Chapter 12, The Origins, Evolution, and Dispersal of Modern People What’s So Modern about Modern Humans? Modern humans are different from archaic humans. Skeletal traits: round, tall skull with vertical forehead, small brow ridges, Small face and teeth with a more gracile postcranial skeleton Some hominid skeletons have a mixture of archaic and modern traits. Modern Homo sapiens: Single Origin and Global Dispersal or Regional Continuity? Homo erectus already established across Africa and southern Europe and Asia Evolution of modern humans began about 350,000 BP with emergence of archaic forms. First modern Homo sapiens appeared in Africa, 200,000 BP. Two main hypotheses: a) Out of Africa Modern Homo sapiens evolved in Africa, and then spread to Asia and Europe, where modern humans replaced the populations there. b) Multiregional Continuity The shift to modern humans took place regionally and did not involve replacement. What Do Early Homo sapiens Fossils Tell Us about Modern Human Origins? Archaic Homo sapiens in Africa (350,000–200,000 BP), Kabwe (Broken Hill) Early Archaic Homo sapiens in Asia (350,000–130,000 BP), Ngandong (Java) Early Archaic Homo sapiens in Europe (350,000–130,000 BP) , Atapuerca, (Spain) Early Archaic Homo sapiens’ Dietary Adaptations Same tools and material technology as Homo erectus but diverse materials Reduction of dental complex with tool use Late Archaic Homo sapiens in Asia (60,000–40,000 BP) Neanderthals found in Amud, Kebara, and Tabun Shanidar, several individuals with interesting life histories seen in their skeletons Late Archaic Homo sapiens in Europe (130,000–24,000 BP) Some Neanderthal bones suggest cannibalism The Neandertal Body Plan: Aberrant or Adapted? La Chapelle-aux-Saints studied first by Boule Boule influenced thinking about the Neandertals Thought Neandertals had bent-kneed gait Thought Neandertals were primitive and stupid Cold adaptation of Neandertals Large nasal aperture Large infraorbital foramina Stocky build, short limbs Short, heavy body plan Emphasis on megafauna Extensive use of fire Use of caves Neanderthals Hunting: Inefficient or Successful? Established Mousterian tool tradition Butchered animals, processed for food Chemical evidence indicates importance of meat in diet. Neandertals Buried Their Dead European, western Asian sites indicate burial; others do not. Neanderthals Talked Some argue Neandertals could not produce range of sound necessary for language. Kebara hyoid bone suggests that Neandertals could talk. Review of Major Neanderthal traits 1) Physical appearance: Skull: Heavy brow ridges Elongated skull with occipital bun Large brains Body: Shorter legs, longer arms Barrel chest Heavy bone structure 2) Behavior: Lived in small family groups Roamed close to home Buried their dead Cared for injured and elderly Hunted large animals (megafauna) 3) Material culture: Sharp spear points, thrusting spear Used adhesives, wore skin clothing Wide variety of stone knives, etc. Possible musical instrument (flute) Possible bone needle, necklaces Early Modern Homo sapiens in Africa (200,000–6,000 BP) Herto, Aduma, Awash Valley, Omo Klasies River Mouth Cave, Hofmeyr Early Modern Homo sapiens in Asia (90,000–18,000 BP) Skhul, Tianyuan Cave, Zhoukoudian Early Modern Homo sapiens in Europe (35,000–15,000 BP) Oase, Predmostí, Dolni Vestnice Modern Behavioral and Cultural Transitions Happened First in Africa Painting, hunting, fishing, tool production, jewelry Once thought these originated in Europe At Katanda (Africa) catfish remains found from as early as 75,000 BP How Has the Biological Variation in Fossil Homo sapiens Been Interpreted? Ancient DNA: Interbreeding between Neanderthals and Early Modern People? Overlap in dates between H. sapiens and Neanderthals suggests coexistence, interbreeding. Supports “out of Africa” model Living People’s Genetic Record: Settling the Debate on Modern Humans’ Origins mtDNA shows similarity between Neanderthals and dissimilarity with modern humans. Lack of samples, time depth leading challenges to DNA analysis and interpretation. Assimilation Model for Modern Human Variation: Neanderthals Are Still with Us Discordance in the fossil and genetic record suggests that current models do not adequately explain modern human origins. Neanderthals contributed to modern gene pool in Europe and Asia. Modern Humans’ Other Migrations: Colonization of Australia, the Pacific, and the Americas Initial spread of population out of Africa, into Asia and Europe Spread to other parts of the world Down Under and Beyond: The Australian and Pacific Migrations Australia Lake Mungo, 40,000 BP , similarities with Homo erectus Pacific Southeast Asia point of origin, “Hobbit” man discovered in 2003 Arrival in America: The First Americans evidence DNA links Native Americans to East Asian populations Anatomical differences from recent Native Americans