1 COMMUNITY ECOLOGY Introduction In this section, we study the structure of a community. We also study the four main types of interaction within communities. We will investigate and discuss the human influence on the community structure of humans Community structure As we already know a group a organisms of the same species living in the same area is called a population. A number of different populations together are called a community. Example, grassland community near Johannesburg. It may consist grass, different insect and bird populations. A number of communities together make up an ecosystem. Within a community organisms can die classified based on their method of obtaining nutrients or feeding. Autotrophs e.g. plants have the ability to produce their own food by a process called photosynthesis. These autotrophs are also called producers. All plants and animals that obtain food from another source because they cannot make their own food is called heterotrophs e.g. humans. These heterotrophs are also called consumers and directly or directly are dependent on the autotrophs or producers for their food. Consumers can be categorised by what they eat. Herbivores are consumers that only eat plants, e.g. cows and sheep. Carnivores are consumers that only eat meat, e.g. lions and leopards. Omnivores are consumers that eat both plant and meat, e.g. humans and apes. These consumers can be placed into three categories: Primary consumers – they eat producers or plants. Secondary consumers – they eat the primary consumers. Tertiary consumers – they eat other consumers. This can be displayed in a pyramid of numbers, see Fig. 1 below. This indicates the number of organisms at each level to ensure healthy community structure or simply the numbers of organisms required to maintain a food chain. Fig.1 Pyramid of numbers Finally, there are decomposers that include fungi and bacteria. These organisms obtained food or energy from dead organic materials. 2 Interactions within a Community All communities are finely balanced with every individual organism involved in a battle for survival. The four types of interactions to be discussed in this lesson are predation, competition, parasitism, mutualism and commensalism. Predation Predation is the interaction between predator and prey. Predators kill others animals for food, e.g. cheetahs catching springbok. Fig. 2 Illustration of predation Predation also has an added advantage because it controls population size as illustrated below. An example is the lynx (predator) and dassie (prey) in Table Mountain. When the number prey increases there is more food available. That means that all the cubs of a lynx litter will receive food and all will reach maturity. That increases the predator population. As the predator population increases the predation pressure would increase which means that the predators will be killing more prey causing a decrease in prey population. As the prey population decreases the predators will have less food and their numbers will be decreasing. The decrease in predators allows the number of prey to increase. And the cycle continues. 3 Fig. 3 Graph of predator-prey interaction Competition Competition is the interaction between individual organisms for a limited resource, e.g. light, water, space, shelter and food. This interaction can lead to specialization for particular modes of life. Intraspecific competition is the competition between individuals the SAME species. Best known example, lions competing against other lions for the carcass (dead body) of zebra. Because the individuals are from the same species (Panthera leo) it is intraspecific competition. The principle is the same in the case of plants. If you plant a large number of mielies (Zea mays), the individual plants competing for water, nutrients and light will be of the SAME species thus making it intraspecific completion. Interspecific competition is the competition between individuals from DIFFERENT species. Best known example, while lions (Panthera leo) are eating the carcass of the zebra, hyenas (Crocuta crocuta) join to get some food and fighting breaks out between lions and hyenas. These are different species competing for the same resource and that is why it is interspecific competition. Again, the principle is the same in the case of plants. If you plant a large number of mielies (Zea mays) in a field with many weeds, the individual plants, mielies and weeds, competing for water, nutrients and light will be of DIFFERENT species thus making it interspecific completion. Gause's law refers to the principle that when two species compete for the same resources within an environment, one of them will eventually outcompete and displace the other. The displaced species may become locally extinct due to death of all representative individuals or migration to another suitable environment. This is called the competitive exclusion principle as illustrated in the example below. 4 When Paramecium aurelia and Paramecium caudatum species live in separate environments their population thrive as illustrated below. But when present in the same environment , they start competing with each other. Paramecium aurelia which is a smaller, faster growing outcompetes Paramecium caudatum. So Paramecium caudatum become extinct. Fig. 4 Competition in Paramecium Resource partitioning When resources are limited, natural selection will result one of the species utilising different resources. This allows the species to co-exist within the same community. This phenomenon is called resource partitioning. Example is in a forest where different species of plants survive in different environments of light intensity. In the African savannah, lion lived in the wide open grasslands. Leopards hunt in grassland but live in trees. 5 Parasitism Parasitism is a symbiotic relationship in which one species (parasite) benefits while the other species (host) is damaged. Most parasites will limit the damage to prevent death of its food source. There are two types of parasites, namely ectoparasites and endoparasites. Ectoparasites live outside the body of its host and examples include mosquitoes, lice, flea and ticks. Endoparasites live inside the body of its host and examples include tape-worms, round worms and bilharzia. Parasitism is regarded by some scientists as a form of predation. Fig.5 Common ectoparasite (mosquito) Most common South African example of an ectoparasite is the common mosquito shown above. It lives outside human’s bodies and suck blood through the human skin. Other examples include ticks that live on dogs and cows. These ticks suck blood from their hosts and in extreme cases can actually kill the host. Another South African animal parasite is the bilharzia parasite, Schistosoma haematobium , a parasitc flatworm. It is a very successful parasite and completes its life cycle in two hosts, a snail, Bulinus africanus and humans. Most common South African plant parasite is the Dodder (Cuscuta sp .) which lives on wheat. The dodder has small root-like structures, called haustoria which penetrate to the wheat's vascular bundles. The parasite obtains organic nutrients, water and mineral salts directly from the host. With all the nutrients, the parasites then spread to adjacent plants. 6 Fig. 6 Dodder Mutualism Mutualism is a symbiotic relationship in which both species benefit. There are two types facultative and obligate mutualism. In facultative mutualism both organisms benefit by living in close association, but is not essential. Example, the relationship between ants and aphids (plant lice) on rose plants. The ants feed on the sugary fluid released by the aphids, and the aphids are protected by the ants. Fig. 7 Mutualism between ants and aphids 7 In obligate mutualism is a relationship in which both species are forced to live in mutualism. Example, is a lichen which is a “plant” made up of a fungus and an alga living in close association. Lichens are visible everywhere including roof tops, rocks and tree trunks. The fungus absorbs inorganic substances which are then used by the alga to make food during photosynthesis. The fungus then gets food which was made by the alga. Another every common example is the relationship between bees and flowers. Fig. 8 Mutualism in lichens Commensalism Commensalism is a symbiotic relationship between two species in which one species benefits and the other is not benefitted or harmed. The host species provides a home and/or transportation for the other species. Most well-known example is the clown fishes live within the waving mass of tentacles of sea anemones as shown in the movie, “Finding Nemo”. Other, examples include birds nesting in trees. The bird receives benefit because its nest is far from predators. The trees are neither benefitted or harmed. Fig. 9 Commensalism