Skokholm Island Annual Report 2012 Jerry Gillham and Lewis Yates 1 GENERAL INTRODUCTION AND SUMMARY 3 1 5 RECORDING 1.1 FLORA: HABITATS AND COMMUNITIES 1.3 MAMMALS 1.4 BIRDS: INDIVIDUAL SPECIES 1.4.1 BIRDS: ASSEMBLAGES 1.4.2 OTHER BIRDS 1.5 HERPTILES 1.6 INVERTEBRATES 6 14 17 30 36 38 38 2.0 HUMAN IMPACT 39 2.1 VISITOR NUMBERS 2.2 DISTURBANCE TO WILDLIFE 2.3 POLLUTION 39 39 40 3.0 RESERVE ADMINISTRATION 41 3.1 FINANCE 3.2 STAFF 3.3 VOLUNTEERS 3.4 TRAINING 3.5 HEALTH AND SAFETY 3.6 MANAGEMENT PLANNING 3.7 TRUST NEW MEMBER RECRUITING 3.8 FRIENDS OF SKOKHOLM AND SKOMER 41 41 42 42 43 44 45 45 4. 0 RESERVE MANAGEMENT 46 4.1 MEDIA AND PUBLICITY 4.2 LIAISON AND RELATIONS 4.3 HABITAT AND SPECIES MANAGEMENT 4.4 INFRASTRUCTURE 4.5 MACHINERY AND EQUIPMENT 4.6 FOOD DELIVERIES 46 46 46 47 48 49 5.0 ACKNOWLEDGEMENTS 50 6.0 REFERENCES 50 2 General Introduction and Summary 2012 was the second year of accepting guests since being reopened and saw visitors staying in the middle block as well as in the recently completed cottage. Work parties were present in spring and autumn once more and much progress was seen. Core monitoring was carried out, as it has been throughout the years of closure, and a range of other monitoring was able to be completed including some vegetation and invertebrate surveys. Breeding bird surveys could be carried out as well as better recording of migrants. Core management for the islands is still handled from Skomer by warden Chris Taylor, while day-to-day management on Skokholm was carried out by a Skokholm Warden. This year saw the departure of Jerry Gillham with the remainder of the season being covered by William Whittington and Lewis Yates who both first came to the island as long term volunteers. Overall the wardens were in place on the island from 12th March until 26th October. Alongside this there were four long term volunteers who maintained continuity during the changeover of wardening staff. Office work, administration and paperwork have been kept on Skokholm again, although a backup of computer files will be kept on the Skomer external hard drive. Scanning important documents continued as conditions on Skokholm are not ideal for long term paper storage. Breeding Bird Overview Manx Shearwater, Storm Petrel and Lesser Black-backed Gull populations are all still below the lower limits of acceptable change (LAC) set out in the management plan. The majority of species on Skokholm seem to have steady populations. Some of these; Manx Shearwater (in study plots), Lesser Black-backed Gull and Herring Gull, are low numbers compared to recent peaks in the population. Others, including Puffin, Fulmar and Oystercatcher, are as expected for a stable population. Storm Petrel monitoring sites and methods established in 2010 were found to be repeatable and gave a similar figure. Renovation Work Restoration work was carried out for four weeks through April then all through August and September. The majority of the work was performed by volunteers, with supervision and specialist jobs carried out by professionals. Spring work concentrated on getting the cottage habitable, following the replacement of the roof and most walls last autumn. The floor was pulled up and approximately eight tons of earth was dug out. Concrete pillars and some beams were put in before the new floors were placed in the common room and angel loft. Work on the walls was finished, including several layer of lime-putty wash and there was loads of painting inside as well 3 as putting in all the furniture and furnishings. Electric lights were placed in the majority of the rooms. The kitchen was re-arranged to make it more suitable for more people, although the weather meant we were unable to replace the roof as planned. Work in the autumn continued this progress with the kitchen and Officer’s Mess being reroofed alongside the new garage roof, which had collapsed during the previous winter. The new water system was also started with two large tanks being constructed from a modular kit as well as a small, preformed header tank being put in place. These were connected with pipework, with the ram pump being relocated nearer South Haven, but the pipes were not all buried. A new ringing shed was constructed in the main courtyard as the old paint shed was demolished with its contents being relocated to the garage. 3 heligoland traps were constructed by the Wheelhouse, in the cottage garden and by the well pond. These traps need finishing off in the spring with small amounts of meshing to be completed alongside the catching boxes. 4 1 Recording This year the recording of the island’s wildlife was more comprehensive and systematic than last, but there were still limitations from the small number of staff and volunteers and differing expertise. More work was possible on projects such as common bird census, moth trapping and some vegetation surveys and the main bird counts were all carried out including the Spring Puffin count, all Gull counts, Shearwater plots, cliffnesting birds plots, whole island cliff-nesting bird counts, Gull breeding success and Fulmar productivity. Prioritising these tasks was still somewhat arbitrary, relying more on the availability of staff and volunteers with time and expertise, than a drawn-up plan of action. It is hoped that just such a ‘plan of action’ can be created over the next year, identifying other key monitoring areas so large gaps in recording can be avoided. The possibilities of having field courses return to Skokholm are being discussed for the near future. These courses could prove their worth in more ways than one if the students attending carried out some of the smaller projects as part of their learning. The limited diversity of island flora and fauna for most taxa means that misidentification would be limited and if necessary small sections of data could be assessed by staff to check accuracy (however if courses are led by knowledgeable individuals known to the island then even this should prove unnecessary). This kind of cooperation could improve field courses by giving students purpose and a better level of satisfaction whilst helping the island by relieving time pressures on staff and volunteers (and still bringing in new investment and visitors to the island). The management plan identifies targets and limits of acceptable change for the ‘features’ of the reserve, this section indicates whether the features are currently in a favorable condition, and the condition of the operational limits for various factors that are likely to affect the feature. Reading this section along with a copy of the management plan will make it easier to understand, the feature number has been added to enable this. Note that not all features and attributes are checked annually. Each suite of features is detailed in a table with the 2012 data and the state of the operational limits, followed by a summary table of available data from the last five years. Following these two tables are some notes regarding their content and other observations of interest. 5 1.1 Flora: Habitats and communities Feature/mixture component habitat, and attributes Checked in 2012? Within Limits of Acceptable Change? Operational limits (factors affecting feature) Notes on operational limits Coastal grassland (feature 1) Extent of coastal grassland Extent of Thrift % Thrift in transect Extent Silene dominated turf Extent of other maritime grassland dominated communities % Thrift tussocks alive in transect quadrats % Vegetative cover (November) Soil erosion (cm) Coastal grassland transects carried out - Unknown Unknown: Data collected but awaiting analysis Bracken distribution Exceeded 1989 limits by 2000, significant expansion since then. Rabbit density Regular counts of above 50/Ha are above LAC Bare earth footpaths Very few bare earth footpaths, none exceeding 1m width. Golden rod extent Now covers Home Meadow and is becoming widespread in the middle of the island. No unauthorized access off footpaths No instances of unauthorized access on cliffs No person to leave paths, nor to access shore from boats No instances of such behaviour - Maritime cliff and associated crevice habitat and communities. (feature 2) Extent - Unknown Diversity - Unknown Mixture component habitat: Intertidal zone (feature 6) Quality - Unknown 6 Other habitat (not included in SSSI notification) : Heathland (feature 20) Extent of Calluna vulgaris - Unknown Diversity - Unknown - Unknown - Unknown Proportion of bare earth: building Calluna : mature Calluna : degenerate Calluna % cover Calluna in transect 1 No more than 5% of heather will be covered by bracken, bramble or shrubs LBBG will not be tolerated within the heath No rabbits will be tolerated within the heath Bracken removed from exclosure None present The rabbit exclosure has received more work but may need complete replacement in 2013. Summary table – Flora: Habitats and Communities * As no vegetation monitoring was carried out 2008 – 2010 these years have been grouped and data from previous years, where available, has been included. Feature 2006 2007 2008-2010 2011 2012 Within limits? Coastal grassland (feature 1) Extent of maritime grassland Extent of Thrift % Thrift in transect 79.1 Extent Silene dominated turf Extent of other maritime grassland dominated communities % Thrift tussocks alive in transect 93.4 quadrats % Vegetative cover (November) 78.2 (Oct) Soil erosion (cm) Feature 2006 Unknown. Unknown. Unknown. Yes in 2006. Unknown. Unknown. Unknown. Yes in 2006. Unknown. Yes in 2006. Unknown. 2007 2008-2010 2011 2012 Within limits? 7 Maritime Cliff and associated crevice habitat and communities (feature 2) Extent Diversity Unknown Unknown Mixture component habitat: Intertidal zone (feature 6) Quality Unknown Other habitat (not included in SSSI notification) : Heathland (feature 20) Extent of Calluna vulgaris Diversity Proportion of bare earth: building Calluna : mature Calluna : degenerate Calluna % cover Calluna in transect 1 Unknown Unknown Unknown 62 No limits set. 8 1.1 Flora – Habitats and Communities – Notes on tables Coastal / maritime grassland monitoring remains a huge task. Working out its extent is a combination of eight different projects, only one of which had time to be carried out in 2011 – coastal grassland transects. Three of the four were carried out in July rather than August this year due to other work commitments. Time restraints also meant results could not properly be analysed, but some observations are listed below; Thrift – decrease in all transects. Sea Campion – increase, particularly covering any bare earth toward the coast. Bracken – increased in density and extent of its outer limits. Bracken has remained relatively unchanged this year, with a consolidation of colonized areas becoming dense growth and boundaries slowly expanding. This looks set to continue without a coordinated effort to thin out growth and try to push back the encroaching edges through bashing/cutting/pulling of fronds. Bracken was remapped this season with small differences in extent noted since last season, mainly the southern sections recorded as more broken. This is probably due to observer error in both years as much as any real difference and work with aerial photography should give much more detail, this approach is currently being assessed for accuracy levels. Bracken in 2000. 9 Bracken coverage in July / August 2011. Mapping the coastal and maritime grasslands of the island remains a large task and if it is to be completed regularly with any degree of accuracy will need special attention from a long term volunteer or additional staff member as a possible secondment from another reserve. 10 1.2 Flora: Species Features and attributes Checked in 2012? Within limits of acceptable change? Operational limits (factors affecting feature) Notes on operation limits Yes 673 ind. Yes Yes No unauthorized public access off paths No unauthorised access around golden hair lichen Maintain the seabird assemblage (feature 18) Not met – see that section Golden hair lichen (feature 7) Presence Extent Assemblage of saxicolous lichens (feature 16) Quality (no. of species) Diversity (prescience of communities) - Not set No unauthorized access off paths - Not set No management practices that will affect lichens Very few instances of unauthorized access No activities carried out that knowingly affect lichens Assemblage of terricolous lichens (feature 17) Quality (no. of species) - Not set Maintain the seabird assemblage (feature 18) Not met – see that section Unknown. Campion has spread in recent years Very few instances of No unauthorized access off paths unauthorized access No management practices that No activities carried out that will affect lichens knowingly affect lichens Maintain limits of Sea Campion Diversity (prescience of communities) - Not set Other species : Three-lobed Crowfoot (feature 22) Quality (no. plants in November) - Unknown Ensure seasonality of pools continues where Crowfoot occurs Ensure no gulls nest in vicinity of pools Water in Dip Pool in March and November No gulls nesting too close 11 Ensure that Brookweed does not out compete Crowfoot Unknown Summary table – Flora: Species Feature 2006 2007 2008-2010 2011 2012 Within limits? Yes Yes Yes Yes Yes - Yes - Yes 673 ind. Yes Yes Golden hair lichen (feature 7) Presence Extent Assemblage of saxicolous lichens (feature 16) Quality (no of species) Diversity (prescience of communities) None set None set Assemblage of terricolous lichens (feature 17) Quality (no of species) Diversity (prescience of communities) None set None set Other species: Three-lobed Crowfoot (feature 22) Quality (no. plants in November) <50 >50 - Unknown 12 1.2 Flora – Species – Notes on tables No specific species surveys were carried out in 2012, aside from the checking for the presence of golden hair lichen. 13 1.3 Mammals Features and attributes Checked in 2012? Within limits of acceptable change? Operational limits (factors affecting feature) Notes on operational limits House mouse (feature 24) Population size No None set No mammalian ground predator will be tolerated No other small mammal will be tolerated None recorded None recorded Rabbit (feature 25) Population size, whole island (individuals) Population size, study plots (individuals / hectare) No Unknown Approx. 70/Ha No above 50/Ha At least 4 (as of Oct 10th) Yes (LAC of 4 pups) No mammalian ground predator will be tolerated Bracken extent will not exceed that of 1989 None recorded Limit exceeded in 2001 Grey seal (feature 26) Number of pups born annually No unauthorized access off No instances of unauthorized paths access onto shore 14 Summary table - Mammals Feature 2006 2007 2008-2010 2011 2012 Within limits? House mouse (feature 24) Population size None set Rabbit (feature 25) Population size, whole island (estimated individuals) Population size, study plots (individuals / hectare) 3000 1600 - - - Not in 2007 30 16 - - Approx. 70 No 1 0 (incomplete monitoring) 5 (at least) 5 (as of Oct 16th) Yes Grey seal (feature 26) Number of pups born annually 0 15 1.3 – Mammals – Notes on table House mouse Upon return in March there was extensive sign of mouse habitation, particularly in the Pantry and Warden’s Quarters (where six were trapped in the first 24 hours!). After an initial purge only the occasional one was seen through the summer months. During September and October low numbers were caught in the Wheelhouse, Kitchen/Pantry and Shop- around one or two per week. Rabbit A basic monitoring of rabbits was carried out this season with a set hectare on North Plain being surveyed on a regular basis in the evening. The population grew into the Autumn with high numbers during September and October, however no counts were possible in November due to the island being vacated at an earlier date than previously. This means counts are not directly comparable with historic data so cannot be relied upon but still indicate there has been a significant rise in rabbit numbers. The study plots on North Plain regularly reached 70 individuals when 50 is the upper limit of acceptable change as set out in the management plan. No Myxomotosis or Viral Haemorrhagic Disease was noticed this season. Grey Seal Regular counts of seal haul outs were made throughout the season with the peak count being 24 at Crab Bay on May 21st. These counts are largely subject to observer presence – the highest regular counts occurred during May to July, when the island was most occupied with visitors round the island all through the day. As of October 16th (time of writing) there were four recorded births on North Haven beach and one in Peter’s Bay with the sighting of at least two other young seals around the north coast, presumably recently moved from Skomer or the mainland. Unfortunately one of 16 the pups on North Haven did not survive to weaning age. As the pupping season here continues into December it is likely these were not the only pups on Skokholm. Cetaceans Regular cetacean watches were carried out throughout the season, with Harbour Porpoise seen almost every day when conditions were favourable. Common and Bottlenose dolphin were seen, occasionally in large groups. 1.4 Birds: Individual Species Features and attributes Checked in 2012 Within Limits of Acceptable Change? Operational limits (factors affecting feature) Notes on operational limits Manx Shearwater (feature 8) Population size in study plots (AOB) Whole-island population (AOB) 521 Partial (see notes). Mammalian ground predators will not be tolerated. None recorded. Coastal Bracken distribution will not exceed that of 1989. Limit exceeded in 2001and continually since then. No – LAC 550 Unknown. Soil erosion should not exceed 0.5cm/yr. Not measured. 17 Features and attributes Checked in 2012 Within Limits of Acceptable Change? Operational limits (factors affecting feature) Notes on operational limits No unauthorized access off paths. No significant instances of deviation from footpaths. No photography of Manx Shearwaters away from permitted areas. None reported. Storm Petrel. (feature 9) Population size in study walls (AOB) Population size in study transects (AOB) Population size, whole island (AOB) 14 84 - No – LAC 80 None set Unknown Mammalian ground predators will not be tolerated. None recorded. Little Owls will not be tolerated. None recorded. No encroachment of Bracken in contact with walls. Bracken was bashed back around the Cottage Garden walls. No unauthorized access off paths. Very few instances of unauthorized access. Walls will be maintained No work done in 2012. 18 Features and attributes Checked in 2012 Within Limits of Acceptable Change? Population size ( no. of territory holding pairs) 1 Yes Breeding success (fledged young) 2 Yes 2200 No – LAC of < 2500 pairs in 3 consecutive years. Operational limits (factors affecting feature) Notes on operational limits Peregrine (feature 10) No unauthorized access off paths No instances of unauthorized access near Peregrines. No unauthorized access off paths Few instances of unauthorized access. Distribution within specified areas. Largely No mammalian ground predators tolerated. None recorded. Bracken will not be reduced to less than half 1989 cover. Limit exceeded in 2001 and not reduced since. Lesser Black-backed Gull. (feature 11) Population size (AON) Breeding success (fledged young / AON) 0.16 Unlikely: LAC of < 0.4 in 3 consecutive years. Guillemot (feature 12) 19 Features and attributes Checked in 2012 Within Limits of Acceptable Change? Operational limits (factors affecting feature) Population size (individuals) 2330 Yes No unauthorized access off paths Number in study plots (individuals) 625 Yes Breeding success in study plots (fledged young / active site) - Unknown Population size (individuals) 1463 Yes Number in study plots 250 Yes Breeding success in study plots (fledged young / active site) - Unknown No mammalian ground predator Notes on operational limits No instances of unauthorized access onto Guillemot breeding areas. None recorded. Razorbill (feature 13) No unauthorized access off paths No instances of unauthorized access onto Razorbill breeding areas. No mammalian ground predator None recorded Puffin (feature 14) Population size (individuals – optimal spring count) 4637 Breeding success (chicks / active burrow) - Yes Unknown No unauthorized access off paths No mammalian ground predator Bracken will not be tolerated on Puffin slopes Few instances reported, immediately dealt with. None recorded Bracken is present on some slopes. Controlled in some areas. 20 Features and attributes Checked in 2012 Within Limits of Acceptable Change? Operational limits (factors affecting feature) Notes on operational limits No unauthorized access off paths No instances of unauthorized access Rabbit density in June = higher then 8/ha Exceeded; Mean 18.6, Max 70 There should be 65% vegetative cover to bare ground in Nov Unknown (last count 78.2 % cover in Oct. 2006) Chough (feature 15) Population size (apparent pairs) Breeding success (fledged chicks / pair) 3 0 Yes No 21 Summary table – Birds: Individual species Feature 2008 2009 2010 2011 2012 Within limits? Population in study plots (AOBs) 620 525 499 495 521 Whole island population (AOBs) Last completed count in 2001: c.45,000 No – Below LAC of 550 Unknown Manx Shearwater (feature 8) Storm Petrel (feature 9) Population in study walls (AOBs) Last census 2006: 23 AOBs 13 13 14 No – Below lower limit of 80 AOB Population in study transects (AOBs) Sites established 2010 84 78 84 No limit set Thought to be below lower limit of 2500 AOB Whole island population (AOBs) Peregrine Falcon (feature 10) Population size (no. of territoryholding pairs) Breeding success (chicks fledged) 1 1 1 1 1 Yes 2 1 0 4 2 Yes Lesser Black-backed Gull (feature 11) Population size (AONs) 2763 2396 2468 2412 2200 Breeding success 0.06 - - 0.03 0.16 No – Below LAC of 2500 pairs for 3 consecutive years. Unlikely 22 Feature 2008 2009 2010 2011 2012 Within limits? 1538 333 1697 556 1795 557 2212 613 2330 625 Yes Yes - - - - - Unknown 946 180 950 236 1140 257 1486 298 1463 250 Yes Yes - - - - - Unknown Population size (IND) 3198 (unreliable count) 3055 (unreliable count) - 4553 4637 Yes Breeding success (young / active burrow) - - - - - Unknown 2 4 (best guess) 2 3 (best guess) 2 2 3 Yes 2 4 0 Yes Guillemot (feature 12) Population size (IND) Number in study plots (IND) Breeding success in study plots (young / active site). Razorbill (feature 13) Population size (IND) Number in study plots (IND) Breeding success in study plots (young / active site). Puffin (feature 14) Chough (feature 15) Population size (apparent pairs) Breeding success (total fledged young) 23 Birds – Individual Species – Notes on table Manx Shearwater Study plots B to K (excluding the discontinued sites E and G) were monitored by callresponse around the first week of June. A full-island count was partially completed and at the time of writing it appears that the figures are similar to those in the 2001 (45,000 AOBs), although further work is needed (C. Perrins, pers. comm.). As part of the full census we carried out a study on 100 burrows to establish a response rate, giving us a conversion factor that is used to estimate the number of AOBs from the number of responses recorded. This factor was 2.06, slightly higher than the previously used 1.98. The total of apparently occupied burrows (AOBs) in the study plots is 521. This is the fourth consecutive year it is below the lower LAC of 550 AOBs. It is perhaps worth a more detailed look at some of the study plots to try and see what is happening to the island as a whole. 24 The weather, and in particular the rain, has probably been the biggest factor affecting Skokholm this year, but how it affected the shearwater breeding season is unclear. There was standing water in several burrows, particularly in plots D and J (located near North Pond and The Dip respectively), while others were clearly wetter and muddier than usual. While some well drained areas showed increases (plot C) and poorly drained ones decreases (plots B and D) the others are not so simple. Plots J and K look like they should be well-drained but results would indicate not. Plots F and H are on similar brackencovered ground, with F slightly more inland; they had wildly differing results. My personal feelings as to how the rain has affected the shearwaters, based on observations but no data, is that while some have been flooded out of their burrows, mostly those from the island’s interior, a greater number than expected have remained. As these birds are so long-lived it may be reasoned that the more mature breeders have witnessed similar breeding seasons in the past, and established themselves in more resistant burrows in better-drained parts of the island. 25 Storm Petrel Transect monitoring took place between 25th June and 22nd July. 98 AOBs in the transect study areas is a very similar count to the last two years that the study has been running. While this could easily be accounted for by natural variation or observer experience there also appears to be some movement among some of the more fragile areas of the quarry, in particular the lower end of transect 2. Storm Petrel table 1 shows this in clearer detail. Year 2010 2011 2012 North Pond Wall 4 7 5 Little Bay Wall 9 6 9 North Haven Gully 18 19 21 Quarry Transect 1 8 11 12 Quarry Transect 2 15 13 8 Quarry Transect 3 11 10 10 Quarry Transect 4 32 25 33 Total 97 91 98 Storm Petrel table 1. AOBs in each transect for the past three years. Storm Petrel table 2 shows that for the second successive year response rates are low, with birds calling, on average, less than 3 times in the 10 visits. There were several sites known to be occupied (researchers could see the bird) but no response was forthcoming. This figure emphasizes the need for not rushing each transect and listening intently for any response. Year 2011 2012 North Haven Gully 3.56 2.76 Quarry Transect 1 3.36 3.67 Quarry Transect 2 1.77 3.10 Quarry Transect 3 2.40 2.30 Quarry Transect 4 3.32 2.64 Average 2.88 2.89 Storm Petrel table 2. Response rates (number of responses by each bird out of 10 visits). The wet weather meant that Storm Petrel monitoring was delayed (access to the quarry was often deemed too risky) so was still being carried out at a time when several chicks could be heard. Curiously this didn’t lead to lower response rates later in the season (as adults spent more time at sea), with some of the peak visits occurring late on. Peregrine Falcon One pair nested on the Bluffs, overlooking Wallsend Bay. The same place they nested in 2009 and 2011 (and possibly 2010). They raised two chicks which successfully fledged. This is the average for the last ten years. Lesser black-backed Gull Walk-through and vantage-point counts of all colonies were carried out 20th to 23rd May. The cold April weather meant that it was early in the nesting season, although the lower bracken made vantage counts easier. This gave a figure of 2412 AONs. 26 This is the fourth successive year that the population has been under 2,500 AONs, the lowest acceptable limit set in the management plan. 2012 saw a drop in numbers to the lowest level yet. While the major colonies held similar numbers it was clear that the smaller ones were spread very thinly. Productivity was calculated by counting fledged chicks within specific colonies. This may lead to bias as it is carried out at the smaller, easier to observe colonies. The figure of 0.16 fledged young / AON is very low, yet higher than last year. It is higher than expected as much of the year was very wet, and chicks spending a long time hiding in damp bracken and long grass tend not to survive. Guillemot Once again there were record counts of guillemots in the study plots (625 ind.) and on the whole island (2,330 ind.). The study plots have been counted for 10 years now and show a 239% increase since 2003 (the whole island counts have increased 217% in the same time). Unfortunately, due to rough seas, we were unable to carry out a count by boat this year, so there remains a section of north coast that was missed, while other parts of the north coast and The Bluffs were counted less accurately at a distance. Attempts to carry out productivity studies proved unsuccessful due to a lack of time. 27 Razorbill Counts of the whole island and study plots (1,463 and 250 individuals respectively) were marginally lower than 2011s record numbers. The study plots in particular showed massive variation in the extremely mixed weather conditions; from 164 ind. on 7th to 338 the next day when the wind died down and the sun came out. Attempts to carry out productivity studies proved unsuccessful due to a lack of time. Puffin An island-wide count on 21st April gave us a figure of 4,637 individuals. The above graph includes counts of around 3,000 birds in 2008 and 2009. These should probably be ignored as they were carried out on sub-optimal evenings when the island was mainly closed. With this in mind it appears the population has been steady between 4,000 and 5,000 for the last ten years. 64 birds were fitted with colour rings, making 196 individuals with those done last year. 99 of the 2011 birds and 44 of the 2012 ones were seen, a total of 143 individuals, 73%. 28 Chough Three pair of Chough were regularly seen around Skokholm, but their nesting statuses were very difficult to pin down and no chicks were recorded. It appeared as if one pair nested in their usual place in Steep Bay, with the second pair again trying on the south coast, east of The Dip. This second pair seemed to start quite late and fail or abandon quite early, both adults being regularly seen associating with another pair around the quarry from early June. This last pair were often seen in that location but didn’t appear to make any attempt to nest. This is the sixth successive year that two pairs have attempted to nest on Skokholm, although they were not as successful as last year. Chough can be particularly difficult to monitor and with the infrequent occupation of Skokholm in the three years before reopening records can not be 100% reliable. 29 1.4.1 Birds: Assemblages Features and attributes Checked in 2012? Within limits of acceptable change? Operational limits (factors affecting feature) Notes on operational limits Seabird assemblage, species other than those as features in their own right (feature 18) Fulmar Population size (AOS) Breeding success (fledged young / active site) 136 Yes 0.59 Yes 274 Yes 1.15 Yes Herring Gull Population size (AON) Breeding success (fledged young / active site) Great Black-backed Gull Population size (AON) Breeding success (fledged young / active site) 73 No – Upper LAC 70 - Unknown Seabirds assemblage, feature as a whole No due to Manx Shearwater, Storm Petrel & Gulls. Note: This feature includes those seabird species previously identified as features. For this feature to be in a favorable condition, all the components must be within limits. No LBBG will nest outside permitted areas No nesting outside of permitted areas Bracken will not encroach on 1989 limits Bracken exceeded 1989 limits in 2001 and continue to do so. Soil erosion not exceeds 0.5 cm/yr Not measured Little owls will not be tolerated None recorded Mammalian ground predators will not be tolerated None recorded No unauthorized access off Few instances of paths unauthorized access. Other species; assemblage of important ground-nesting birds (feature 23) Oystercatcher Population size (AOT) 32 Yes Bracken will not encroach on 1989 limits Bracken exceeds 1989 limits (measured in 2001) 30 Breeding success (fledged young / active site) - Rabbit density in June above 20/ha Gull counting methods will be developed to cause less disturbance Mammalian ground predators will not be tolerated. Unknown Lapwing Population size (AOT) 0 No – LAC 2 Skylark Population size (AOT) 7 No – LAC 8 Wheatear Population size (AOT) 10 Exceeded; Max 70 Walk through counts kept to a minimum, more vantage point counts None recorded No unauthorized access off Few instances of paths unauthorized access Yes Summary table; Birds - assemblages Feature 2008 2009 2010 2011 2012 Within limits? Seabird assemblage, species other than those as features in their own right (feature 18) Fulmar Population size (AOS) Breeding success (fledged young / active site) 119 116 118 0.36 (estimate) 128 136 Yes - - 0.49 0.59 Yes Population size (AON) 287 353 312 257 274 Yes – LAC <270 in 3 consecutive years Breeding success (fledged young / active site) - - 0.82 0.67 1.15 Yes 61 69 80 84 73 No – Upper LAC 70 Herring Gull Great Black-backed Gull Population size (AON) 31 Breeding success (fledged young / active site) 0.97 - 0.71 0.89 - Unknown Other species; assemblage of important ground-nesting birds (feature 23) Oystercatcher Population size (AOT) 25 31 31 28 35 Yes – LAC 30 in 3 consecutive years. Breeding success (fledged young / active site) - - - - - Unknown 0 0 0 0 0 No - - 4 (estimate) 6 7 No – LAC 8 in 2 consecutive years - - 8 (estimate) 14 10 Yes Lapwing Population size (AOT) Skylark Population size (AOT) Wheatear Population size (AOT) 32 1.4 Birds – Assemblages - Notes Fulmars Study plot counts and five whole-island counts were carried out during the first ten days of June, giving figures of 20 and 136 AOSs respectively. While the number in study plots is the same as it has been the last five years, the whole-island population appears to be rising again after a slight dip. It is likely that some fulmars were missed this year as no boat count was possible. The productivity of 0.59 is established from 56 sites in Little Bay, North Gully, Twinlet Bay and Peter’s Bay. A total of 37 chicks hatched in these and 33 fledged from these nests. This is slightly above the average of 0.42, taken from the years 1981 to 2004 and again higher than last year’s figure of 0.49. This is significantly above the Lower Acceptable Limit set and shows a continuation of the recent short term rise in numbers. Herring Gulls Counts were carried out on the cliffs and within other gull colonies in mid-May. This gave us a figure of 274 AONs. This is low compared to the Herring Gull population 35 years ago, but the mean AONs since 1984, when the figure first fell below 500, is 320, and for the last ten years it is 285, so this years count is within the expected range. 33 Like the Lesser Black-backed Gulls and other gull populations across the UK, the Herring Gull population experienced a big increase between the late 1950s and early 1980s, reaching a peak of 1400 AONs in 1975. It is interesting that the Skokholm population appears to be stabilizing around its 1930s population of 250 – 300 AONs, this year the island total being 273 AONs. An estimated productivity of 1.15 fledged young / AON (based on the population on the Neck) is significantly higher than the average for the years 1989 – 2011. It is also significantly higher than that of the Lesser Black-backed Gulls, thought to be a result of the birds (at least those being monitored) nesting on the cliffs rather than amongst dense vegetation so in-between rain showers the young birds had a chance to dry off and warm up rather than be surrounded by wet grass and bracken. Great Black-backed Gulls Nesting pairs were noted throughout April and May, giving a whole-island total of 73. This is a slight drop on the last two years, perhaps reflecting the increased occupancy of the island and the disturbance that comes with that; four pairs nested on Little Bay Wall in 2011 compared with two this year. The steepest rise in breeding numbers was 20072011 so a target to maintain pre-closure numbers of 50-60 pairs may prove to be more sensible than maintaining present numbers alongside increased disturbance from renewed activity on the island. The upper limit to acceptable change as stated in the management plan is 70 AONs so this new dip in population should remove the need for limiting numbers. Counts of roosting birds stayed high through to October with around 100 birds present on North Plain in the twilight hours. This shows the continued importance of the island to 34 the birds and increased disturbance in future seasons will hopefully be managed to limit any reduction in numbers. Oystercatcher There were 35 AOTs, well within the expected range. Since 2002 there has been an average 34 AOTs, with a maximum of 48 and minimum of 25. Skylark Seven territories is below the lower LAC of 8. Wheatear Ten AOTs is low both in terms of historic and recent populations, although recording of the latter has been sporadic. 35 1.4.2 Other Birds Canada Geese 16 AONs (excluding relocations of disturbed nests) were found in 2012, although up to 50 birds were regularly present. These totals are fewer than the last few years, suggesting that increased occupancy and nesting disturbance (see 2011 report) is being successful. Up to 17 goslings were seen on North Pond but they were either predated or failed in some other way as only one appeared to fledge. Raven For the fourth year in succession three pairs of Raven were present, but they had very different levels of success; a pair near Twinlets appeared to abandon their nest very early on; a pair in Crab Bay used the same nest as last year which collapsed in mid-April, dropping the one remaining chick that was then lost to large waves; and the pair near Blacksmith’s fledged three chicks. Assuming the three pairs are nesting in roughly the same place each year, it is interesting to note that the pair that were successful this year were the pair that failed in 2011. Similarly, the pair at Twinlet Bay have successfully bred or abandoned early in alternate years since 2009. 36 Common bird census Other breeding birds are summed up in the following table. Many of these were recorded during common bird census. Numbers for Jackdaw, Meadow Pipit, Rock Pipit and Wren are best estimates – there may be more birds within gull colonies, areas of fragile ground or on cliff edges. Species Pairs / territories Shelduck 2 Shag 1 Water Rail 1 Moorhen 2 Buzzard 1 Swallow 5 Meadow Pipit 42 Rock Pipit 35 Pied Wagtail 3 Wren 52 Blackbird 4 Sedge Warbler 4 Jackdaw 15 Carrion Crow 5 Reed Bunting 2 Notes Two sets of parents with young ducklings were seen two weeks apart in June. It is unclear how well they fared but on was seen taking five young down to the sea. A pair nested on the north side of The Neck, possibly the first Shag pair to attempt nesting on Skokholm since 1987. Despite getting three chicks to quite a large size they all died in one rain storm in early June. Regularly heard and occasionally seen around Well Pond. Territories in North Pond and Well Pond, although there could be more than one pair in each place. One pair was successful and raised two chicks. Two around buildings, one at red hut, one at lighthouse, one in North Pond hide. Appeared to have a very successful year. Apparent territories, although much of the island was not checked e.g. close to gull colonies, over fragile ground. Very similar to 2011 count. Present all around the coast and seemed to have a very successful year raising chicks. Pairs around the Garage, East Bog and the Lighthouse. Very abundant – singing from every bay and boulder. Very similar to 2011 count. Behind Wheelhouse, near The Well, North Haven and East Bog. After the exceptional year in 2011 a return to more familiar numbers. Very difficult to estimate but this is conservative approximation is very similar to the 14 pairs last census (in 1996). Five definite nest but can be a difficult species to census. Pairs at Well Pond and East Bog. 37 1.5 Herptiles Slow worms were present underneath corrugated sheets throughout the season. 1.6 Invertebrates Butterfly transects were not carried out weekly, but occasional ones were done. Wet weather for much of the summer meant that numbers were low. The following species were noted as present at least once in 2012: Small White, Large White, Small Copper, Red Admiral, Painted Lady, Small Tortoiseshell, Peacock, Meadow Brown, Small Heath. After the purchase of a battery powered trap, recording effort was increased this year and varying locations across the island were sampled. Wet and windy weather meant results were often limited but hopefully more regular and systematic trapping can occur next season. A full list of moths is available separately. 38 2.0 Human Impact 2.1 Visitor numbers Overnight guests Skokholm was open for up to sixteen overnight guests, staying in the Middle Block and newly-refurbished Cottage, between 11th May and 3st August. With changeover boats on Mondays and Fridays stays were in blocks of three of four days, with a handful of visitors staying a full week. The maximum number of people that could stay on Skokholm this season was 432- if all 16 beds were full and we were fully booked with no one staying a full week to ensure maximum turnover of guests. 202 guests were booked on to stay in 2012, but some of these did stay a full week and several guests paid extra for single occupancy of rooms so the figures are not quite as stark as they seem. The largest problem this season was bad weather disrupting boat services with 72 guests being affected with either total or partial cancellations of their stays. These lost days (and relevant refunds) represent over 15%* of the island’s capacity at present, and when combined with rooms that were never booked out in the first place it shows there is considerable margin for increasing revenue from residential visits. Obviously it has been an extreme summer of weather but these figures alone justify further time and effort being spent on maintaining and improving the landings at South Haven and Blacksmith Ledges. The latter of these will require significant work on both the path leading to it and the steps of the landing if it is to be used safely on a semiregular basis. *16.2% of “bed-nights” out of a maximum 1,520 (16 beds x 95 nights) were refunded in 2012 2.2 Disturbance to Wildlife Island visitors There have been no serious incidents of visitors disturbing wildlife this season. The reduction of day trips has undoubtedly helped with this. The significantly increased number of people combined with a brief introduction to the island and day-trippers need to cover the whole island in a few hours frequently led to disturbance, mainly from people leaving the paths. Confusion arising from day-trippers and overnight guests arriving on the same boat also led to problems in previous years. The Founders day trip 39 this season did have a few minor incidents with groups straying slightly from paths but no real damage was done. As done last season, the South Coast path was left in place from the lighthouse and diverted off to the main track before reaching Crab Bay. This path was open but not advertised and the start/end of the park was unmarked to minimise interest. This led to little disturbance from its use but it may be wise to make it entirely off-limits to guests due to disturbance and health and safety considerations (although this will inevitably draw some criticism from long term visitors of the island). Boats During the time Skokholm was inhabited there was no serious disturbance noted from boats. Sea Safari, dive boats and kayakers all come in close, especially to South Haven, but those observed have appeared aware of the marine code and done their best not to disturb seals or puffins. As these are what is drawing these visitors out here they are treating them with respect, keeping a reasonable distance away, moving quietly and slowly. Moving with less care is the occasional lobster-potting boat coming in close; however this was not noted during the main breeding season. Low flying aircraft There were no sightings of aircraft while wardens were on the island. The lighthouse was visited several times by Trinity House employees, each time the helicopter arrived over the sea. 2.3 Pollution No incidents of pollution were encountered or reported to the island staff. 40 3.0 Reserve Administration 3.1 Finance Island Budget Budgeting was handled by the mainland and Skomer staff. Fees Regular day trips were not run in 2012 but a Founders Day trip did proceed, this was free of charge as usual. Overnight guests were charged £125 for a three or four night stay (Monday to Friday or Friday to Monday). This did not include boat fares. Volunteers were not charged for staying on the island. Tuck shop The shop was stocked with a range of dried and tinned goods so self-catering guests would not have to bring everything for themselves. There was also a range of snacks that sold well. There was a large variation in the amount that guests used the shop, with some relying on it, others barely using it. This year the shop was almost emptied at the end of season to generate a turnover of stock, and will be restocked in the spring. Island Shop The Friends’ shop was set up in the library, having a range of books, sweatshirts, hats and keyrings available in return for donations. Money from this went to the Friends of the Islands. Donations Donations of any items, many of which have been made for the restoration project, have been handled by the Friends of Skokholm and Skomer. 3.2 Staff Jerry Gillham was employed as Skokholm Warden from 2nd March – 29th November. William Whittington was employed as Skokholm Warden from 19th August - 30th November. Lewis Yates was employed as Skokholm Warden from 20th August – 26th October. Chris Taylor was employed as Skomer Island Warden with responsibility to oversee work carried out on Skokholm. 41 3.3 Volunteers A series of Long Term Volunteers was vital to the running of Skokholm in 2012. Their stays were staggered so through peak season there was always two volunteers, one of whom had at least a months experience when the other arrived. This reduced the amount of pressure on the warden and meant one could learn from another. The initial long term volunteer also stayed on for the entire season so that three volunteers were present for much of the year, which helped with the transition between staff members. Alongside helping with monitoring, visitor work, daily cleaning and maintenance, most volunteers had their own specialist interests and carried out individual projects. Name Dates present Will Whittington March 12th to Sep 27th Molly Heal Start of April to June 1st Teresa Donohue June 1st to Oct 26th Leighton Newman July 9th to Sep 22nd Notes Helped with getting island up and running in Spring, work party projects, running of the island and took on Warden position for two months as cover. Helped with running and maintenance of the island and did a large portion of bird counts. Helped with running and maintenance of the island, joined work party groups and stayed on after to close down island and keep records of birds present. Also aided and wrote up much of the bird monitoring- Gulls, Fulmars etc. Helped with running and maintenance of the island and stayed for all but one week of the work parties in the Autumn. Did a lot of bird monitoring and vegetation work, writing up a report on the status of the Heather remaining on the island. The volunteers who carried out all the restoration work are too many to list here (a full report will be produced separately) but thanks must go to all who put in vital weeks of hard labour. In addition there were many volunteers from Skomer, Dale Sailing and the Marloes / Dale local community who aided with the delivery of materials for the building project. In particular John Walmsley and Steve Sutcliffe, who organized the purchasing of materials, their delivery, the work-parties and the restoration plans. 3.4 Training Minimal official training was needed this year, with Jerry Gillham already holding the First Aid at Work and Powerboat Level II certificates. Neither of these was needed but could have been. 42 First Aid at Work training was received by Lewis Yates, Will Whittington and Teresa Donohue. In house training was given to all volunteers associated with day trips, monitoring and restoration work. 3.5 Health and Safety General All Risk Assessments were reviewed and in some cases rewritten by J. Gillham at the beginning of the season. These were read by all Long Term Volunteers in the first week of their stay. First aid kits were replenished at the beginning of the year. Fire blankets were visually checked and extinguishers were taken to Skomer and checked by the fire brigade. Working equipment, including helmets, harnesses and ropes, were checked by J. Gillham before use and sent off for professional checks in October. Accident forms have been completed for any incidents or near misses that occurred during the year. The health and safety record was very good, especially when considering the nature of work party tasks. Visitors All visitors were given an introductory talk, which highlighted the dangers of the fragile ground and cliffs. Any specific areas of fragile ground were pointed out during the compulsory guided walk. The south coast footpath was closed to visitors as it has eroded somewhat and become particularly difficult to follow. Other footpaths had become somewhat overgrown so a large effort was made to remark them with white stones and temporary canes. Night walks to observe Storm Petrels in the Quarry were carried out only in the company of staff and volunteers. Everyone was instructed to take torches and warm clothes and were warned about the terrain. Future consideration may be made as to putting ropes and handholds down there. The landing was scrubbed with Dairy HypoChlorite to keep it free from algae and seaweed growth, ensuring it was safe to walk on for all visitors and for deliveries of building material. In addition to this it was painted with anti-fouling paint which restricted algae regrowth and significantly reduced the quantity of hypochlorite required. 43 Work parties Risk assessments were drawn up by those organizing the restoration work. Personal protective equipment and clothing was supplied and training was given as required. Some specific jobs, including removing asbestos roofing and rebuilding the roof, were carried out by professionals. 3.6 Management planning There has been no development of the management plan since 2007 and it is in need of a re-write. The register of conservation projects to take place on Skokholm is also in need of rewriting as it is very comprehensive and would require additional staff to be completed to proper standards. Many of these projects are in need of updating whilst some could very possibly be done without. 44 3.7 Trust New Member Recruiting Trust membership leaflets were available in the Wheelhouse on day-trips. The map that is given to visitors contained revised information about the restoration project and details of the Trust. 3.8 Friends of Skokholm and Skomer Friends of the islands leaflets left on the island have suffered in the damp conditions and had to be removed. However the publicity of the blog and general information have been prominent in the Wheelhouse and many visitors showed a great interest in the group. Recent newsletters were displayed in the Wheelhouse and information about the Friends was on the reverse of the visitor map. 45 4. 0 RESERVE MANAGEMENT 4.1 Media and publicity Several programmes were filmed out on the islands this year with the majority taking place on Skomer, there was one exception however. Dream Island was a six part documentary on the history, wildlife and life of Skokholm. It had been filmed over the previous two seasons by Gordon Main and an associate. It was aired in between episodes of Coronation Street at 8pm on Fridays in September and October. The program seemed to get good feedback and combined with the large range of media coverage from 2011 the profile of Skokholm should continue to increase. Social media efforts were also expanded with a new Facebook profile for the warden and page for the island increasing routes for communications. This builds upon the Skokholm blog which continues to get relatively high viewing ratings. Visiting photographers were again encouraged to attach links from their blogs to the island’s website, with a reciprocal arrangement from the Skokholm blog proving mutually beneficial. Bird sightings were publicized in the back of Bird Watching magazine as well as on the Pembrokeshire Birds blog. In the future this could be expanded to a Twitter feed and very possibly small articles in bird watching and general wildlife magazines. Journalists should continue to be encouraged to visit as there are many angles to conservation and island life, and most individuals have a very positive on first visiting the island. 4.2 Liaison and relations A good relationship was maintained with all relevant organizations and individuals and effort was put into improving communication between the island and mainland staff and other organizations. 4.3 Habitat and species management Bracken Bracken was cut back mainly from the paths, using a combination of bashing, pulling and cutting with shears and the brush-cutter. Some cutting of bracken on puffin slopes was accomplished. 46 The distribution of Bracken is very similar to that of last year – see section 1.1. and above acceptable limits of the management plan. Heather Despite repeated repairs the exclosure is still not fully rabbit-proof and has further deteriorated. This is something that needs to be addressed urgently and a complete rebuild may be necessary in 2013. No gulls nested in the exclosure and bracken was removed by pulling but this needs repeat efforts. Small mesh exclosures were made for individual plants and a volunteer project was to assess the state of the heather. 4.4 Infrastructure Buildings – visitor accommodation and storerooms See introductory section for report of works carried out on the buildings as part of the restoration project. Communal buildings cleaned daily, visitor rooms cleaned before new arrivals. Small jobs done as necessary. Buildings – hides Minor repairs were needed to the North Pond, Neck and South Haven hides, mainly on the doors and flaps. Some re-painting and wood-staining was needed. This work may well need repeating in the spring as some sections of wood are deteriorating quite rapidly. Water and sewage Rainwater tanks were not used for anything more than casual hand-washing and kitwashing. Water from the well was used for all cooking and washing. The semi-compost toilets were emptied into a spoil pit, this one is not on the hill so will not drain into the well catchment area. This pit was filled at the end of the season so a new one was dug, also in a separate catchment area. Rubbish disposal All visitors and volunteers were asked to take away their recyclables (metal, glass, thick plastics). Of the rest, that which could be composted or safely burnt was done so. This worked well but needs to be done with every group to prevent build up. A large amount of refuse collected due to building works and should be shipped off the island when possible. Paths All path markers were checked and re-painted at the beginning of the season, with more markers added when needed. Bracken was cut from edges of paths on an irregular basis. Again the South Coast path was open but not publicized, and a stretch toward the Crab Bay end was altered to avoid the most fragile ground. 47 Landing places Landing cleaned of algae on a fortnightly to three-week basis during the visitor season. The new steps received more layers of white anti-fouling paint which lasted several months last season. This seems to be a worthwhile effort and reduces the amount of hypochlorite required to remove algal growth. Separate efforts were required to reduce seaweed growth in the more open waters in South Haven, which consisted of cutting by hand the longer fronds to stop them interfering with the Dale Princess’s water intakes and propeller. 4.5 Machinery and equipment Island computer The island laptop has continued to function well, even in the sometimes damp conditions. The dongle has also provided sufficient internet access with signal being acceptable in the new cottage office. Backups of computer data are still as important as ever. Dumper truck The dumper truck worked throughout the season with very few problems, only a few idiosyncrasies. The steering ram is functioning but in need of replacement, the parts for which have been acquired and are awaiting delivery to the island. Regular repainting and liberal application of duck oil at the end of season should see it well through another winter. In addition the garage has been re-roofed and should be a considerably better shelter this time around. Gator The Gator is a small petrol powered buggy which was included in the sale of the lighthouse. It has proved relatively reliable and much more suitable for moving lighter items around the island, especially to and from the lighthouse. Two slow punctures in the rear tyres meant that it was put away early for the winter but these should be fixed for next season and this vehicle will certainly prove its worth. Power barrow The power barrow had been temperamental in 2011 and in need of regular use. This did not occur in 2012 and the decision was made to take it off island. The acquisition of the small Gator vehicle with the lighthouse provides a much better alternative for smaller jobs. Gas system and appliances Gas system checked by the Warden at start of season and some minor works carried out. The gas system was checked and adapted by a qualified contractor and may need some small changes for next season as one bottle is very prominent in the courtyard. 48 All of the gas lights have now been removed. The two cookers in the kitchen continue to function well. All gas appliances, isolator switches and gas bottles turned off at the end of the season. Electricity generators and system The electricity system installed last April has provided ample power through even the worst weather again this season. This is despite it being extended so that the solar panels now power all of the main buildings. All of the accommodation now has LED lighting and charging sockets in the library have been supplemented with two more in the Wheelhouse and some in the office for staff/volunteer use. Generators were used when using power tools, cement mixer, etc. but outside of work party weeks this is minimal. Old system was removed during autumn work parties and the batteries used to supplement items such as extractor fans in toilets. Batteries were left connected over winter to maintain condition. Hydraulic RAM pump The RAM pump was dismantled and serviced by a long term volunteer at the start of the year and has been relocated during the autumn work parties. This included it in the new water system and should give it enough header pressure to ensure efficient working. Radio The handheld VHF radio has continued to be essential for communicating with boatmen and coastguard if necessary. It has remained in good working order and battery life has stayed high. The old VHF remains unmounted and boxed up but could well be reinstated for future use in the office in the cottage. 4.6 Food deliveries Food deliveries to restock the shop were made approximately once a fortnight. Stock levels were checked casually each week and more thoroughly roughly every three weeks. The system in place (warden emailing a list of what is needed) worked well although it needed some fine tuning and undoubtedly will need more. 49 5.0 Acknowledgements As well as those mentioned in the volunteers section above, thanks must go in particular go to Wendy Barnes-Jones for her expert organizational skills with regards the day trips. Thanks to the boatmen – Kenny, Karl, Peter, Tom and John - for their help with some tricky landings. 6.0 References J. Milborrow (2007) – Storm Petrel Pilot Study on Skokholm Island. Report for the Countryside Council for Wales, contract no. FC73-01-583 S. Smith, G. Thompson & C.M. Perrins (2001) - An estimate of the Manx shearwater population on Skomer, Skokholm and Middleholm islands. Report for the Countryside Council for Wales, contract science report no. 342 G. Thompson (2007) – The Natural History of Skokholm Island. Trafford Publishing. 50