Pancreas – enzymes that complete food digestion, ducted to duodenum via pancreatic ducts Main duct -> Hepatopancreatic ampulla Accessory pancreatic duct -> minor duodenal papilla, pancreatic juice when no bile Duct – bicarbonate mucus, acini secrete enzymes Figure 14.6 Trypsinogen, chymotrypsinogen, procarboxypeptidase Trypsinogen converted to Trypsin by enterokinase in lumen of duodenum Trypsin cleaves other two to make chymotrypsin and carboxypeptidase, and digests dietary protein Pancreatic amylase, pancreatic lipase, ribonuclease, deoxyribonuclease – active after exposure to bile and ions in lumen Bile and pancreatic juice secreted in response to vagus stimulation (parasympathetic), inhibited by sympathetic Both stimulated by Cholecystokinin (CCK), gastrin, and secretin CCK released by duodenum in response to fat and acid from stomach. Contraction of gallbladder – bile into duct Secretion of pancreatic enzymes Relaxation of hepatopancreatic sphincter Secretin – stimulates bile and pancreatic duct secretion of bicarbonate, neutralize acid 4 lobes – rt, lt, quadrate, caudate, sep by falciform ligament = mesentary Round ligament = remnant of umbilical vein, blod from umbilical cord to liver of fetus Portal hepatitis – entry of hepatic portal vein, proper hepatic artery, exit of bile passages Gallbladder associated Hepatic lobules – central veins, cuboidal hepatocytes Hepatic Triad – artery + vein (branches of proper hepatic artery and hepatic portal vein), bile ductule Hepatic sinusoid – fenestrated endothelium sep hepatocyte from bloodstream, blood plasma fills, blood directly from intestine Liver gets first dibs on glucose, AA, iron, vitamis, nutrients Removes hormones, toxins, bile pigments, drugs Secretes some blood proteins Bile canaliculi, bile ductules of triads, left and right hepatic ducts >common hepatic duct Hepatic and cystic duct join = bile duct Bile duct joins pancreatic duct = hepatopancreatic ampulla Enter duodenum=major duodenal papilla, heptopancreatic sphincter (sp of Oddi) Yellow-green color Watery solution contain bile salts and pigments, cholesterol, phospholipids, electrolytes Bilirubin – breakdown product of Hb Only bile salts and phospholipids aid digestion – emulsify fats, more SA for fat digesting enzymes to work on Thin walled green sac When not eating bile stored here Concentrated by water removal Fatty meal enters duodenum, bile released due to hormonal stimulus http://gogogojiteam.com/gallstone.html Bile stored for too long or too much water removed, cholesterol crystallizes Blockage of common hepatic or bile ducts prevent bile into small intestine, backs up into liver Bile pigments enter blood and circulate, yellowing jaundice MAJOR digestive organ 3 subdivision Duodenum Jejunum Ileum – meets large intestine at ileocecal valve Only able to process small amount of food at time, pyloric sphincter controls food movement Nearly all food absorption 3 structures in wall increase absorptive surface Microvilli – plasma membrane – brush border enzymes – digestion of protein and carbs Villi – mucosa, w/in each are capillaries and lacteal duct (fat), food absorbed, absorptive and goblet cells Circular folds – deep fold of mucosa and submucosa, don’t disappear as fill with food All decrease as move along Peyer’s Patches increase – collections of lymphatic tissue Bacteria in food! Figure 14.7 Absorptive cells line villi, have microvilli Contain brush border enzymes – enterokinase – remember pancreatic Trypsinogen Crypts of Lieberkuhn – glands at base of villi, extend to muscularis mucosa – absorptive, goblet cells, stem cells, Paneth cells (lysozyme, phospholipase, defensins) Brunner glands – submucosa – bicarbonate mucus (unique to duodenum) Peyer patches – lymphatic nodules - ileum Segmentation vs. Peristalsis Mix nutrients Churn chyme, contact with mucosa for contact digestion and absorption Move to large intestine Ileocecal valve to anus, appendix Absorb water and eliminate indigestible food as feces No villi Goblet cells in mucosa produce HCO-3 rich mucus, lubricant Divisions Cecum, Appendix, Colon, Rectum, Anal Canal Colon – ascending, transverse, descending, sigmoid Anus – internal and external sphincters, anal columns and sinuses Figure 14.8 Bacterial flora – ferment cellulose and other undigested nutrients Synthesize B vitamins, vitamin K which are absorbed by colon Don’t get enough vitamin K in diet Expel about 500ml of gas Some from swallowed air, some from bacterial digestion 12-24 from meal to feces Reabsorbs water and electrolytes Feces – 75% water and 25% solids Solids – 30% bacteria, 30% undigested fiber, 10-20% fat (not diet), epithelial cells, salts, mucus, etc Stretching of rectum stimulates defecation Intrinsic defecation reflex – stretch signals travel myenteric nerve plexus to muscularis of descending, sigmoid colon, rectum Stimulates peristaltic waves Internal sphincter relaxes – defecation if external also relaxed Need cooperation of parasympathetic defecation reflex – spinal cord Stretch of rectum, signal to sacral of spinal cord Return via parasympathetic fibers, intensifies peristalsis Parasympathetic fibers relax internal sphincter Food must enter the system to be acted upon Active and voluntary Figure 14.11 Foods must be processed my multiple organs – move from one to next Swallowing is example Movement depends mostly on peristalsis vs. segmentation Segmentation – small intestine, alternating segments contract, mechanical digestion Mixing of food in mouth by tongue Churning of food in stomach Segmentation in small intestine Prepare food for enzymatic digestion by breaking into smaller pieces Food molecules broken down into building blocks by enzymes Reactions called hydrolysis rxn, water added as bond breaks Carbs (saccharides) we digest – sucrose, lactose, maltose, and starch. Eat cellulose but can’t digest, fiber Proteins (a.a.) – polypeptides or peptides Lipids (fats) – fatty acids and glycerol Transport of digested food into blood from lumen of GI tract Must enter mucosal cells by active or passive transport. Small intestine major site of absorption Elimination of indigestible food from GI tract via anus in form of feces Figure 14.13 (1 of 3 Figure 14.13 (2 of 3)) Figure 14.13 (3 of 3)