47.2 Cerastium L. (1753), Sp. Pl. 437. [VP]
Comments:
(1)
The order of species has been changed from Petrovsky's draft to keep together the
obviously closely related taxa of the C. alpinum group. (Elven)
(2)
Petrovsky proposes a subgeneric classification into two subgenera and two sections of
the latter subgenus:
I Subg. Dichodon (Bartl.) Fenzl in Ledeb. (1842), Fl. Ross. 1: 496; basionym:
Dichodon Bartl. in Rchb. (1841), Nomencl. 205. [C. cerastoides].
II Subg. Eucerastium (Boiss.) Pax in Engl. & Prantl (1884), Nat. Pflanzenfam. 2,1:
80; basionym: sect. Eucerastium Boiss. (1867), Fl. Or. 1: 713.
a Sect. Strephodon Ser. in DC. (1824), Prodr. 1: 414. [C. maximum and C.
dahuricum].
b Sect. Orthodon Ser. in DC. (1824), Prodr. 1: 415. [all the others].
There are two reasons for omitting this classification: (a) Cerastium
maximum and C. dahuricum have also been descibed as Dichodon; and (b) There is
documented hybridisation between Dichodon and Eucerastium with fertile polyploid
offspring, which blurs the limit. Cerastium cerastoides forms a yet unnamed fertile
allopolyploid with C. alpinum locally in S Norway. The formalistic characters used
for this separation obviously have a fairly simple genetic background. I have
therefore omitted the classificaton. (Elven)
(3)
Excluded species: Cerastium glomeratum Thuill. (1799), Fl. Env. Paris, éd. 2, 226;
synonym: C. viscosum L. (1753), Sp. Pl. 437, p.p. The single report in Petrovsky's
draft, from Jan Mayen, is erroneous. Obviously casual in N Alaska (Brooks Range)
where it is mapped by Hultén (1968). This species must therefore be excluded from
the checklist if there are no other records. (Elven)
47.2.1 Cerastium cerastoides (L.) Britton (1894), Mem. Torrey Bot. Club 5: 150.
B
Stellaria cerastoides L. (1753), Sp. Pl. 422.
S
Dichodon cerastoides (L.) Rchb. (1841), Icon. Fl. Germ. Helv. 5: f. 4915; C.
trigynum Vill. (1779), Hist. Pl. Dauphiné 48.
T
Linnaean herbarium 584.8 (LINN) lectotype, selected by Jonsell & Jarvis (1994,
Nord. J. Bot. 14: 159).
2n=
38 (2x).
2nD
Löve & Löve (1975) list numerous counts, some as arctic.
G
ICE NOR RUS CAN GRL
Comments:
47.2.2
S
T
2n=
2nD
Cerastium maximum L. (1753), Sp. Pl. 439.
Dichodon maximum (L.) Á. Löve & D. Löve (1976), Bot. Not. 128: 507.
[Described from Siberia.]
38 (2x).
Löve & Löve (1975) list numerous counts, some as arctic from Russia/Siberia and
North America; Petrovsky & Zhukova (1981 Wrangel Isl).
G
RUS SIB RFE ALA CAN
Comments:
(1)
If the subgeneric classification is retained, there must be some comment on the
placement of this section and species, in Dichodon or Eucerastium, see Löve & Löve
(1976: 507). I have added the Dichodon synonym just to remind you of this. (Elven)
47.2.3 Cerastium dahuricum Fisch. ex Spreng. (1815), Pugillus 2: 65.
S
Dichodon dahuricum (Fisch. ex Spreng.) Á. Löve & D. Löve (1976), Bot. Not. 128:
507.
2n=
38 (2x).
2nD
Löve & Löve (1975) list five counts, one as arctic.
G
RUS SIB
Comments:
(1)
Added to Petrovsky's draft. Mapped into arctic parts of Polar Ural by Jalas &
Suominen (1983) and perhaps also in Siberia along Jenisei (Hultén & Fries 1986).
This is also a Dichodon taxon in Löve's concept. (Elven)
(2)
A border case in Siberia. (Petrovsky)
47.2.4 Cerastium arvense L. (1753), Sp. Pl. 438.
S
C. arvense L. var. taimyrense Tolm. (1971), Fl. Arct. URSS 6: 49; C. strictum L.
(1753), nom. dub., Sp. Pl. 439.
T
Sweden: Skåne, Linnaean herbarium 603.9 (LINN) lectotype, selected by Ugborogho
(1977, Phyton (Buenos Aires) 35: 177).
2n=
(1) 36 (2x). (2) 72 (4x).
2nD
(1) Löve & Löve (1975) list numerous counts, one as arctic (Zhukova & Petrovsky
1971 Wrangel I.); Zhukova & Petrovsky (1981 Wrangel I.). (2) Löve & Löve (1975)
list numerous non-arctic counts.
G
RUS SIB RFE CAN GRL
Comments:
(1)
An extremely heterogeneous species. Is there any information about subspecies (etc.)
present in the Arctic?
Löve & Löve (1975) treated the ploidy levels as two species, C. arvense s.
str. (2n=72) and C. strictum (L.) Haenke (2n=36). There is a C. arvense L. var.
strictum Koch (1835), Syn. Fl. Germ. Helv. 124, synonym: C. arvense L. subsp.
strictum (Koch) Schinz & R. Keller (1905), Fl. Schweiz, ed. 2, 1: 182, but I believe
that Löve's name rather is based on the dubious Linnaean taxon included in Species
Plantarum, not on the Koch basionym. Flora Europaea reports their (Koch's) strictum
with 2n=36, arvense s. str. with 2n=72. In any case, this strictum taxon is a southern,
thermophilous plant. (Elven)
(2)
The Tolmachev synonym is added as it is an accepted name in Fl. Arct. URSS.
(Elven)
The Cerastium alpinum L. aggregate (C. alpinum, C. arcticum, C. beeringianum, C.
fischerianum, C. nigrescens, C. regelii)
Comments:
(1)
Most or all of the entities currently recognized as species in this aggregate are fairly
distinct. However, the frequent use of 'C. alpinum' in a very collective meaning in
American and Greenlandic literature may make this aggregate necessary for some
areas. (Elven)
47.2.5 Cerastium alpinum L. (1753), Sp. Pl. 438.
S
T
Lapland herbarium 192 (LAPP) lectotype, selected by Jonsell & Jarvis (1994, Nord.
J. Bot. 14: 156).
2n=
For the collective species. (1) 54 (3x) see comment. (2) 72 (4x).
2nD
(1) Sokolovskaya & Strelkova (1960 Kola Pen., referred by Löve & Löve 1975 as C.
hyperboreum). (2) Löve & Löve (1975) list numerous counts for the collective
species, including all subspecies and many arctic areas.
Comments:
(1)
Currently three subspecies are accepted in the North Atlantic area and all reach the
Arctic: subsp. alpinum, subsp. glabratum (Hartm.) Á. Löve & D. Löve and subsp.
lanatum (Lam.) Asch. & Graebn. See Jonsell (in prep., Flora Nordica 2) and Grundt
et al. (2000) for discussions. These entities seem to encompass the major variation in
the mainland European and Iceland parts but not in Svalbard, Jan Mayen, Greenland
og eastern Canada. A fourth, arctic entity may be present in these areas but is still
unproved. It is preliminarily here included in subsp. alpinum and not in the
(2)
comparatively southern and thermophilous subsp. lanatum as done by Hultén (1956).
There is not much similarity between these arctic plants and the subsp. lanatum that
Hultén refers them to. Subsp. lanatum is originally described from C Europe. (Elven
& Grundt)
The triploid count from Kola Peninsula is strongly deviating from the very numerous
tetraploid counts of this species. The voucher specimen (if any) should be inspected.
One possible explanation is a hybrid C. alpinum x C. cerastoides. (Elven)
47.2.5.1 Cerastium alpinum L. subsp. alpinum
S
2n=
72 (4x).
2nD
Jonsell (in prep., Flora Nordica 2, S and N Norw, Sb); Brysting et al. (2000 N Eur).
G
ICE NOR RUS CAN? GRL?
Comments:
(1)
The question marks for Canada and Greenland do not indicate doubt about the
presence (frequent in NE Canada and S-C Greenland) but about the identity with
subsp. alpinum. At least major parts of the Greenland material belongs to the
'unnamed' entity mentioned above. (Brysting & Elven)
47.2.5.2 Cerastium alpinum L. subsp. lanatum (Lam.) Asch. & Graebn. (1917), Syn.
Mitteleur. Fl. 5, 1: 619.
B
C. lanatum Lam. (1785), Encycl. 1: 680.
S
C. alpinum L. var. lanatum (Lam.) Hegetschw. (1825), ***
T
Lamarck s.n. (P-LA) lectotype, selected by Boscaiu et al. (1997, Willdenowia 27:
41).
2n=
72 (4x).
2nD
Jonsell (in prep., Flora Nordica 2, Icel); Brysting et al. (2000 N Eur).
G
ICE NOR RUS?
Comments:
(1)
The question mark for European Russia results from doubts about occurrence in Kola
Peninsula. The subspecies occur straight up to the border river on the Norwegian side
and is expected to occur at least in Rybachi Peninsula. (Elven)
47.2.5.3 Cerastium alpinum L. subsp. glabratum (Hartm.) Á. Löve & D. Löve (1956), Acta
Horti Gothob. 20: 141.
B
C. glabratum Hartm. (1820), Handb. Skand. Fl. 180.
S
T
Ill. in Vahl (1790), Fl. Dan. 6, 17, Fig. 979, lectotype, selected by Jonsell (1996,
Nord. J. Bot. 16: 5).
2n=
72 (4x).
2nD
Jonsell (in prep., Flora Nordica 2, N Norw); Brysting et al. (2000 N Eur).
G
ICE NOR RUS
Comments:
(1)
There is a previous C. alpinum L. var. (b) glabrum Wahlenb. (1812, Fl. Lapp. 136),
the first valid name if the entity is treated as a variety. (Elven)
47.2.6 Cerastium nigrescens (H.C. Watson) Edmondston ex H.C. Watson (1860), Cybele
Brit., Suppl. 1: 81.
B
C. latifolium L. var. nigrescens H.C. Watson (1847), Cybele Brit. 1: 233.
S
C. arcticum auct. plur., non Lange (1880).
T
Shetland: Baltasound, 08.1844, leg. Edmondston, herb. H.C. Watson H221/90 10 (K)
lectotype, selected by Brummit et al. (1987, Watsonia 16: 295).
2n=
108 (6x).
2nD
Brysting (1999 Norw, Icel, Brit. I.), also with references to previous reliable counts.
G
ICE GRL?
Comments:
(1)
Now well documented to be as different from the truly arctic plants (C. arcticum s.
str.) as to deserve status as a distinct species. It reaches the Arctic in Iceland and
possibly in SW Greenland.
The plants that reach the Arctic are var. laxum (Lindbl.) Brysting & Elven
(2000), based on C. latifolium L. var. laxum Lindbl. (1837-38), Physiogr. Sällsk.
Tidskr. 1: 334, with type: Norway: Dovre, Kongsvold, ad Södra Nystubäck infra
terminum betulinum, 07.1837, leg. Lindblom (O) lectotype, selected by Brysting &
Elven (2000). Var. nigrescens is confined to the serpentine areas in Shetland. The
treatment as variety is necessary because the type of the species belong to a very local
serpentine race - ecotype - that should not be given higher status than as variety. We
should perhaps enter var. laxum formally in the list. (Brysting & Elven)
47.2.7 Cerastium arcticum Lange (1880), Fl. Danica 17, 50: 7, t. 2963.
S
C. nigrescens (H.C. Watson) Edmondston ex H.C. Watson subsp. arcticum (Lange)
P.S. Lusby in Brummitt et al. (1987), Watsonia 16: 295; C. arcticum Lange var.
procerum (Lange) Hultén (1956), Sv. Bot. Tidskr. 50: 451; C. arcticum Lange var.
vestitum Hultén (1956), Sv. Bot. Tidskr. 50: 453; C. arcticum Lange var. sordidum
Hultén (1956), Sv. Bot. Tidskr. 50: 456; C. hyperboreum Tolm. (1930), Skr. Svalb.
Ishavet 34: 6.
T
W Greenland: Upernivik, leg. J. Vahl (UPS) lectotype, selected by Hultén (1956, Sv.
Bot. Tidskr. 50: 459).
2n=
(1) 54 (3x). (2) 108 (6x). (3) c.130.
2nD
(1) Böcher & Larsen (1950 Grl, a single count). (2) Brysting (1999 Grl, Sb), with
references to previous reliable counts. (3) Brysting (1999 SW Grl, possibly an
aneuploid number found in several plants in one population).
G
NOR RUS CAN GRL
Comments:
(1)
A morphologically and genetically heterogeneous high-ploid, but the intricate system
of subspecific categories presented by Hultén (1956), and his hypothesis of
widespread introgression, is not supported by recent studies (Brysting 1999, Brysting
et al. 2000). Neither is the subspecies proposed by Böcher (1977). At present it is best
treated as one variable species that reaches from the western Canadian Arctic (Banks
Island) across Greenland and Svalbard to the western Russian Arctic (at least to
Novaya Zemlya). (Brysting & Elven)
47.2.8 Cerastium beeringianum Cham. & Schltdl. (1826), Linnaea 1: 62.
S
C. beeringianum Cham. & Schltdl. subsp. beeringianum; C. beeringianum Cham. &
Schltdl. var. beeringianum; C. beeringianum Cham. & Schltdl. var. grandiflorum
(Fenzl) Hultén (1937), Fl. Aleut. Isl. 165; C. bialynickii Tolm. (1927), Trav. Mus.
Bot. Acad. Sci. URSS 21: 81; C. beeringianum Cham. & Schltdl. subsp. bialynickii
(Tolm.) Tolm. (1971), Fl. Arct. URSS 6: 45.
T
Alaska: Sin. Kotzebue (Eschscholtz Bay), 1816, leg. Chamisso (B) holotype?
[cotype? in LE].
2n=
72 (4x).
2nD
Löve & Löve (1975) list several counts, at least one each from rctic NE Asia and
North America; Zhukova (1980 S Chuk); Zhukova (1965b, 1968, 1969 Sib, NE As
for C. bialynickii).
G
RUS? SIB RFE ALA CAN GRL
Comments:
(1)
The species seems to be fairly well separated from C. arcticum (and the more distant
C. alpinum), at least in the Canadian Arctic where all three meet, but its range must
be extended eastwards at least to NW Greenland as at least one specimen from here is
available in HbO. (Brysting & Elven)
(2)
(3)
(4)
(5)
Cerastium bialynickii have to be reduced to synonymy. The material available for
typification (in LE) is not significantly different from C. beeringianum as typified
(from Dikson close to the Yenisey estuary). Another matter is the application of the
name. It has been generally applied in Russia for more northern plants than 'typical'
C. beeringianum, including a distinctive dwarf 'type' from the northernmost areas
(Severnaya Zemlya, northernmost Taimyr Peninsula, parts of the Novosiberian
Islands). This dwarf 'type' is also present throughout the northernmost parts of arctic
Canada and is the one proved from Greenland.
The 'arctic dwarf' might be recognised as a taxon, if one wanted to do some
splitting, but C. bialnynickii would not be the name for it. The preliminary decision is
therefore to reduce C. bialynickii to synonymy and to leave the 'arctic dwarf'
unnamed. (Brysting, Petrovsky & Elven)
The two Hultén (1956) varieties have been included in the synonymy. We have,
however, problems with a prostrate, nearly glabrous plant in the southern parts of
arctic Canada, mainly on the mainland. (Brysting & Elven)
'RUS?' is based on one record from the W coast of Novaya Zemlya, mapped and
mentioned by Fl. Arct. URSS 6. (Elven)
The 'grandiflorum' entity is possibly worthy of status as a separate taxon (variety,
subspecies). (Petrovsky & Elven)
47.2.9 Cerastium fischerianum Ser. in DC. (1824), Prodr. 1: 419.
S
T
Kamtchatka, Herbarium De Candolle, leg. Merk (Geneva) holotype. Type specimen
shown in Hultén (1930) Fl. Kamtsch. 4: 249.]
2n=
72 (4x).
2nD
Sokolovskaya (1960 NE As).
G
RFE? ALA
Comments:
(1)
Doubts about occurrence of this species in arctic Chukotka. (Petrovsky)
47.2.10 Cerastium regelii Ostenf. (1910), Skr. Norske Vidensk.-Akad. Christ., Math.Naturvidensk. Kl. 1908, 8: 10.
S
C. regelii subsp. caespitosum (Malmgr.) Tolm. (1971), Fl. Arct. URSS 6: 41
[basionym: C. alpinum L. var. (gamma) caespitosum Malmgr. (1862), 'Spetsb.
Fanerogamfl.' 242]; C. jenisejense Hultén (1956), Svensk Bot. Tidskr. 50: 473.
T
Canada: King William Land, 31.07.1904, leg. Lindström (O) lectotype, selected by
Jonsell (1996, Nordic J. Bot. 16: 6).
2n=
72 (4x).
2nD
Löve & Löve (1975) list numerous counts both for C. jenisejense and for C. regelii
(both subspecies), all as arctic; Engelskjøn (1979 Sb); Zhukova (1980, 1982 Chuk).
G
NOR RUS SIB RFE ALA CAN GRL
Comments:
(1)
Heide et al. (1990, Nord. J. Bot. 10: 141-147) convincingly proved that C. jenisejense
is conspecific with C. regelii, only a phenotypic response to day length. Similar
southern and regularly flowering plants are found on the American side. It should
therefore be reduced to synonymy. It should also be a warning against too easy
acceptance of 'zonal' taxa. (Elven)
(2)
A subspecific division has been indicated (e.g., by Tolmachev 1971) into a mainly
European - W Siberian subsp. caespitosum and an American - E Siberian subsp.
regelii. Northern arctic American material (from where 'regelii' s. str. is described)
confirms completely with the Svalbard materials (from where 'caespitosum' is
described). There is no difference whatsoever, and this subspecific division is
impossible to uphold. (Elven)
47.2.11 Cerastium fontanum Baumg. (1816), Enum. Stirp. Transsilv. 1: 425.
S
T
Romania: Transsylvania, "Fogarasser Alpen, am Zirnaflusse", 07.1851, leg. P.J.F.
Svhur (E) neotype, selected by Wyse Jackson (2000, Nord. J. Bot. 20: ***).
Comments:
(1)
The name C. caespitosum Gilib. (1782) must be rejected because binary
nomenclature was not consistently applied in 'Fl. Lit. Inch.' (acc. to Stafleu). When
the Linnaean C. vulgatum (illegitimate) and C. caespitosum are rejected, the oldest
valid name for the collective species seems to be C. fontanum. See also Wyse Jackson
(1995). (Elven)
(2)
A polymorphic species. At least two distinct races, hete treated as subspecies, reach
the Arctic in the N Atlantic area. The native race in arctic N Europe and Greenland is
C. fontanum subsp. fontanum [syn: C. fontanum Baumg. subsp. scandicum Gartn.].
The anthropochorous race is C. fontanum subsp. vulgare (Hartm.) Greuter & Burdet.
(Elven)
47.2.11.1 Cerastium fontanum Baumg. subsp. fontanum
S
C. fontanum Baumg. subsp. scandicum Gartn. (1939), Feddes Repert., Beih. 113: 68.
2n=
144 (8x).
2nD
Böcher (1938b Scand); Löve & Löve (1956b Icel).
G
ICE NOR RUS GRL
Comments:
(1)
Even if this race is not usually recognized in Russian floras it must be frequent in
parts of European Russia. Mapped as such for Kola Peninsula by Fl. Murm. Obl. 3
and reported eastwards to Kanin Peninsula. (Elven)
47.2.11.2 Cerastium fontanum Baumg. subsp. vulgare (Hartm.) Greuter & Burdet (1982),
Willdenowia 12: 37.
B
C. vulgare Hartm. (1820), Handb. Skand. Fl. 182.
S
C. caespitosum Gilib. (1782), nom. illegit., Fl. Lit. Inch. 2: 159 [see comment under
the species; alternative: Aschers. (1860), Fl. Prov. Brandenb. 1: 102]; C. triviale Link
(1821), Enum. Hort. Berol. Alt. 1: 433; C. vulgatum L. (1762), non L. (1755), nom.
illegit.; C. holosteoides Fr. (1817), Novit. Fl. Suec. 4: 52.
T
Sweden: Gästrikland, 'Gefle' (Gävle), 1833, leg. C.J. Hartman (UPS) neotype,
selected by Wyse Jackson (1995, Nordic J. Bot. 15: 562).
2n=
144 (8x).
2nD
Löve & Löve (1975) list numerous counts, one Icelandic; Lövkvist & Hultgård (1999
S Sweden, 11 counts).
G
NOR RUS CAN* GRL*
Comments:
(1)
Possibly an archeophyte in N Norway and N Russia and to be indicated as introduced
also here. (Elven)
47.3 Sagina L. (1753), Sp. Pl. 128. [VP]
S
Spergella Rchb. in Moessl. (1827), Handb., ed. 2, 1: lxv.
Comments:
(1)
The order of species is changed from Petrovsky's draft as that did not reflect the
current subgeneric classification. Three of the included species belong to sect.
Saginella Koch (1843), Syn., d. 2, 117 [x normaniana, procumbens, saginoides,
subulata], three to sect. Spergella (Rchb.) *** [cespitosa, intermedia, nodosa].
Subgeneric classification is omitted as there are only seven taxa. (Elven)
47.3.1 Sagina saginoides (L.) H. Karst. (1882), Deut. Fl. 539.
B
Spergula saginoides L. (1753), Sp. Pl. 441.
S
T
S. linnaei Presl (1831), Reliq. Haenk. 2: 14.
Linnaean herbarium 604.6 (LINN) lectotype, selected by Crow (1978, Rhodora 80:
25).
2n=
22 (2x).
2nD
Löve & Löve (1975) list several counts, some as arctic.
G
ICE NOR RUS SIB RFE CAN GRL
Comments:
47.3.2 Sagina procumbens L. (1753), Sp. Pl. 128.
S
T
Séguier, Tabula 5, Fig. 3 in Plantae Veronenses 1: 421 (1745), selected by Jonsell &
Jarvis in Jarvis et al. (1993, Regnum Vegetabile 127: 83).
2n=
22 (2x).
2nD
Löve & Löve (1975) list numerous counts, a few from close to the Arctic.
G
ICE* NOR* RUS* GRL*
Comments:
(1)
Possibly (or probably?) a stable archaeophyte in most or all northern areas and
therefore to be treated as introduced. (Elven)
47.3.3 Sagina x normaniana Lagerh. (1898), Kongel. Norske Vidensk. Selsk. Skr. 1898, 1:
4.
S
S. procumbens L. x S. saginoides (L.) Karst.
T
Cultivated material from N Norway, Tromsø, collected by Lagerheim, 28.06.1898,
leg. Holmberg (LD) lectotype, selected by Nilsson in Jonsell (1996, Nord. J. Bot. 16:
6).
2n=
2nD
G
NOR
Comments:
(1)
Added to Petrovsky's draft. Occurs independent of its parents and reaches the Arctic
at least in N Norway. The hybrid is partly fertile and reproduces both vegetatively
and by seeds. (Elven)
47.3.4 Sagina subulata (Sw.) C. Presl (1826), Fl. Sicul. 158.
B
Spergula subulata Sw. (1789), Kongl. Vetensk. Acad. Nya Handl. 10: 45.
S
T
Sweden, Halland, leg. P. Osbeck (BM) lectotype, selected by Crow (1978, Rhodora
80: 51).
2n=
22 (2x).
2nD
Löve & Löve (1975) list several counts, one Icelandic.
G
ICE
Comments:
(1)
Added to Petrovsky's draft as it occurs in the arctic parts of Iceland (Kristinsson 1987
and his later list and maps). (Elven)
47.3.5 Sagina nodosa (L.) Fenzl (1833), Vers. Darstell. Alsin., tab. ad 18.
B
Spergula nodosa L. (1753), Sp. Pl. 440.
S
T
Linnaean herbarium 604.4 (LINN) lectotype, selected by Crow (1978, Rhodora 80:
25).
2n=
(1) c. 52. (2) 56.
2nD
(1) Jonsell (in prep., Flora Nordica 2, S Sweden). (2) Löve & Löve (1975) list several
counts, some Icelandic; Lövkvist & Hultgård (1999 N Norw, S Sweden); Jonsell (in
prep., Flora nordica 2, N Norw).
G
ICE NOR RUS SIB CAN GRL
Comments:
(1)
Two subspecies are recognised by Crow (1978) and Karlsson in Jonsell (in prep.,
Flora Nordica 2): subsp. nodosa and subsp. borealis G.E. Crow (1978), Rhodora 80:
28. These are more or less sympatric in the Nordic area and I have no very
determined opinion about them. The Nordic material has recently been revised by Th.
Karlsson. Both occur (with intermediates) in the arctic parts of North Norway. Subsp.
nodosa is, acc. to Crow (1978), mainly European and may have been introduced in
America. Subsp. borealis is mainly American, only found in glaciated parts of N
Europe. The reported differences (Crow, Karlsson) are: 'borealis' - glabrous stems,
often prostrate, shorter stems and rosette leaves, few-flowered, with numerous
bulbils; 'nodosa' s. str. - glandular hairs on stems, often erect, longer stems and rosette
leaves, many-flowered, with few or without bulbils. Type of subsp. borealis: Canada:
Quebec, Gaspé County, 19.08.1905, leg. Collins & Fernald 75 (MSC) holotype. We
must decide between acceptance of them, with normal entries in the checklist, or only
reference in synonymy. A decision depends on whether the subspecies can be applied
consistently throughout the circumpolar area. (Elven)
47.3.6 Sagina nivalis (Lindblom) Fr. (1842), Mant. 3, 2: 31.
B
Spergula saginoides L. var. nivalis Lindblom (1838), Physiogr. Sällsk. Tidskr. 1: 328.
S
S. intermedia Fenzl in Ledeb. (1842), Fl. Ross. 1: 339; Spergella intermedia (Fenzl)
Á. Löve & D. Löve (1976), Bot. Not. 128: 508.
T
Norway: Sør-Trøndelag, Oppdal, Kongsvold, 18.09.1839, Lindblom scr. (LD)
neotype, selected by Nilsson in Jonsell (1996, Nordic J. Bot. 16: 6).
2n=
56.
2nD
Löve & Löve (1975) list several counts, all as arctic.
G
ICE NOR RUS SIB RFE ALA CAN GRL
Comments:
(1)
Sagina nivalis (July 1842) is now generally accepted to have priority before S.
intermedia (October 1842), see Stafleu. (Elven)
47.3.7
B
S
T
2n=
2nD
Sagina cespitosa (J. Vahl) Lange in Rink (1857), Grønl. Geogr. Stat. Beskr. 2, 6: 133.
Arenaria cespitosa J. Vahl (1840), Fl. Dan. 39, t. 2289.
Spergella cespitosa (J. Vahl) Á. Löve & D. Löve (1976), Bot. Not. 128: 508.
Greenland, leg. Vahl (C) lectotype, selected by Crow (1978, Rhodora 80: 57).
(1) 84. (2) 88.
(1) Löve & Löve in Löve (1975c). (2) Knaben (1950 Norw); Löve & Löve (1965b
Icel?).
G
ICE NOR CAN GRL
Comments:
(1)
Epithet changed from the 'caespitosa' applied by J. Vahl to 'cespitosa'. Is this
legitime? (Elven)