction state of muscle. position of body & limbs. movements of joints. Neuromuscular Fundamentals 2-33 Manual of Structural Kinesiology Proprioception Proprioceptors specific to muscles Muscles spindles concentrated primarily in muscle belly between the fibers sensitive to stretch & rate of stretch Insert into connective tissue within muscle & run parallel with muscle fibers Golgi tendon organs (GTO) found serially in the tendon close to muscle tendon junction sensitive to both muscle tension & active contraction much less sensitive to stretch than muscle spindles require a greater stretch to be activated Muscle spindle Muscle spindles & myotatic or stretch reflex 1. Rapid muscle stretch occurs 2. Afferent Impulse is sent to the CNS 3. CNS activates motor neurons of agonist muscle while inhibiting the antagonist muscle. 4. Stretched or agonistic muscle contracts Monosynaptic Reflex Ex. Knee jerk or patella tendon reflex Neuromuscular Fundamentals 2-36 Manual of Structural Kinesiology Golgi Tendon Organ Golgi tendon organ Tension in tendons & GTO increases as muscle contract, which activates GTO 1. 2. 3. 4. GTO stretch threshold is reached Impulse is sent to CNS CNS causes muscle to relax facilitates activation of antagonists as a protective mechanism GTO protects us from an excessive contraction by causing its muscle to relax Manual of Structural Kinesiology Chapter 3 Basic Biomechanical Factors & Concepts Basic Biomechanical Factors & Concepts 3-38 Manual of Structural Kinesiology Biomechanics Biomechanics - study of the mechanics as it relates to the functional and anatomical analysis of biological systems Necessary to study the body’s mechanical characteristics & principles to understand its movements Mechanics - study of physical actions of forces Musculoskeletal system may be thought of as a series of simple machines Machines - used to increase mechanical advantage Consider mechanical aspect of each component in analysis with respect to components’ machine-like function Levers Humans moves through a system of levers Levers cannot be changed, but they can be utilized more efficiently lever - a rigid bar that turns about an axis of rotation or a fulcrum axis - point of rotation about which lever moves Levers rotate about an axis as a result of force (effort, E) being applied to cause its movement against a resistance or weight In the body bones represent the bars joints are the axes muscles contract to apply force Manual of Structural Kinesiology Levers Three points determine type of lever & for which kind of motion it is best suited Axis (A)- fulcrum - the point of rotation Point (F) of force application (usually muscle insertion) - effort Point (R) of resistance application (center of gravity of lever) or (location of an external resistance) Basic Biomechanical Factors & Concepts 3-41 Manual of Structural Kinesiology Levers Systems 1st class lever – axis (A) somewhere between force (F) & resistance (R) 2nd class lever – resistance (R) somewhere between axis (A) & force (F) 3rd class lever – force (F) somewhere between axis (A) & resistance (R) Manual of Structural Kinesiology Levers • FAR 1st Force Arm | || Resistance Arm F | R A • ARF 2nd | Resistance Arm | | Force Arm R | F A • AFR 3rd | | Force Arm | Resistance Arm F A Basic Biomechanical Factors & Concepts 3-43 | R Manual of Structural Kinesiology First-class Levers Produce balanced movements when axis is midway between force & resistance (e.g., seesaw or scissors) Produce speed & range of motion when axis is close to force, (triceps in elbow extension) Produce force motion when axis is close to resistance (crowbar) Manual of Structural Kinesiology First-class Levers Head balanced on neck in flexing/extending Elbow extension in triceps applying force to olecranon (F) in extending the nonsupported forearm (R) at the elbow (A) Agonist & antagonist muscle groups are contracting simultaneously on either side of a joint axis agonist produces force while antagonist supplies resistance First-class Levers Force is applied where muscle inserts in bone, not in belly of muscle Ex. in elbow extension with shoulder fully flexed & arm beside the ear, the triceps applies force to the olecranon of ulna behind the axis of elbow joint As the applied force exceeds the amount of forearm resistance, the elbow extends 3-46 Manual of Structural Kinesiology Levers The mechanical advantage of levers may be determined using the following equations: Mechanical advantage = Resistance Force or Mechanical advantage = Length of force arm Length of resistance arm Torque and length of lever arms First class levers A, If the force arm & resistance arm are equal in length, a force equal to the resistance is required to balance it; B, As the force arm becomes longer, a decreasing amount of force is required to move a relatively larger resistance; C, As the force arm becomes shorter, an increasing amount of force is required to move a relatively smaller resistance Second-class Levers Produces force movements, since a large resistance can be moved by a relatively small force Wheelbarrow Nutcracker Loosening a lug nut Raising the body up on the toes Second-class Levers Plantar flexion of foot to raise the body up on the toes where ball (A) of the foot serves as the axis as ankle plantar flexors apply force to the calcaneus (F) to lift the resistance of the body at the tibial articulation (R) with the foot Relatively few 2nd class levers in body Torque and length of lever arms Second class levers A, Placing the resistance halfway between the axis & the point of force application provides a MA of 2; B, Moving the resistance closer to the axis increases the MA, but decreases the distance that the resistance is moved; C, the closer the resistance is positioned to the point of force application the less of a MA, but the greater the distance it is moved Third-class Levers Produce speed & range-of-motion movements Most common in human body Requires a great deal of force to move even a small resistance Paddling a boat Shoveling - application of lifting force to a shovel handle with lower hand while upper hand on shovel handle serves as axis of rotation Basic Biomechanical Factors & Concepts 3-52 Manual of Structural Kinesiology Third-class Levers Biceps brachii in elbow flexion Using the elbow joint (A) as the axis, the biceps brachii applies force at its insertion on radial tuberosity (F) to rotate forearm up, with its center of gravity (R) serving as the point of resistance application Torque and length of lever arms Third class levers A, a force greater than the resistance, regardless of the point of force application, is required due to the resistance arm always being longer; B, Moving the point of force application closer to the axis increases the range of motion & speed; C, Moving the point of force application closer to the resistance decreases the force needed Torque and length of lever arms Long levers produce more linear force and thus better performance in some sports such as baseball, hockey, golf, field hockey, etc. For quickness, it is desirable to have a short lever arm 3-55 Pulleys Single pulleys function to change effective direction of force application Mechanical advantage = 1 Pulleys may be combined to form compound pulleys to increase mechanical advantage Each additional rope increases mechanical advantage by 1 Pulleys Ex. lateral malleolus acting as a pulley around which tendon of peroneus longus runs As peroneus longus contracts, it pulls toward it belly (toward the knee) Using the lateral malleolus as a pulley, force is transmitted to plantar aspect of foot resulting in eversion/plantarflexion Force Muscles are the main source of force that produces or changes movement of a body segment, the entire body, or some object thrown, struck, or stopped Strong muscles are able to produce more force than weak muscles both maximum and sustained exertion over a period of time Basic Biomechanical Factors & Concepts 3-58 Manual of Structural Kinesiology Force Forces either push or pull on an object in an attempt to affect motion or shape Without forces acting on an object there would be no motion Force - product of mass times acceleration Mass - amount of matter in a body Basic Biomechanical Factors & Concepts 3-59 Manual of Structural Kinesiology Force The weight of a body segment or the entire body X the speed of acceleration determines the force Important in football Also important in activities using only a part of the body In throwing a ball, the force applied to the ball is equal to the weight of the arm times the speed of acceleration of the arm Leverage factors are also important Basic Biomechanical Factors & Concepts 3-60 Manual of Structural Kinesiology Mechanical loading basics Significant mechanical loads are generated & absorbed by the tissues of the body Internal or external forces may cause these loads Only muscles can actively generate internal force, but tension in tendons, connective tissues, ligaments and joints capsules may generate passive internal forces Basic Biomechanical Factors & Concepts 3-61 Manual of Structural Kinesiology Web Sites Biomechanics: The Magazine of Body Movement and Medicine www.biomech.com/ Biomechanics World Wide www.uni-essen.de/~qpd800/WSITECOPY.html This site enables the reader to search the biomechanics journals for recent information regarding mechanism of injury. Kinesiology Biomechanics Classes www.uoregon.edu/~karduna/biomechanics/kinesiology.htm A listing of numerous biomechanics and kinesiology class site on the web with many downloadable presentations and notes The Physics Classroom www.physicsclassroom.com/ Numerous topics including the laws of motion and other physics principles Basic Biomechanical Factors & Concepts 3-62 Manual of Structural Kinesiology Web Sites Edquest www.edquest.ca/pdf/sia84notes.pdf Text, pictures, and illustrations on simple and complex machines COSI Hands-on science centers www.cosi.org/files/Flash/simpMach/sm1.swf A Flash site demonstrating simple machine explanations EuclideanSpace - building a 3D world www.euclideanspace.com Information on how to simulate physical objects with computer programs Physics Homework Help http://tutor4physics.com/index.htm Physics formulas, principles, tutorials Basic Biomechanical Factors & Concepts 3-63 Manual of Structural Kinesiology Web Sites GRD Training Corporation www.physchem.co.za Explanations of physics principles for in motion with quizzes International Society of Biomechanics www.isbweb.org/ Software, data, information, resources, yellow pages, conferences Sports Coach—Levers www.brianmac.demon.co.uk/levers.htm A basic review of levers with excellent links to the study of muscle training & function. Integrated Publishing www.tpub.com/content/engine/14037/index.htm Engine mechanics Basic Biomechanical Factors & Concepts 3-64 Manual of Structural Kinesiology Web Sites Human Anatomy Online http://innerbody.com/image/musfov.html An interactive site with details on the muscular and nervous systems. BBC Science & Nature http://bbc.co.uk/science/humanbody/body/interactives/3djigs aw_02/index.shtml An interactive site allowing you to select the innervation for the muscles. Functions of the Muscular System http://training.seer.cancer.gov/anatomy/muscular Several pages with information on skeletal muscle structure, types, and groups, along with unit review and quizzes. Neuromuscular Fundamentals 2-65 Manual of Structural Kinesiology Web Sites U.S. National Library of Medicine’s Visible Human Project AnatQuest Anatomic Images Online http://anatline.nlm.nih.gov/AnatQuest/AwtCsViewer/aq-cutaway.html A great resource using a cut-away viewer to see cadaver images throughout the body to identify the relevant anatomy. The Physician and Sportsmedicine www.physsportsmed.com/index.php?article=1476 Refining rehabilitation with proprioception training: expediting return to play. Get Body Smart www.getbodysmart.com/index.htm An online interactive tutorial on the muscular and nervous systems. Neuromuscular Fundamentals 2-66 Manual of Structural Kinesiology Web Sites Neurologic Exam: An anatomical approach http://medlib.med.utah.edu/neurologicexam/home_exam.html A very thorough site on neurological exams, including numerous movies with both normal and pathological results. Cranial Nerves: Review info www.gwc.maricopa.edu/class/bio201/cn/cranial.htm A good resource on the cranial nerves. University of Arkansas for Medical Sciences Nerve tables http://anatomy.uams.edu/anatomyhtml/nerves.html Numerous tables of all nerves throughout the body. Dermatomes www.meddean.luc.edu/lumen/MedEd/GrossAnatomy/learnem/dermat/m ain_der.htm An interactive review of the body’s dermatomes. Neuromuscular Fundamentals 2-67 Manual of Structural Kinesiology Web Sites Loyola University Medical Education Network Master Muscle List www.meddean.luc.edu/lumen/MedEd/GrossAnatomy/dissector/mml An interactive and graphical review of the muscles indexed alphabetically and by region. Proprioception Exercises Can Improve Balance http://sportsmedicine.about.com/library/weekly/aa062200.htm Proprioception. Muscular System www.bio.psu.edu/faculty/strauss/anatomy/musc/muscular.htm Cadavaric photos of the cat muscular system. Functions of the Nervous System http://training.seer.cancer.gov/anatomy/nervous Several pages with information on the nervous system organization, nerve structure, unit review, and quizzes. Neuromuscular Fundamentals 2-68 Manual of Structural Kinesiology Web Sites Training for Proprioception & Function www.coachr.org/proprio.htm Information on improving body awareness movement efficiency. Neuromuscular Fundamentals 2-69 Manual of Structural Kinesiology