Terrestrial Biomes Biome – defined Concept of “climate space” Climate diagrams Potential vs. actual vegetation A brief biome tour Readings: Chapter 2 – esp. pages 14-18 Terrestrial Biome defined • Major division of the terrestrial biosphere (broader than “ecosystem” or “landscape”) • Large-scale classification of the terrestrial habitat, distinguished by predominant plants, and associated with particular climates. – Examples include: desert, rain forest, tundra • Globally, there are about 6-9 major terrestrial biomes (depending upon who’s counting) • Similar terms: life zone, ecoregion, biogeographic province Terrestrial Biomes Campbell What determines the distribution of the terrestrial biomes? Distribution of annual temperature ranges Krebs 2001, Figure 7.2 Annual precipitation - a function of temperature and moisture patterns. Krebs 2001, Figure 7.3 What determines the global distribution of temperature and precipitation? Atmospheric circulation patterns (e.g. “cells”) create zones of temperature and precipitation Distribution of Earth’s major dry & wet areas Related to Hadley cells of air circulation • These circulation patterns are further modified by the Coriolis effect due to the spinning of the Earth. • Note the distribution of land and ocean masses also affect temperature and moisture At a regional scale, topography and distance from water bodies influence vegetation distribution (California) Bakker 1971 Temperature (energy) and moisture (precipitation) are primary determinants of terrestrial biome distribution Additional phenomenon and processes determine vegetation distribution at smaller scales (e.g. topography, distance from water bodies, ElNino/Southern Oscillation and other periodic climate phenomena). Convergent evolution Unrelated plants from different parts of the world often show similar appearances when exposed to similar climates Senecio sp. on Mt. Kilimanjaro Espeletia sp., Andes Mts. Molles & Cahill Fig. 2.40 …This introduces the concept of “potential vegetation” according to “climate space” Campbell & Reece 2002 Biomes “Climate Space” • Each region of the Earth can be defined by it’s particular “climate diagram” or location in “climate space.” • Temperature and moisture are primary determinants of distribution. • Local organisms are adapted to that particular climate (temperature-moisture space). Climate diagram for Blythe, California (Sonoran Desert) Krebs 2001, Figure 7.4a Climate diagram for Great Smoky Mountain National Park, Tennessee Krebs 2001, Figure 7.4b Terrestrial Biomes: distribution of “potential” vegetation according to climate space Campbell Problems with this concept: • Rapid climate change poses problems – can species distribution keep up? • Humans also alter vegetation, transforming “potential” vegetation to “actual” vegetation – how do we measure this? Deforestation in Rondonia, Brazil wet precipitation 7 4 2 dry 6 5 1 cold 3 temperature warm