Plantae Seed Plants Vascular Plants Formation of vascular tissue – Xylem (water) – Phloem (food) – True leaves, roots, and stems Lignin Sporophyte generation dominate Alternation of Generation Alternation of Generation Sporophyte dependent on gametophyte – mosses Large sporophyte and small independent gametophyte – ferns Gametophyte dependent on sporophyte – seed plants Why be Sporophyte Dominant? Reduced mutations – UV light harmful to DNA – Diploid (2n) form copes better with mutations two alleles Why Retain Gametophyte Generation? Ability to screen alleles – doesn’t require a large amount of energy Sporophyte embryos rely on some gametophyte tissue Seeds A seed is a sporophyte in a package – spores are only single cells – packaged with food All seed plants are heterosporous – megasporangia – microsporangia From Ovule to Seed Overview of Seed Plants Produce Seeds – Can remain dormant for years – Pollination replaces swimming sperm Gametophyte generation reduced – Gymnosperms lack antheridium – Angiosperms lack both archegonium and antheridium Phylogeny Gymnosperms (Naked Seed) Division: Division: Division: Division: Cycadophyta Ginkgophyta Gnetophyta Coniferophyta Ginkgophyta Ginkgo or Maidenhair Tree Characteristic leaves Only one species Only males are planted Cycadophyta Cycads Palm-like plants – Sago Palms Leaves in cluster at top of trunks True Secondary growth Gnetophyta 3 Genera Ephedra Mormon Tea – Ephedrine raises heart rate raises blood pressure Coniferophyta Coniferophyta Pine tree is the sporophyte generation Contains both male and female cones – Pollen (staminate) cones (low in tree) produces pollen – Ovulate cones (high in tree) with scales produces seeds Pine Life Cycle No Antheridium (microsporangia) produce pollen grain (4 cells) –2 prothallial cells –1 generative cell • become a sterile cell and a spermatogenous cell • produces 2 sperm –1 tube cell –wings for dispersal Pine Life Cycle Ovule in a ovulate cone – integument (seed coat) (2n) – megasporangia (meiosis) or nucellus (nutrition) (2n) – produces 4 megaspores (3 die) develops into female gametophyte –archegonium with eggs (n) Angiosperms Angiosperm Flower Sepals Petals Receptacle Stamen – Anther – Filament Carpel – Stigma – Style – Ovary with ovule Angiosperm Life Cycle Angiosperm Life Cycle No Antheridium (microsporangia) produce pollen grain –1 generative cell • produces 2 sperm –1 tube cell Angiosperm Life Cycle Ovule in Ovary – megasporangia – produces 4 megaspores (3 die) develops into female gametophyte called the embryo sac Angiosperm Life Cycle Inside embryo sac – 7 cells (eight nuclei) due to 3 mitotic divisions 3 antipodals 2 polar nuclei (one cell) 2 synergids 1 egg Angiosperm Life Cycle Double fertilization – one sperm unites with egg – one sperm unites with polar nuclei develops into endosperm (3n) Fruit and Seed development – ovule = seed – ovary = fruit Cross Pollination Most flowers do not self-pollinate – stamen and carpal may develop at different times – stamen and carpal may be arranged in flower to avoid contact Angiosperm Radiation Begins the Cenozoic era (65 mya) Most closely related to the Gnetophyta Coevolution – the mutual influence of two species on each other – plants and animals (insects, birds, bats) Pollination Dispersal The Global Impact Transformed atmosphere – reduced carbon dioxide – cooled the earth Nonrenewable resource