86 Boraginaceae (Draft: V.V. Petrovsky)
Comments:
(1)
Three genera in Petrovsky's draft are excluded (Amsinckia, Cryptantha and Lycopsis)
and two more are in danger (Asperugo, Lappula).
Amsinckia Lehm. (1831), Sem. Hort. Bot. Hamburg. 1: 7, with two species.
Both are most probably casual in the Arctic and should be excluded: A. menziesii
(Lehm.) Nelson & Macbride (1916), Bot. Gaz. (Crawfordsville) 61: 36 [basionym:
Echium menziesii Lehm. (1830), Nov. Stirp. Pug. 2: 29] - given from ALA; and A.
lycopsoides (Lehm.) Lehm. (1831), Sem. Hort. Bot. Hamburg. 1831: 3 [basionym:
Lithospermum lycopsoides Lehm. (1830), Nov. Stirp. Pug. 2: 28] - also given from
ALA. (Elven)
Cryptantha Lehm. ex Fisch. & Mey. (1836), Ind. Sem. Hort. Petrop. 2: 35,
with C. spiculifera (Piper) Payson (1927), Ann. Miss. Bot. Gard. 14: 298 [basionym:
Oreocarya spiculifera Piper (1906), Contr. U.S. Nat. Herb. 11: 481]. Reported from
RFE. The Chukotka record is based on a single casual occurrence, but probably as a
native (or non-anthropogenic) casual. The identity of the Chukotka plant with this
species is doubted by D.F. Murray. This only Asian specimen of this American genus
should be checked by some American expert before acceptance.
Lycopsis L. (1753), Sp. Pl. 138, with L. arvensis L. (1753), Sp. Pl. 139
[synonym: Anchusa arvensis (L.) M. Bieb. (1808), Fl. Taur.-Caucas. 1: 123]. Given
with 2n=48 (8x); Lövkvist (1963 Sweden); Gadella & Kliphuis (1970 S Eur).
Reported from N Fennoscandia as a casual weed. It is obviously a much too
thermophilous species to be stable anywhere in the Arctic. Therefore omitted, and
then we need not decide between Lycopsis or Anchusa.
For Asperugo and Lappula, see comments below. (Elven)
86.1 Eritrichium Schrad. (1820), Comm. Gotting. 4: 186, in obs.
Comments:
(1)
Löve & Löve (1975) list another species, E. czekanowskii Trautv. (1877), Acta Hort.
Petrop. 5: 88, from Siberia. The name is absent from Fl. Arct. URSS 8. Is it a
worthwhile taxon, and if so does it occur within (or near) the Arctic? (Elven)
86.1.1
B
S
2n=
2nD
Eritrichium villosum (Ledeb.) Bunge (1836), Verz. Pfl. Altai Gebirge 14.
Myosotis villosa Ledeb. (1815), Mém. Acad. Sci. Pétersb. 5: 516.
For the collective species. (1) 24 (4x, x=6). (2) 48 (8x, x=6).
(1) Zhukova (1968, 1969); Zhukova & Tikhonova (1973); Zhukova et al. (1973). (2)
Zhukova et al. (1973).
Comments:
(1)
The chromosome counts were published before the recognition of subsp. pulvinatum.
Petrovsky should try to sort the numbers on subspecies. (Elven)
(2)
The two subspecies are nearly sympatric - subsp. pulvinatum occurs within the range
of subsp. villosum. How different are they and what keeps them apart? (Elven)
WARNING! Subspecies might be omitted if not sufficiently well differentiated.
86.1.1.1 Eritrichium villosum (Ledeb.) Bunge subsp. villosum
S
2n=
2nD
G
RUS SIB RFE
Comments:
86.1.1.2 Eritrichium villosum (Ledeb.) Bunge subsp. pulvinatum Petrovsky in Tolm. &
Jurtz. (1980), Fl. Arct. URSS 8: 241.
S
T
Type: Siberia: Pars orientalis peninsulae Taimyr, sinus Mariae Pronczisczeviae,
26.07.1972, leg. N. Matveeva 380 (LE) holotypus.
2n=
2nD
G
SIB RFE
Comments:
86.1.2 Eritrichium sericeum (Lehm.) DC. (1846), Prodr. 10: 126.
B
Myosotis sericea Lehm. (1818), Pl. Asperif. Nucif. 2: 98.
S
2n=
For the collective species. 28 (4x, x=7).
2nD
Zhukova & Petrovsky (1971); Zhukova et al. (1973).
Comments:
(1)
The chromosome reports counts were published before the recognition of subsp.
arctisibiricum. Petrovsky should try to find out to which of the subspecies they
belong. (Elven)
(2)
The distribution of subsp. arctisibiricum is very strange, disjunct from Taimyr with
surroundings to Wrangel Island whereas the taxon between these areas is subsp.
sericeum. This is not a consistent phytogeographic pattern and needs some
explanation before the taxa are fully accepted as significantly different. (Elven)
WARNING! Subspecies might be omitted if not justified as sufficiently differentiated.
86.1.2.1 Eritrichium sericeum (Lehm.) DC. subsp. sericeum
S
2n=
2nD
G
SIB RFE
Comments:
86.1.2.2 Eritrichium sericeum (Lehm.) DC. subsp. arctisibiricum Petrovsky in Tolm. &
Jurtz. (1980), Fl. Arct. URSS 8: 240.
S
T
Type: Russian Far East: Insula Wrangelii (pars occidentalis) in cursu medio fl.
Gusinaja, in valli rivuli Korall, 02.07.1970, leg. P. Zhukova & V.V. Petrovsky 70-48
(LE) holotype.
2n=
28 (4x, x=7).
2nD
Petrovsky in Tolmachev & Yurtsev (1980 NE As).
G
SIB RFE
Comments:
86.1.3 Eritrichium splendens Kearney ex W.F. Wight (1902), Bull. Torrey Bot. Club 1920:
410.
S
T
Alaska: Old Man Creek, 09.07.1901, leg. Mendenhall (US) holotype.
2n=
36 (6x, x=6).
2nD
Johnson & Packer (1968 Ala).
G
ALA
Comments:
86.1.4 Eritrichium aretioides (Cham.) DC. (1846), Prodr. 10: 125.
B
Myosotis aretioides Cham. (1829), Linnaea 4: 443.
S
T
[Described from the Bering Strait area: "Ad sinum St Laurentii ... in insula Sancti
Laurentii", probably erroneous in St. Lawrence Island.]
Comments:
86.1.4.1 Eritrichium aretioides (Cham.) DC. var. aretioides
S
2n=
24 (4x, x=6).
2nD
Löve & Löve in Löve (1975d).
G
RFE ALA
Comments:
(1)
This variety seems to be excluded from the arctic in Fl. Arct. URSS 8 (p. 243).
(Elven)
86.1.4.2 Eritrichium aretioides (Cham.) DC. var. chamissonis (DC.) Petrovsky in Tolm. &
Jurtz. (1980), Fl. Arct. URSS 8: 243.
B
E. chamissonis DC. (1846), Prodr. 10: 125.
S
T
[Described from Russian Far East: "Ad Sinum Sancti Laurentii ad fretum Beering ...
et Kamtchatka".]
2n=
48 (8x, x=6).
2nD
Johnson & Packer (1968 Ala); Zhukova et al. (1973 NE As).
G
RFE ALA
Comments:
86.1.5 Eritrichium tschuktschorum Jutz. & Petrovsky in Tolm. Jurtz. (1980), Fl. Arct.
URSS 8: 244.
S
T
Type: Russian Far East: Districtus autonomicus Tschukotskij, planiteis elata
Anjuensis, in systemate fl. Malyi Anjui, ad fontes fl. Koralveem, prope opidulum
Bezymyannyi, 23.06.1966, leg. P. Zhukova & V.V. Petrovsky 66-44 (LE) holotype.
2n=
20 (x=?).
2nD
Petrovsky in Tolmachev (1980 NE As).
G
RFE
Comments:
86.2 Mertensia Roth (1797), Catal. Bot. 1: 34.
86.2.1 Mertensia maritima (L.) Gray (1821), Nat. Arr. Brit. Pl. 2: 354.
B
Pulmonaria maritima L. (1753), Sp. Pl. 136.
S
T
[Described from Britain.]
2n=
24 (4x).
2nD
Löve & Löve (1975) list several counts for the species, most as arctic.
Comments:
(1)
The chromosome counts listed by Löve & Löve (1975) probably include both
varieties. (Elven)
(2)
The arctic race - var. tenella - seems to be both morphologically and genetically
distinct from the more southern var. maritima. Skarpås (1999) found total identity in
genetical markers (enzymes) in var. maritima from southernmost to northernmost
mainland Norway and clear differences from arctic (Svalbard) material og var.
tenella. Var. tenella is described from Svalbard. The arctic race is also proved
(morphologically) from Jan Mayen, Greenland and arctic Canada whereas, e.g., all
the Icelandic material I have seen is var. maritima.
The identity of the north Alaskan and east Russian material should also be
checked. The plant illustrated in Hultén (1968) is a typical var. tenella and the
drawing is made from an Alaskan plant. The west Russian (Kola) material is var.
maritima. The arctic race probably deserves status as subspecies but does not seem to
have been described as such yet. (Elven)
WARNING! The varieties should probably be raised to rank of subspecies.
86.2.1.1 Mertensia maritima (L.) Gray var. maritima
S
2n=
24 (4x).
2nD
Löve & Löve (1956 Icel).
G
ICE NOR RUS RFE? ALA? CAN?
Comments:
(1)
The question marks for RFE, ALA and CAN refer to which race; the species is
present in all areas. (Elven)
86.2.1.2 Mertensia maritima (L.) Gray var. tenella Th. Fr. (1869), Öfvers. K. Vet.-Akad.
Förh. 26: ***
S
T
[Described from Svalbard.]
2n=
2nD
G
NOR RFE? ALA? CAN GRL
Comments:
(1)
The question marks for RFE, ALA and CAN refer to which race. (Elven)
86.2.2 Mertensia drummondii (Lehm.) G. Don (1838), Gen. Hist. 4: 319.
B
Lithospermum drummondii Lehm. (1830), Nov. Stirp. Pug. 2: 26.
S
2n=
2nD
G
ALA CAN
Comments:
86.2.3 Mertensia sibirica (L.) G. Don (1838), Syst. Veg. 4: 319.
B
Pulmonaria sibirica L. (1753), Sp. Pl. 135.
S
2n=
2nD
G
SIB
Comments:
86.2.4 Mertensia pubescens (Willd. ex Roem. & Schult.) DC. (1846), Prodr. 10: 90.
B
Pulmonaria pubescens Willd. ex Roem. & Schult. (1819), Syst. Veg. 4: 744.
S
M. kamczatica (Turcz.) DC. (1846), Prodr. 10: 100.
2n=
2nD
G
RFE
Comments:
WARNING! Might be reduced to a race of M. paniculata.
86.2.5 Mertensia paniculata (Ait.) G. Don (1838), Gen. Hist. 4: 318.
B
Pulmonaria paniculata Ait. (1789), Hort. Kew., ed. 1, 1: 181.
S
2n=
For the collective species. 24 (4x).
2nD
Löve & Löve in Löve (1975d).
Comments:
(1)
Three varieties are accepted in American standard sources and must either be
accepted as races or included in the synonymy. Two of these are var. paniculata and
var. alaskana (Hultén 1968, Porsild & Cody 1980, Cody 1996). The third, var.
eastwoodae, is accepted as a species in Petrovsky's draft but should be justified as
such. Due to Hultén's arguments (Hultén 1968, Ark. Bot.) I have reduced it to a
variety in this draft version. Mertensia pubescens belongs in the same complex and
should be re-evaluated with the others. (Elven)
86.2.5.1 Mertensia paniculata (Ait.) G. Don var. paniculata
S
2n=
2nD
G
ALA CAN
Comments:
86.2.5.2 Mertensia paniculata (Ait.) G. Don var. alaskana (Britt.) O.L. Williams (1937),
Ann. Miss. Bot. Gard. 24: 48.
B
Mertensia alaskana Britt. (1901), Bull. New York Bot. Gard. 2, 6: 181.
S
T
[Described from Alaska.]
2n=
2nD
G
ALA? CAN
Comments:
(1)
A border case in the Porcupine River valley and Richardson Mts in Yukon Territory,
possibly also on the Alaskan side. (Elven)
86.2.5.3 Mertensia paniculata (Ait.) G. Don var. eastwoodae (Macbr.) Hultén (1968), Ark.
Bot., ser. 2, 7, 1: 116.
B
Based on M. alaskana Eastwood (1902) non Britton (1901).
S
M. eastwoodae Macbride (1917), Contrib. Gray Herb., n. s., 49: 18.
T
[Described from Alaska: Nome.]
2n=
2nD
G
ALA
Comments: