Chapter 17 Classification Why is classification necessary? •Are you an organized person? – Consider – do you organize your room? Your clothes? Your school work? CDs? •What would happen if nothing was organized? – Would you be able to find anything? Scientists have identified more than 2 MILLION SPECIES!!! • Every year, thousands of new species are discovered. • There may be millions of undiscovered species • especially microbes, plants & insects • In the Tropical Rain Forest & Oceans Classification helps scientists understand & study living things. I. Biodiversity • Biodiversity is the variety of organisms on the earth. • Considered at all levels from populations to ecosystems. A. Taxonomy • Taxonomy is the science of describing, naming, and classifying organisms. • The branch of biology that names & groups organisms -according to their characteristics & evolutionary history. • A Universal System was designed to Eliminate the use of Common Names and Confusion in the Scientific World. The History of Taxonomy Aristotle – a Greek philosopher who lived more than 2,000 years ago –Classified things- Plant or Animal • Grouped Animals into Land Dwellers, Water Dwellers, and Air Dwellers. • Also grouped Plants into 3 categories, based on differences in their Stems. Aristotle’s classification system was replaced • As modern science Aristotle's system was found to be INADEQUATE. • Aristotle's categories did not work for all organisms. • his use of common names was problematic. Use of Common Names • COMMON NAMES, such as “robin” or “fir tree”, for organisms created some problems • common names varied from one locale to next & did not describe species accurately. Use of long Latin names • Used by scientists before 1700’s,- did not show relationships between species & were inconvenient, hard to understand. Describe a problem with each of these common names : -Starfish -Seahorse -Jelly fish -Peanut -Catfish -Tiger shark How are their names misleading? Discuss problems when 1 organism has 2 common names: Example- firefly & lightening bug Carl Linnaeus (1707-1778) “Father of Modern Taxonomy” • He used morphology (which is the organism’s structure & form) • Grouped organisms into hierarchical categories Carl Linnaeus • Formed “Taxa” (groups of organisms) • (Used Latin for the Names because it was the language of educated people) • Morphology -the STRUCTURAL SIMILARITIES BETWEEN ORGANISMS • Series of hierarchical categories used to show relationships • He had 2 KINGDOMS: PLANTAE & ANIMALIA. B. Modern classification system is based on morphological similarities. • Hierarchy of eight groups (Taxa) – Domains- include all six kingdoms – Kingdom – a taxon of similar phyla or divisions – Phylum – (phyla-plural)- taxon of similar classes – Class – taxon of similar orders – Order – taxon of similar families – Family – group of similar genera – Genus – group of similar species – species – most exclusive, specific group. Members of this grouping can mate, produce viable offspring – (varieties) – same species but with slight differences – (subspecies) – same species, different location 8 Modern Levels of Classification (From the most general to the most specific) • Domain • Kingdom • Phylum • Class • Order • Family • Genus • Species Species- is the smallest, most specific groupcontains only 1 kind of organism. Taxon - A particular group in a taxonomic system Most specific Taxon (group) Most general taxon Pneumonic Devices pick one to help you remember the taxa • Keep Penguins Cool Or Find Good Shelter or • Kennywood Park Can Open For Good Summers. or • King Phillip Comes Over For Good Spaghetti Classification Hierarchy of Organisms 3 Domains: (Most modern level by scientist Carl Woese) • These are Broad groups above the kingdom level. Archaea Bacteria Eukarya Binomial Nomenclature means Two Name Naming • Uses the last 2 categories (the most specific) to name things. uses the Genus & Species for the 2 parts of the name • Always Capitalize the Genus but Not the Species Identifier. • Both are either underlined or italicized. Using Binomial Nomenclature • Acer rubrum - RED MAPLE TREE • • • Acer is the Latin name for Maple (genus) rubrum is the Latin word for Red(species) Can be Abreviated: A.rubrum. • Homo sapiens - HUMANS • Homo -large brain & upright posture. sapiens for our intelligence & ability to speak. Abbreviated H. sapiens Additional Categories • Zoologists – use term “SUBSPECIES” for variations that may occur in species from different geographical locations- ie, timber wolf and the northern timber wolf- ex Canis Lupus ssp occidentalis • Botanists – May use the term “division” instead of phylum – sometimes split species into Subsets known as VARIETIES. Example: peaches & nectarines are varieties of Prunus persica var. Additional Categories • Microbiologists- Bacteria are also broken into subsets called STRAINS. Example: Escherichia Coli – some strains are harmless, even helpful;- live in our intestines, but strain E. coli 157 is responsible for food poisoning deaths. According to the CDC there are an estimated 73,000 cases of E. Coli infection every year in the United States. The typical symptoms are bloody diarrhea and (if severe) kidney failure. These symptoms most commonly appear when a person has eaten undercooked or contaminated ground beef. Testbook Assignment: • Read chapter 17, section 2 • do end of section questions • & Define: – Systematics – Phylogeny – Phylogenic Trees – Embryology, -blastula, -gastrula – Cladistics – derived characters, – cladogram II. Systematics • The way we group organisms continues to change Today these methods reflect the evolutionary history of organisms (What’s in their genes). This is called Phylogeny. A. Phylogeny • organizes the diversity of living organisms in the context of evolution. • are based on several types of evidence: 1. Fossil Record 2. Morphology 3. Embryology 4. Chromosomes & Macromolecules Phylogenic Trees • A family tree that shows evolutionary relationships thought to exist among organisms. • Is a hypothesis about the relationships. • Is subject to change - as more evidence is learned. II. Evidences for Evolutionary Relationships 1. Fossil Record • a useful tool for ancient organisms. • Record is incomplete – Some organisms overrepresented – Some organisms may be missing • Need other evidences to verify phylogenic relationships Fossils Types: actual preservation, petrification, imprints, molds, casts, footprints • Dated by radioactive isotopes in fossil or geological formation in which fossils are found • Requires long periods of time and unusual conditions for fossil preservation II. Evidences for Evolutionary Relationships 2. Morphology • Examine structure & function • Homologous structures- similar features that originated from similar ancestors. (forelimbs on bat, human, penguin) • Analogous structures- features that serve similar functions & look alike but originated from different embryonic tissues. (wings- on butterfly, bat, hummingbird) • Vestigial structures -serve no useful function any longer in the organism Homologous structures Comparing the structural features found in different organisms reveals a basic similarity. example is the forelimb of mammals - Although function is quite different, they are similar structurally. Analogous structures • We must look at structures that look & function the same but are not derived from the same embryonic tissue. • These features do not show recent, related ancestory. Vestigial structures • Features which serve no useful function any longer in the organism. • Examples: the pelvis bone in the whale, tailbone & appendix in humans, pelvis & leg bones in some snakes, etc II. Evidences for Evolutionary Relationships 3. Embryology • SIMILARITIES IN EARLY EMBRYOLOGICAL DEVELOPEMNT OF VERTEBRATES • CAN BE TAKEN AS ANOTHER INDICATION THAT VERTEBRATES MAY SHARE A COMMON ANCESTOR. At the blastula stage, scientists begin to look for differences in the ways organisms develop. Blastula - An early embryonic form produced by cleavage of a fertilized ovum - a spherical layer of cells surrounding a fluid-filled cavity.(think of a basketball) Gastrula - double-walled stage of the embryo succeeding the blastula; the outer layer of cells is the ectoderm and the inner layer differentiates into the mesoderm and endoderm Embryology exampleAt the blastula stage- what happens if a scientist separates a cell from the ball? • In Vertebrates (animals with a backbone) & Echinoderms animals like starfish & sand dollars)- any cell separated can produce a “twin”. • But blastula cells in a fruit fly cannot- the separated cells are already specialized to form a part & will die. • Conclusion- we are more closely related to starfish than insects II. Evidences for Evolutionary Relationships 4. Chromosomes & Macromolecules • Taxonomists compare Macromolecules like DNA, RNA & Proteins. • Example- the number of differences in amino acids is a clue to how long ago 2 species “Diverged” • Divergent Evolution- 2 species become more and more dissimilar. • Convergent Evolution- Species which have different ancestors, but have become more similar Proteins indicate degree of relatedness. Differences - Amino Acids in Protein Cytochrome C Number of different amino acids found in human cytochrome C as opposed to selected organisms Organism # of amino acids different compared to humans Human 0 Self (Family Hominidae, Order Primates) Monkey 1 Different family (Pongidae), same order (Primates) Pig, bovine, sheep 10 Horse 12 Dog 11 Rabbit 9 Chicken, Turkey 13 Duck 11 Rattlesnake 14 Turtle 15 Tuna 21 Different class (Ostheichthys), same phylum (Chordata) poikilothermic Moth 31 Different phylum (Arthropoda), same Kingdom (Animalia) Candida fungus 51 Different Kingdom (Fungi) • Different order (Carnivora), same class (Mammalia) Different class (Aves), same phylum (Chordata) homeothermic Different class (Reptilia), same phylum (Chordata) poikilothermic (From Atlas of Protein Sequence and Structure, 1967-68 by Margaret O. Dayhoff Chromosome comparisons • For example, cauliflower, cabbage, kale, and broccoli look different but have chromosomes that are almost identical in structure. Martin (1993). B. Cladistics • uses shared, derived characters as the only criterion for grouping taxa. • Shared character - A feature that all members of a group have in common • Derived character - A feature that evolved only within the group under consideration • is a newer way to display relationships • Derived Characters- are special features that apparently have only developed in that group. Examples: – feathers in birds – larger brains. Homosapiens have larger brains than the 'outgroup' (monkeys). The larger brain of homosapiens is a derived characteristic. • Clade – A Group of organisms that includes an ancestor plus all of its descendants Cladogram • Diagrams which show derived characters • Shared & derived characters are strong indicators of common ancestry. • Cladograms can show non-traditional conclusions about which organisms are “close cousins”. Cladogram From this cladogram, we can figure out that brown bears have more derived characters in common with sun bears than with dogs & lesser pandas are more closely related to racoons than giant pandas. Linnaean Taxonomy - compared to Phylogenetic Nomenclature: • Linnaean Taxonomy - Primary goal is to group species based on morphological similarities (who has 6 legs) • Phylogeny- Primary goal is to reflect the process of evolution (who’s close cousins) Dichotomous Key • graphically organizes data. • You start with a main idea, split that into two major pieces. • Those pieces are then split again into two major pieces. You continue splitting until you reach only one possible answer. • Each set of questions is called a couplet, & contains instructions for which couplet to go to next. Example of Dichotomous Key Dichotomous Key Use • Can be used by any one to classify anything. • Are used by taxonomists to classify organisms. III. Taxonomic Groups 3 domains, 6 kingdoms 1. Domain Archaea – aligns with Kingdom Archaebacteria, are single-celled prokaryotic cells that have distinctive cell walls & are “ancient bacteria” 2. Domain Bacteria – aligns with Kingdom Eubacteria, are singlecelled prokaryotic cells that are “true bacteria”. 3. Domain Eukarya – Domain Eukarya includes the kingdoms Protista, Fungi, Plantae, & Animalia. – All members of this domain have eukaryotic cells. • SIX KINGDOMS GROUP ORGANISMS TOGETHER THAT HAVE SIMILARITIES SUCH AS MAJOR CELLULAR STRUCTURE, METHODS OF OBTAINING NUTRIENTS, AND METABOLISM. A closer look at each of the 6 kingdoms: Domain: Archaea, kingdom Archaea vs. Domain: Bacteria, Kingdom Eubacteria • Organisms in the Kingdoms Eubacteria & Archaea are very different from each other, both genetically & biochemically! • Archaea have been found in temperatures above the boiling point and in cold that would freeze your blood. • Eubacteria are the “regular” bacteria. Domain: Archaea, Kingdom Archaea 1. Archaea • “archae”- from the Greek for "ANCIENT".Scientists think these are similar to Earth’s First Organisms • “Extremophiles” – Many types of Archaeans live in HARSH ENVIRONMENTS Some types are • Methanogens • Themoacidophiles • Extreme Halophiles Methanogens Live in ANAEROBIC Environments, • Produce Methane Gas, a byproduct of metabolism in conditions of very low oxygen • Includes Chemosynthetic Bacteria. • Lives in the Intestines of Mammals. • Methanobrevibacter smithii is the prominent methanogen in the human gut, where it helps digest polysaccharides (sugars). http://www.nature.com/nrgastro/journal/v8/n10/full/nrgastro.2011.159.html Thermoacidophiles - living in Sulfurous Hot Springs & Volcanic Vents They “love”“acid” & “heat” Extreme Halophiles Live in Very Salty Places (like the great Salt lake & the Dead Sea) (Halite is the mineral name for NaCl) Domain: Bacteria, Kingdom Eubacteria 2. “true” or Eubacteria • The first thing you probably think of when you say this word is “disease”something like strep. throat or maybe an infection in a cut. • Most of the bacteria that are disease causing are Eubacteria. • NOTE: Only about 1% of bacteria are disease causing. Domain: Bacteria, Kingdom Eubacteria • Many bacteria are very helpful organisms. • Actinomycetes, produce antibiotics such as streptomycin and nocardicin; • others live symbiotically in the guts of animals (including humans) or elsewhere in their bodies, or on the roots of certain plants, converting nitrogen into a usable form. • Bacteria put the tang in yogurt and the sour in sourdough bread; • help to break down dead organic matter; & make up the base of the food web in many environments. Domain: Bacteria, Kingdom Eubacteria “True” Bacteria: So remember- Most bacteria are beneficial • Benign (benign = good, friendly, kind) • Pathogens (means disease causing -only a few are “bad guys”) • Bacteria occur in 3 basic shapes: cocci, bacilli & spiral. Many are named by their shape. biology.clc.uc.edu Examples of common bacteria Lactobacillus acidophilus Streptococcus pneumoniae Escherichia coli answersingenesis.org http://www.oley.org/lifeline/Probiotics.html genome.microbio.uab.edu Domain: Eukarya, Kingdom Protista 3. Protists • Made of a variety of organisms that don’t “fit” anywhere else. (Some are not very much like the others in this group.) • EUKARYOTIC (has a true nucleus) • 50,000 species- many unicellular, some are like fungi, some like plants or animals. • Includes protozoans, unicellular algae, slime molds & water molds Examples of Protists includes slime molds & protozoans like Euglena, Paramecium, Ameoba Domain: Eukarya, Kingdom Fungi 4. Fungi • Can be Unicellular or multicellular • HETEROTROPHIC (eats something else) –NOT like plants (photosyntheic) (this is why they were kicked out of the plant kingdom) 100,000 species of mushrooms, puffballs, rusts, mildew & molds Agaricus bisporus: The Button Mushroom Fungi Examples: Stink horn Candida albicanscan cause yeast Infections- like this mouth thrush fcps.edu reference.medscape.com http://www.mushroomexpert.com/agaricus_bisporus.html Domain: Eukarya, Kingdom Plantae 5. Plantae • Multicellular & • Photosynthetic (Autotrophs) (They make their own food & are the chief food producers of the world.) • Found in all the types of environments: aquatic algae, amphibian mosses, and terrestrial ferns and seed-bearing plants. • 350,000 species identified. http://plantspages.com/typesofplants.htm Cladogram of the plant kingdom Domain: Eukarya, Kingdom Animalia 6. Animalia • Eukaryotic, • Multicellular • Heterotrophic • Most animals are symmetrical • Movement at some time in their life cycle. Examples of animals biology.kenyon.edu What is on the ch 17 test? • 1-6 Matching the 6 kingdoms or 3 domains with the correct description. • 7-40. Multiple choice. Review you notes. *Know the classification groups, the correct way to write binomial nomenclature & the morphology terms (homologous, analogous, vestigial structures) & phylogenic tree info. • 41-50. True/ False • 10-55. short answer. Practice these short answer questions for the test: • Write a pneumatic devise to remember the classification categories, in order (this can be one we used in class or your own): • What are the 6 kingdoms recognized today? • What do plants & fungi have in common with animals? *You can do online flash cards for the vocab in this chapter at: • http://quizlet.com/2096160/chapter-17classification-flash-cards/