Plant Science Introduction - Basic Plant Structure Assessment Statement 9.1.1 9.1.2 9.1.3 9.1.4 9.1.5 9.1.6 9.1.7 Draw and label plan diagrams to show the distribution of tissues in the stem and leaf of a dicotyledonous plant Outline three differences between the structures of dicotyledonous and monocotyledonous plants Explain the relationship between the distribution of tissues in the leaf and the functions of these tissues Identify modifications of roots, stems, and leaves for different functions: bulbs, tubers, storage roots and tendrils State that dicotyledonous plants have apical and lateral meristems Compare the growth due to apical and lateral meristems in dicotyledonous plants Explain the role of auxin in phototropism as an example of the control of plant growth Green plants make up one of the five kingdoms of living things. The kingdom is Plantae. Plants are different than other organisms because: Each has a cell wall around each cell made of cellulose, a polysaccharide, which is an extremely tough and protective material. Cell organelles called chloroplasts are the site of photosynthesis, which the plant takes carbon dioxide and water and produces organic molecules (carbohydrate) and oxygen. It is thought that plants evolved from aquatic, single-celled organisms called green algae. Today, the dominant terrestrial plants are the angiospermophytes (flowering plants). Fossil records indicate that the angiosperms took over about 100 mya. Angiosperms, or flowering plants, produce seeds enclosed inside fruits. Angiosperm comes from the word angerion – a container and sperma – a seed and phyton – a plant. Angiosperms are divided into two large groups: 1. Monocotyledons – include palms, gingers, lilies, irises, grasses, orchids, palms, bananas, and others 2. Dicotyledons – include most trees and shrubs, and many non-woody plants. These names refer to the number of leaves contained in the embryo, called cotyledons. The differences between the two are listed below: Number of Cotyledons Numbers of Organs in the Flowers Meristems Leaf Attachment Method of Formation of new roots Vascular Tissue in the Stem Leaf Veins Monocots 1 3 or 6 Dicots 2 4 or 5 Apical only – stems cannot grow wider Around the whole stem circumference By formation of roots from the stem Vascular Bundles spread throughout Parallel Apical and lateral so stems can widen To one side of the stem by a leaf stalk By branching from other roots Branching to form a network Branching to form a network Draw the typical forms of each. Basic Plant Structure and Growth Whether the plant (angiosperm) is woody or not (herbaceous), the plant consists of a stem, leaves and root. Plants retain groups of stem cells throughout their lives, allowing them to continue to grow indefinitely. These groups of cells are called ________. Cells in meristems are small and go through the cell cycle repeatedly to produce more cells, by mitosis and cytokinesis. These new cells absorb nutrients and water and increase in volume and mass. Primary meristems are found at the tips of stems and roots. They are called apical meristems. The root apical meristem is responsible for the growth of the root. The shoot apical meristem, at the tip of the stem, is more complex. It throws off the cells that are needed for the growth of the stem and also produces the groups of cells that grow and develop into leaves. Meristems Plants grow throughout their lifetime. This pattern of growth is called indeterminate. Plants, of course, will die, based upon their life cycle. Annuals complete their life cycle in one year, whereas perennials live many years and usually die due to a disease or environmental factor. Indeterminate growth is due to the meristematic tissue. In dicots, there are apical and lateral, which have been mentioned before. Apical Meristems Sometimes referred to as primary meristems, these occur at the tips of roots and stems. Primary tissue is produced and causes primary growth. In the root this allows the structure to extend through the soil. In the stem, it allows the stem to grow longer and increases the exposure to light and carbon dioxide. This type of growth results in herbaceous, non-woody stems and roots. Lateral Meristems Lateral meristems allow growth in thickness of plants. This is referred to as secondary growth. Most trees and shrubs have active lateral meristems. These plants have two types of lateral meristems: Vascular Cambium produces secondary vascular tissue. It lies between the xylem and phloem in the vascular bundles. On the inside is produces secondary xylem, which is a major component of wood. One the outside it produces secondary phloem. Cork cambium occurs within the bark of a plant and produces the cork cells of the outer bark. Make a table to compare Apical and Lateral Meristems Stems The stem supports the leaves in the sunlight, and transports organic materials (such as sugar and amino acids), ions and water between the roots and leaves. As the plant grows, the stem has to grow larger. Terrestrial plants support themselves: With turgid cells, which are almost rigid because of their high pressure With cells that have thickened cellulose walls With xylem tissue, which has cells walls impregnated with lignin, making it woody and hard. The stem is surrounded or contained by a layer called the epidermis. Inside is vascular tissue, which is composed of a system of veins or vascular bundles. The bundles are made of xylem that transports water and phloem that transports organic solutes. In the stem, the vascular bundles are arranged in a ring, towards the outside of the stem. The vascular the tissue extends into the leaf with some specialized leaf tissues. Besides the apical meristems at the tips, the stems of dicots can grow wider and the plant grows taller. This is done by a lateral meristem, which develops in a complete circle around the stem. The lateral meristem develops between two tissues called xylem and phloem, and the cells produced develop into both of these tissues. Xylem develops on the inner side of the lateral meristem, gradually adding to the width of the stem. The plants that produce the largest amount of xylem can grow the tallest and compete for light the most. Naturally, we refer to these as trees. Monocots have stems with an apical meristem at the tip of the stem, but the tissues of the stem differentiate in a different arrangement from those of dicots. Draw the Plan Diagram of the Stem Leaf Structure A leaf consists of a leaf blade connected to the stem by a leaf stalk. It grows from the apical meristem at the tips of the stem. Leaves must have a large surface area and a small space between the upper and lower surfaces. The leaf is an organ specialized for photosynthesis. The leaf is composed of: 1. The outer structure - Epidermis It is a tough, transparent layer. The upper epidermis has an external waxy cuticle on its surface. The cuticle is impervious and effectively reduces water vapour loss from the leaf surface. The lower epidermis has a thinner cuticle, with specialized epidermal cells called guard cells and pores called stomata (stoma). They are the sites of inward diffusion of carbon dioxide. Within the inner structures are air spaces that enhance the diffusion of carbon dioxide to the cells. 2. Inner part of the Leaf The inner part of the cell consists of mesophyll cells that are spread out over a wide area, so the amount of light absorbed is maximized. In the upper layer, the mesophyll cells are called palisade mesophyll cells and are tightly packed close to the upper surface of the epidermis. They are packed with chloroplasts. In the lower half, there are spongy mesophyll cells. These are loosely packed in the lower portion of the leaf, with large air spaces in between and fewer chloroplasts. Between the mesophyll cells lies a network of vascular bundles or the veins. The vascular bundles contain xylem and phloem. This brings water (xylem) to the cells and transports the sugars made (phloem). The network of bundles has another function; to support the leaf. Mesophyll tissue is supported by the turgidity of all its cells, contained in the non-elastic epidermis and reinforced by the network of bundles. Draw the Plan Diagram of a Leaf Root Systems One of the earliest stages in the development of an embryo plant is the formation of a root. When seeds germinate, the embryonic root grows out of the seed coat and downwards into the soil. The root anchors the plant and is the site of absorption of water and ions from the soil. These roots go deep into the soil, and are called tap roots. When the roots branch out to the side, to search for water outside their canopy, these are called lateral roots. The seedling root develops branches in some plants and hairs, which absorb more water in dry soil. The branching helps with absorption by increasing the surface area. The water flows in by osmosis. Roots have an epidermis that forms a protective outer layer. The inner layer is called the cortex. This is used for conducting water and can be modified to store nutrients. There is an endodermis that surrounds the vascular layer that contains xylem and phloem. Draw Patterns of Root Branching and tissue plan diagram Modifications and Adaptations of roots, stems and leaves. The structures of the root, leaf and stem are very similar, with some differences. As the principle plant organs, they are used to build the structures that they need for water and mineral absorption, support and photosynthesis. Other functions can be carried out if the principle structures are modified. Roots Roots can be modified, based upon the needs of the plant. Besides tap roots and lateral (fibrous) roots there are: Prop Roots – thick adventitious roots that grow from the lower part of the stem and brace the plant. Ex. Corn Storage Roots – specialized cells within the roots store large quantities of carbohydrates and water. Ex. Carrots and Beets. Pneumatophores – produce by plants that live in wet places, these roots extend above the soil or water surface and facilitate oxygen uptake. Ex. Mangroves and cypress trees. Buttress Roots – large roots that develop near the bottom of trees to provide stability. Ex. Fig Stems Stems are capable of great modification. Bulbs – vertical, underground stems consisting of enlarged bases of leaves that store food. Ex. Onions Tubers – horizontally growing stems below ground that are modified as carbohydrate- storage structures. Ex. Potatoes Rhizomes – horizontal stems that grow just below the surface to allow the plant to spread. Ex. Ginger plant Stolons – horizontal stems growing above the ground that allow a plant to reproduce asexually. Ex. Strawberry plant Leaves They still function as photosynthesizing factories, but can adapt. Tendrils – structures that coil around objects to aid in support and climbing. Ex. Pea plants Reproductive Leaves – produce tiny plants along the leaf margins that fall to the ground and take root in the soil. Ex. Kalanchoe plants Bracts or floral leaves – coloured modified leaves that surround flowers and attract insects for pollination. Ex. Poinsettia Spines – reduce water loss, may be associated with modified stems that carry out photosynthesis. Ex. Cacti Know at least one example of each. Growth and Phototropism Plants use hormones to control the growth of stems and roots. Rate and direction are controlled. Both light and gravity influence growth. In light they grow towards the brightest source and in the dark, grow in the opposite direction to gravity. The responses are called tropisms. The growth towards light is called phototropism. The steps of phototropism are: Photoreceptors (proteins called phototropins) absorb light. Light of an appropriate wavelength is absorbed which stimulates the photoreceptors to bind in the cells and transcribe glycoproteins which will aid in the transport of a plant hormone where it is needed. The stimulus of the light will also transcribe a protein hormone to be produced called auxin. Auxin does many things in the plant: o Auxin synthesized in the tips of growing stems and transported down to stimulate growth. o It promotes the elongation of cells in stems, by loosening the connections between the cell walls and cellulose microfibrils o If phototropins detect a greater light intensity on one side of the stem than the other, auxin is transported to the shadier side. This promotes the stem to grow more on the shadier side and go towards the light. This allows the leaves on the sunny side to get more light and photosynthesize at a greater rate. Do the worksheet on Phototropism Transport in Angiosperms Assessment Statement 9.2.1 9.2.2 9.2.3 9.2.4 9.2.5 9.2.6 9.2.7 9.2.8 9.2.9 9.2.10 9.2.11 Outline how the root system provides a large surface area for mineral ion and water uptake by means of branching and root hairs List ways in which mineral ions in the soil move to the root Explain the process of mineral ion absorption from the soil into rots by active transport State that terrestrial plants support themselves by means of thickened cellulose, cell tugor and lignified xylem Define transpiration Explain how water is carried by the transpiration stream, including the structure of xylem vessels, transpiration pull, cohesion, adhesion, and evaporation State that guard cell can regulate transpiration by opening and closing of stomata State that the plant hormone abscisic acid causes the closing of stomata Explain how the abiotic factors light, temperature, wind and humidity, affect the rate of transpiration in a typical terrestrial plant Outline four adaptations of xerophytes that help to reduce transpiration Outline the role of phloem in active translocation of sugars (sucrose) and amino acids from source (photosynthetic tissue and storage organs) to sink (fruit, seeds, roots) Root system, absorption and uptake The roots provide a huge surface area to draw up essential ions and water. Contact with the soil is vastly increased by root hairs that occur just behind the growing tip of each root. The water flows in due to osmosis. The uptake of minerals is done via active transport. 1. Water uptake Water uptake occurs from the soil in contact with the root hairs by osmosis. Uptake is largely by mass flow through the interconnecting free spaces in the cellulose cell walls to the xylem. There are three possible routes of water movement through the plant cells and tissues. The cortex structure of the root also facilitates the water uptake. Apoplast Pathway (Mass Flow) Water does not enter the cell. It moves through the cell walls until it reached the endodermis. Cells of the endodermis have a Casparian Strip around them that is impermeable to water. The water is diverted to the cytoplasm of cells (momentarily), and then eventually to the xylem. The Casparian Strip is thought to be a measure of control so water cannot move directly into the xylem, so the cell can divert it out of the cell, if it is contaminated. Symplast Pathway o Water enters the cytoplasm but not the vacuole. It passes from cell to cell via connections between cellular cytoplasm of adjacent cells, called plasmodesmata. The organelles are packed together in cells, and as a result, block significant progress of water. It is not the major pathway for water. Minerals mainly move through this pathway. Vacuolar Pathway o Water enters the cell and move into the vacuole. It then travels through the cytoplasm and the cell wall to the next cell. Once in the endodermis, water can move into the xylem and pulled via transpiration forces. 2. Uptake of Minerals There are three ways that minerals can move from the soil to the root. 1. 2. 3. Active Transport using protein pumps in the plasma membranes of root cells. Mass Flow, or the water carrying the ions goes into the root. Fungal hyphae – fungus grows on the surface of roots and sometimes into the cells of the roots. The hyphae grow into the soil and absorb minerals ions and phosphate from the surface of soil particles. The ions are supplied to the roots and allow the plant to grow in mineral poor soil. Some plants supply sugars, and other nutrients to the fungus. This is an example of a mutualistic relationship. Transpiration Find the definition of Transpiration Transport up the stem Water is taken up the stem, from the roots by the Mass Flow or Apoplast Pathway, into the xylem. In the root, the xylem is centrally located, in the stem, the xylem occurs in the ring of vascular tissue. Xylem, therefore runs from the roots to stem and then to leaf. Xylem begins as elongated cells with cellulose walls and living contents, connected end to end. During development, the end walls dissolve away, and the mature xylem is a long hollow tube. As the tubes extend, the tissue dies. Therefore, xylem is dead tissue, and is composed of two elements: Tracheids – narrow cells arranged in columns, overlapping at tapered ends giving some support to the plant. The ends have pits for water to move rapidly from one cell to the next. Due to their structure, they are less efficient that xylem vessels. Xylem vessels – larger columns of cells. When the dead cell walls disappear, they become wider and transport water more efficiently. Since they are so wide, they are reinforced by ligin for support of the plant. This only occurs in angiosperms. Mechanism of water transport Transpiration controls the flow of water. But, how does water move against gravity? First, in the leaf are stomata. These open and close to control the amount of water present in the leaf. Around each stomata are guard cells, which open and close, depending on the tugor of the cells. When the plant is well hydrated, the guard cells are swollen, causing them to open, due to the pressure on the cells walls. When the plant dries out, the guard cells sag and the stomata close. Water loss is stopped and gas exchange is halted. Other external factors that affect the opening and closing of stomata are: Light causes stomata to open Low CO2 levels in the air spaces in the cause the stomata to open Shortage of water causes the stomata to close When leaves are deficient in water, they synthesize a hormone called abscisic acid. This closes the stomata and overrides any external stimuli – the stomata close. This allows the plant to avoid dehydration and death. When the stomata are open, water evaporates out of the leaves, diffusing water vapour out. This maintains a concentration gradient that requires more water. The water in the stem, which is connected by xylem, moves up to replace the water lost by transpiration. As a result, the water is pulled up the plant. Water is polar and is held together by its cohesive forces. Therefore, the water molecules stick together and flow up the stem together. This is called transpiration pull. Factors affecting Transpiration The rate of transpiration is the amount of water vapour that a plant loses from its leaves and stems per unit of time. The rate depends on the: Size of the plant The thickness of the cuticle How widely spaced the stomata are Whether the stomata are open or closed These are the biotic factors. These are the factors the plant can control, based upon the species of plant. There are four abiotic factors that affect the rate of transpiration. 1. Temperature – affects the rate at which water evaporates from the surfaces inside the leaf. At higher temperatures, evaporation increases. The higher temperatures also increase the rate of diffusion between the inside of the leaf and the outside. The increase in temperature reduces the air’s ability to hold more moisture and reduces the relative humidity of air outside the leaf. The concentration gradient increases, doubling the rate for every 10oC increase in temperature 2. Humidity – is the water vapour content of the air. It is measured as a percentage of the maximum amount of water vapour the air can hold. The humidity inside the leaf is always around 100%. The lower humidity outside the leaf causes the evaporation to be higher, increasing transpiration. Evaporation is much higher in dry air, so on hot / cold, dry days, transpiration increases dramatically 3. Wind – air currents take water vapour away from the leaf surface, keeping the concentration gradient large, and increasing the rate of transpiration. On calm days, the humidity around the leaf increases, slowing down transpiration. 4. Light – in light, photosynthesis increases and stoma open to allow CO2 in, but also water out. Therefore, transpiration increases. In darkness, there is no need to absorb CO2, and water can be conserved, as the stoma close and photosynthesis decreases. What else does water do? The take up of water, allows plants to grow very tall. Plants do not have a skeleton. Woody trees and shrubs have xylem to support them. Herbaceous plants have xylem, but depend on water to provide turgor pressure. Turgor Pressure is the swelling of cells with water. As the water is absorbed into the cells, from the xylem, the vacuole takes in water. As a result, the vacuole swells until its membrane presses against the cell wall. The pressure provides that stiffness, or crispness, that we see in vegetables, like celery. Moving Food in Plants Find the definition of Translocation is the movement of manufactured food (sugars and amino acids). This occurs in the phloem tissues of the vascular bundles. Sugars are made in the leaves (in the light) by photosynthesis and transported as sucrose. The first formed leaves transport sugars to sites of new growth (new stems, new leaves, and new roots). In older plants, sucrose is increasingly transported to sites or storage, such as the cortex of roots or stems, and in seeds and fruits. Amino acids are mostly made in the root tips. Here is where the absorption of nitrates takes place. After they are made, amino acids are transported to sites where protein synthesis is occurring. These are mostly in buds, young leaves and young roots, and in developing fruits. Translocation is not just limited to organic compounds. Chemicals that are applied to plants by spraying, and are then absorbed by the leaves may be carried all over the organism. Pesticides are called systemic for this reason. Phloem tissue consists of sieve tubes and companion cells. Sieve tubes are narrow, elongated elements connected end to end to form tubes. The end walls, know as sieve plates, are perforated by pores. The cytoplasm of mature sieve tubes has no nuclei, or many of the other organelles in a cell. But each sieve tube is connected to a companion cell by strands of cytoplasm passing through gaps (called pits) in the walls. The companion cells are believed to service and maintain the cytoplasm of the sieve tube, which has lost its nucleus. Phloem is a living tissue, and has a relatively high rate of aerobic respiration during transport. In fact, transport of manufactured food in the phloem is an active process, using energy from metabolism. Phloem transport may occur in either direction in stem leaves and roots, and is believed to move by mass flow. How this works is: Solutes are loaded into the phloem sieve tubes, requiring ATP and then the solutes flow through the phloem from a region of high hydrostatic pressure to low hydrostatic pressure. Hydrostatic pressure is high around photosynthetic cells in the light (mesophyll of the leaf), and in the phloem sieve tubes nearby. It is the presence of the sugars, which concentrate the fluid and creates a high osmotic pressure. Water flows in, raising the hydrostatic pressure further. This is called a source area. Hydrostatic pressure is low in cells where sugar is converted to starch and stored. Areas of storage are the cortex of the root, stem, seeds and in the nearby phloem tissue. Here the removal of sugars lowers the osmotic pressure, and water flows away. These storage areas are called sink areas. Below summarizes the sources (areas where sugars and amino acids are loaded into the phloem) and sinks (where the sugars and amino acids are unloaded and used). Sources Photosynthetic tissues: Mature green leaves Green stems Storage organs that are unloading their stores: Storage tissues in germinating seeds Tap roots or tubers at the start of the growth season Sinks Roots that are growing or absorbing mineral ions using energy from cell respiration Parts of the plant that are growing or developing food stores: Developing fruit Developing seeds Growing leaves Sometimes sinks turn into sources and visa versa, and therefore, phloem must be able to transport in both directions. Unlike the vessels in animals, there are no valves or a central pump. However, they are similar because in both, fluid flows inside tubes due to pressure gradients. Energy is needed for both, and therefore, both are active processes. The movement of substances in phloem is called active translocation for this reason. Adaptations of xerophytes Xerophytes are plants that have adapted to arid climates. Examples are cacti are an example. In order to adapt to dry climates, xerophytes must decrease water loss due to transpiration. Therefore, the plants have adapted by: Small, thick leaves reducing the water loss by deceasing surface area (needles or green stems) Reducing the number of stomata Having the stomata located in crypts or pits on the leaf surface, which causes higher humidity near the stomata Having a thickened, waxy cuticle Having hair-like cells on the surface to trap water vapour Becoming dormant in the dry months Storing water in the fleshy stems and restore the water in the rainy season Using alternative photosynthetic processes called CAM photosynthesis (Crassulacean acid metabolism) and C4 photosynthesis. CAM plants close stomata during the day and incorporate carbon dioxide at night. C4 plants have stomata open during the day, but take in carbon dioxide more rapidly than nonspecialized plants. Reproduction in Flowering Plants Assessment Statement 9.3.1 9.3.2 9.3.3 9.3.4 9.3.5 9.3.6 Draw and label a diagram showing the structure of a dicotyledonous animal pollinated flower Distinguish between pollination, fertilization, and seed dispersal Draw and label a diagram showing the external and internal structure of a named dicotyledonous seed Explain the conditions needed for the germination of a typical dicotyledonous seed Outline the metabolic processes during the germination of a starchy seed Explain how flowering is controlled in long-day and short-day plants, including the role of phytochrome Flowering plants contain their reproductive organs in the flower. Flowers are often hermaphrodite structures, carrying both male and female parts. The parts of flowers occur in rings or whorls, attached to the tip of the flower stalk, called the receptacle. Find the definition of the following parts of a typical dicotyledonous flower Sepal (together with the receptacle, called the calyx) – Petals (together called the corolla) – Stamen – male part of the flower, which consists of: Anthers – Filament or stalk – Carpels – female part of the flower, and they may be on their own or fused together. Each carpel consists or: Ovary – Stigma – Connecting style – Draw the Structure of a Dicotyledonous Flower with annotated labels Pollination and Fertilization Find the definition of Pollination The pollen may come from the anthers or the same flower or flowers of the same plant, which is called self-pollination. When pollen comes from flowers on a different plant of the same species, which is transferred, is called cross-pollination. Transfer of pollen is usually by insects, wind, birds or bats. Flowers that attract insects usually produce nectar. Find the definition of Fertilization The pollen produces a tube, which grows down between the cells of the style, and through the ovule. The pollen tube delivers two male nuclei. One of these male nuclei then fuses with the egg nucleus in the embryo sac, forming a diploid zygote. The other fuses with the other nucleus, which triggers formation of the food store for the developing embryo. Seed formation and dispersal The seed develops from the fertilized ovule and contains and embryo plant and a food store. After fertilization: The zygote grows by mitosis. The cells form the embryonic plant, consisting of an embryo root and stem. A seed leaf or cotyledon forms. The seed leaf has two forms, as angiosperms have two classes. o Monocotyledons – have a single seed leaf o Dicotyledons – have two seed leaves The formation of stored food reserves is triggered. In many seeds the food store is absorbed into the cotyledons. As the seed matures, the outer layers of the ovule become the protective seed coat, or testa. The micropyle is a small hole through the testa, where it was attached to the parent plant. The whole ovary develops into the fruit. The water content decreases and the seed moves into a dormancy period. Water only makes up 10 –15% or the seed weight. Dispersal of the seed is needed to make sure there are not many seeds close together, as they will compete for resources. Therefore they travel long distances from then parent plant. The type of seed dispersal depends on the structure of the fruit; dry and explosive, fleshy and attractive for animals to eat, leathery and winged to catch the wind, or covered in hooks to catch the coats of animals. Draw the External and Internal Structure of a Bean Seed Seed Germination Seeds are in suspended animation and there is very little metabolic activity occurring. When the metabolic activity starts, this is the start of germination. The seed stays dormant due to: 1. 2. 3. Incomplete seed development Presence of a plant growth regulator – abscisic acid Impervious seed coat The conditions that are essential for germination are: 1. 2. 3. Water – hydrates plant and activates amylase and removes the abscisic acid Oxygen – for Cellular respiration Period of warm temperatures as this is important for enzyme production. Outline the metabolic processes during the germination of a seed. The seedling develops and functional leaves appear. Photosynthesis takes over and becomes less dependent on the maltose. The seed will eventually become a mature plant and produce seeds of its own, starting the whole process over again. Control of Flowering in Angiosperms Light is required for photosynthesis and controls aspects of growth and development. Plants are able to detect the presence of light, its direction, wavelength and intensity. Photoperiodism is the plant’s response to light involving the lengths of day and night. It is the length of day and night that controls flowers. When the plant is young and structures, such as leaves, stem and roots grow, this is called the vegetative phase. When the plant is ready to reproduce, this is called the reproductive phase and flowers form. As mentioned before, flowers are very important, as they attract pollinators. There are three categories of plant in relation to light and flowering. Plant Type Long-day plants Short-day plants Day-neutral plants Flowering and Light Bloom when days are longest and nights are shortest (midsummer) Bloom in spring, late summer and autumn when days are shorter Flower without regard to day length Examples Radishes, spinach and lettuce Poinsettias, chrysanthemums and asters Roses, dandelions, and tomatoes It is actually the length of night that controls the flowering process in the long-day and short-day types. The control by light is brought about by a special blue-green pigment called phytochrome. Phytochrome is a large protein that is not a plant growth hormone, but a photoreceptor pigment. There are two forms of phytochrome: inactive form Pr active form Pfr When red light (600 nm) is present in available light, the Pr is converted to Pfr. It is done rapidly. The active phytochrome can absorb far-red light (730 nm). In darkness, the active form (Pfr) slowly converts back to Pr. The slow conversion allows the plant to time the dark period and controls the flowering in short-day and long-day plants. What does this all mean? In long-day plants, the remaining Pfr at the end of a short night, stimulates the plant to flower. It acts as a promoter in these plants. The long day, long period of daylight, causes the accumulation of Pfr. In short-day plants, enough Pfr acts as an inhibitor. It has been converted to Pr, to allow flowering to occur. In other words, the very long nights required by short-day plants allow the concentration of Pfr to fall to a low level, removing the inhibition.