Biology 1 Outline and Objectives Unit 1 The Science of Biology Chapter 1 General Outcome 1.0 The students should be able to recognize the basic characteristics of life and describe the nature of science. Specific Learning Outcomes: Upon successful completion of this unit, the students should be able to: 1.1 List the major properties of life. 1.2 Explain how science is distinguished from other ways of seeking understanding of life. 1.3 Explain the significance of major unifying themes of modern biology. 1.4 Explain the limitations of science. 1.5 Identify how the steps in the scientific process were used to determine the effects of CFCs on the earth’s ozone layer. 1.6 Be able to define the following terms. The Scientific Process Deductive reasoning Inductive reasoning Stages of scientific investigation Observation Hypothesis Predictions Controlled experiment (testing) Control experiment 1 Variable experiment Scientific Theory Properties Cellular organization Metabolism Homeostasis Reproduction Heredity Biological Themes Evolution The flow of energy Cooperation Structure determines function Homeostasis Explain this statement: The process of science does not work to discover truth. Biology 1 106736530 1 Unit 2 The Chemistry of Life Chapter 3 General Outcome: 2.0 The students should be able to explain the structure of atoms, chemical bonding, properties of water, and the groups of organic molecules associated with life. Specific Learning Outcomes: Upon successful completion of this unit, the students should be able to: 2.1 Explain how the structure of an atom determines its chemical properties and the kinds of bonds it can form. 2.2 Describe and explain ionic and covalent bonding. 2.3 Name the elements that make up the majority of all living matter. 2.4 Recognize the structure of the water molecule, showing areas of positive and negative charge. 2.5 Describe a hydrogen bond. 2.6 List the major chemical and physical properties of water which result from the hydrogen bonding between water molecules 2.7 Describe the ionization of water and describe the pH scale. 2.8 Explain why the carbon atom plays a central role in the formation of organic molecules. 2.9 Describe the condensation and hydrolysis of carbohydrates. List examples of monosaccharides, disaccharides, and polysaccharides. 2.10 Describe the condensation and hydrolysis of triglycerides and the role of other lipids such as steroids and phospholipids. 2.11 Describe the structure of an amino acid and how polypeptides are formed. Explain protein variety in terms of amino acid arrangement. 2.12 Define primary, secondary, tertiary and quaternary structure of proteins and relate the structures to protein function. 2.13 Describe nucleic acid structure and function. 2.14 Describe theories and significant experiments regarding the origin of life on earth. 2.15 Define the following terms: Basic Chemistry Atom Nucleus Protons Neutrons Electron Orbitals Energy of position (potential energy) Ions Isotope = variations in # of neutrons Atomic number = # of protons Atomic mass = protons + neutrons Chemical bond Molecule 3 types of bonds Ionic bond- strong when solid, weaker in H2O Covalent bond- strong Hydrogen bond- weak Water Properties of water Polar molecule Cohesion Adhesion Solvent Heat storage = high Biology 1 106736530 2 High heat of vaporization = high, so carries away a lot of heat Lower density of ice- ice floats in water! Hydrophobic property of nonpolar compounds that are not water soluble pH pH scale Hydrogen ion (H+) Hydroxide ion (OH-) H2O OH- + H+ Acid Base Buffer Carbonic acid/bicarbonate H2O + CO2 H2CO3 HCO-3 + H+ Macromolecules Polymer Dehydration synthesis = making a polymer Hydrolysis = breaking down a polymer Organic molecule- backbone of carbon Functional group = groups of atoms with special properties that are added to a carbon backbone Carbohydrate C6H12O6 (H2O) Monosaccharides Examples: glucose, fructose Disaccharides Examples: sucrose Polysaccharides Examples: starches, glycogen, cellulose Functions: energy storage and structure Lipids- mostly C-H bonds, little O Fats Subunits: Fatty acids (3), glycerol (1) = triglyceride Saturated: usually animal fat, hard fat Unsaturated: usually plant oil Phospholipids Steroids- four-carbon ring structure Examples: cholesterol, hormones Functions: energy storage, cell membranes, hormones Protein Subunit: amino acids, 20 common ones Peptide bonds bind amino acids into chains Polypeptides = chains of amino acids Examples: Structural proteins Examples: collagen, keratin Enzymes- makes possible most chemical reactions of life Example: amylase, lipase Structure 1. Primary = the sequence of amino acids 2. Secondary 3. Tertiary 4. Quaternary Denaturation Functions: structural, enzymes (catalyst) Nucleic acids Biology 1 106736530 3 Subunit: nucleotide = 5-carbon sugar (2 types), phosphate (PO4), nitrogen-containing base (4 types) Function: Information storage for the message of life Deoxyribonucleic acid (DNA) = chemical blue print of an organism Double helix- base pairing Adenine – thymine (A-T) Guanine – cytosine (G-C) Ribonucleic acid (RNA) Unit 3 Cells / Energy and Life / How Cells Acquire Energy Chapter 4, 5, 6 General Outcome 3.0 The students should be able to describe a theory of the origin of cells, distinguish prokaryotic and eukaryotic cells, list cell organelles and their functions, describe membrane function, and detail the phases of mitosis and their significance. 3.1 The students should be able to explain: 1) the energy requirements of cells, 2) the central role of ATP, 3) the generation of ATP during cellular respiration, 4) the production of food by photosynthesis, and 5) the role of enzymes in controlling chemical processes in cells. Specific Learning Outcomes: Upon successful completion of this unit, the students should be able to: 3.1 Define the terms heterotroph, autotroph, prokaryote, and eukaryote. 3.2 Describe the structure of a cell membrane and a cell wall. Explain how they differ in function. 3.3 Describe the structure and function of the nucleus and the following cell organelles: ribosome, endoplasmic reticulum, Golgi body, lysosome, chloroplast, mitochondrion, and vacuole. 3.4 Describe microtubules and microfilaments and their role in support and movement. 3.5 Discuss the biological importance of maintaining a chemical composition that is different from that of the surrounding medium. 3.6 Explain the fluid mosaic model of membrane structure. 3.7 Compare and contrast movement through the cell membrane by diffusion, osmosis, facilitated diffusion, and active transport. 3.8 Describe endocytosis and exocytosis. 3.9 Explain how most living things are dependent upon the radiant energy of the sun. 3.10 Describe an oxidation-reduction reaction. 3.11 Describe the biological importance of enzymes and coenzymes and explain how they work. 3.12 Explain why ATP is often called the "universal energy currency" of the cell and describe how it performs its important function. 3.13 Recognize the summary equation for the oxidation of glucose to form carbon dioxide and water. 3.14 Detail the anaerobic process of fermentation in microorganisms and the production of lactic acid in human muscle during vigorous exercise. 3.15 Describe glycolysis, Krebs cycle, and the electron transport chain; list where each occurs in the cell, relative energy yield, and major events of each phase. 3.16 Recognize the overall equation for photosynthesis. 3.17 Recognize that life depends upon the visible portion of the electromagnetic spectrum and the chemical process of photosynthesis. 3.18 Describe the events of the light dependent and light independent reactions (Calvin cycle) of photosynthesis, explaining how the latter reactions depend on the products of the former reactions. 3.19 Recognize that the light-independent reactions of photosynthesis create building block molecules for plant cell macromolecules. 3.20 Compare and contrast photosynthesis and cellular respiration. 3.21 Define the following terms: Chapter 4 Cells Biology 1 106736530 4 Cell theory 4 principals of cell theory Cell surface to volume ratio Cytoplasm Plasma membrane Phospholipid bilayer Membrane proteins Cell surface markers Transmembrane proteins Membrane defects Example: cystic fibrosis Procaryotes Eukaryotes Organelles Cytoskeleton Microfilaments Microtubules Nucleus Nuclear envelope Nuclear pores Chromosomes Ribosomes Endoplasmic reticulum (ER): rough and smooth Vesicles Golgi complex Lysosomes Mitochondria Oxidative metabolism Mitochondrial DNA Chloroplasts Centrioles Flagella and cilia Vacuoles Cell walls Diffusion and osmosis Concentration gradient Net movement of molecules from an area of higher to lower concentration Osmotic pressure Hypertonic Hypotonic Isotonic Endocytosis Phagocytosis Pinocytosis Exocytosis Facilitated diffusion Active transport Sodium potassium pump Proton pump Chapter 5 and 6 Energy and Life / How Cells Acquire Energy Chemical reaction Substrates Products Activation energy Biology 1 106736530 5 Enzymes Specificity Binding site and active site Catalyst Cofactor or coenzyme, example: NAD+ Factors affecting enzymes pH Temperature Biochemical pathway Adenosine triphosphate (ATP) Photosynthesis = energy from the sun 6CO2 + 12H2O + light energy C6H12O6 + 6H2O + 6O2 Chloroplast Grana Thylakoids Stroma Photons Electromagnetic spectrum Visible light UV light Pigments Chlorophyll in photosystem networks Chlorophyll a and b Carotenoids Light dependent reactions occurs inside thylakoid: 3 steps 1. Primary photo event occurs where light is captured by chlorophyll by an excited electron. H2O is split to give off O2 , H+, and a free electron. 2. Electron Transport. The excited electron moves along an electron transport chain. NADPH is produced that provides reducing power to make sugars and other organic compounds. 3. Chemiosmosis. ATP is made with H+ pump providing energy to build organic molecules. Light independent reactions (Calvin cycle, C3 photosynthesis) occur inside stroma surrounding the thylakoid CO2 is fixed into organic molecules ATP provides energy to power reactions. NADPH provides hydrogens with energetic electrons to bind to carbon C3 photosynthesis = Calvin cycle Circular set of reactions. Some molecules are siphoned off to make sugars. Some molecules reform the 5C sugar to restart the cycle. C4 photosynthesis- common in Taft Photorespiration occurs in excessive heat and interferes with C 3 photosynthesis to fix CO2. C4 concentrates CO2 in the leaf so the Calvin cycle can continue but is less efficient than C3. Cellular respiration = energy from organic chemicals Aerobic respiration C6H12O6 + 6O2 6CO2 + 6H2O + energy (heat, ATP) Biochemical pathway NAD+ = coenzyme, an electron carrier 2 steps in cellular respiration Step 1:Glycolysis in the cytoplasm O2 not required 2 Pyruvate molecules, 2 ATP, 2 NADH (2 ATP) produced Step 2: Oxidation in the mitochondria Biology 1 106736530 6 Krebs cycle (2 cycles needed for each glucose molecule) Electron carriers (NADH and FADH2) and 2 ATP are produced 3 CO2 produced as waste Electron transport system (ETS) O2 used as electron acceptor with protons (H+) to form water (H2O) ATP produced at H+ transport pump (channel protein) Grand total of 36 ATP molecules from 1 glucose molecule Fermentation- inefficient but used when O2 not available Pyruvate acts as electron acceptor in absence of O 2 Lactic acid fermentation occurs in muscle Ethanol and CO2 release occur in yeast fermentation Exam I Chapters 1,3-6 Biology 1 106736530 7 Unit 4 Cell Division 4.1 List essential life processes that depend on mitotic production of new cells identical to the parent cell. 4.2 Explain the role mitosis plays in the cell cycle. 4.3 Explain one strategy for curing cancer. 4.4 Discuss the two cell divisions of meiosis and their effect on the chromosome number. 4.5 Describe the importance of these 2 events of meiosis I and II: 1) crossing over, and 2) independent assortment. 4.6 Define the following terms: Chapter 7 Cell Division Binary fission of prokaryotes Mitosis of eukaryotes Somatic cells Chromosomes Homologous chromosomes Diploid cells Sister chromatids Centromere Cell cycle 3 Phases Interphase M- mitosis C- Cytokinesis Stages of mitosis Cytokinesis Programmed cell death Example: Developmental cell death Cancer = cell division out of control Tumor Metastases Strategies for cures Meiosis = 2 cell divisions to make cells for sexual reproduction Germ cells Chromosome number Diploid Haploid Sexual reproduction Gamete Zygote Meiosis reduces the number of chromosomes in gametes to half of parent cell. Meiosis cell division I separates the 2 parental versions of each chromosome Crossing over with DNA exchange occurs during synapsis in prophase I Creates 2 collage chromosomes containing parts of both parental versions. 2 parental chromosomes remain unchanged. Independent assortment of each parental chromosome versions occurs in anaphase I Possibilities = 2 to the power of the number of chromosome pairs Humans = 223 = 8,388,608 No replication of chromosomes occurs between Meiosis I and II, so a reduction in chromosome numbers occurs. Meiosis cell division II separates the 2 replicas of each parental chromosome 4 Gametes are produced, each haploid Random fertilization results in 223 x 223 = 70 trillion possible outcomes Generating diversity provides choices for survival in a changing environment Biology 1 106736530 8 Unit 5 Mendelian Genetics / Genes / Gene Technology / Genomics Chapter 8 General outcome: 5.0 The student should understand the principles of heredity as first worked out by Gregor Mendel and extended by others, both in regard to chromosome behavior and to the statistical ratios of traits among offspring. Specific learning outcomes: Upon successful completion of this unit, the students should be able to: 5.1 State Mendel's first principle of inheritance and give examples. 5.2 Define: dominant, recessive, allele, homozygous, heterozygous, genotype, phenotype, segregation, recombination. 5.3 Explain Mendel's trait ratios in terms of probability. 5.4 Recount Mendel's second law of inheritance, independent assortment, giving an example involving two traits. 5.5 Define and give examples of incomplete dominance and codominance. 5.6 Define and give examples of multiple allele inheritance. 5.7 Define and give examples of polygenic inheritance. 5.8 Explain Mendel's laws of segregation and independent assortment in terms of chromosome behavior during meiosis. 5.9 Explain the connection between meiosis and trisomy-21. 5.10 Give a chromosomal explanation of sex determination. 5.11 Analyze the genetics of sex-linked traits in terms of inheritance. 5.12 Give examples of chromosome mutations. 5.13 Define the following terms: Chapter 8 Heredity Mendel’s genetics Mendel’s experiments P(arental) generation True breeding F1 generation (first filial meaning ‘son’ or ‘daughter’) F2 generation (second filial) Genes Alleles Homozygous Heterozygous Dominant Recessive Phenotype Genotype Punnett square Probability Test cross Mendel’s laws of heredity I: Law of segregation: Only one allele for a trait can be carried in a gamete, and gametes combine randomly to form offspring. II: Law of independent assortment: Genes located on different chromosomes are inherited independently of one another. Epistasis Multiple alleles Codominant e.g. ABO blood groups Polygenic inheritance (continuous variation) Incomplete dominance Biology 1 106736530 9 Environmental effects Chromosomes Karyotype Aneuploidy Nondisjunction Monosomic Trisomic example: Down syndrome- chromosome 21 trisomy Sex chromosomes Autosomes Sex chromosomes X chromosome Y chromosome Abnormal numbers of sex chromosome Examples: Klinefelter syndrome = XXY male Turner’s syndrome = XO female Mutations Autosomal recessive, example: Sickle-cell anemia, Tay-Sachs disease Sex linkage inheritance = when a recessive gene is carried on the X chromosome Example: Hemophilia Pedigree Dominant autosomal, example: Huntington’s disease Genetic counseling Unit 6 Molecular Genetics Chapter 9, 10, 11 General Outcome: 6.0 The students should understand the chemical and physical structure of the gene and its operation in the synthesis of polypeptides. Specific Learning Outcomes: Upon successful completion of this unit, the students should be able to: 6.1 Chronicle the experimental evidence by key researchers leading to the Watson/Crick Model of DNA. 6.2 Chronicle the experimental evidence leading to an understanding of gene function. 6.3 Describe the Watson and Crick Model of DNA and the Central Dogma of molecular biology. 6.4 Describe DNA replication. 6.5 Define introns and exons. 6.6 Contrast an RNA nucleotide with one of DNA. 6.7 Describe the transcription of RNA. 6.8 Discuss the structure and function of tRNA, mRNA, and rRNA. 6.9 Define codon and grasp the significance of the tabulated genetic code. 6.10 Describe the role of the ribosome in translation of genetic information. 6.11 Discuss control of gene expression. 6.12 Define: 1) genetic mutation in terms of a point mutation, 2) mutagen. State examples of common mutagens. 6.13 Describe the 4 steps in transferring a gene from one organism to another. 6.14 Describe the role that restriction and ligase enzymes play in gene transfer. 6.15 Describe the uses of the polymerase chain reaction (PCR) in biology and society. 6.16 Describe the use of a Ti plasmid in agriculture for genetically altered plants. 6.17 Explain the benefits of a glycophosate resistance gene. 6.18 Describe the use of piggyback vaccines in treating genetic defects. 6.19 Give examples of genetically engineered drugs. 6.20 Describe the goal of the Human Genome Project. 6.21 Define the following terms: Chapter 9 Transformation: Griffith (1928) and Avery(1944) experiments Biology 1 106736530 10 Hershey-Chase experiment (1952): Radioactive isotopes in protein vs DNA DNA structure Double helix: The Watson and Crick model (1953) Nucleotides Purines- two large bases: adenine, guanine Pyrimidines- two small bases: thymine, cytosine Base pairing: A-T G-C Chargaff’s rule states amount of A = T, G = C DNA replication (semiconservative) DNA polymerase adds complementary nucleotides at each position DNA repair mechanisms limit errors Central dogma of gene expression: from gene to protein Gene expression: DNA RNA protein Transcription: creating a functional copy of a gene in the form of mRNA Messenger RNA (mRNA) RNA polymerase: reads a gene to make a complementary RNA version Note: T(hymine) is replaced with U(racil) Genetic code Codon- a three base sequence corresponding to an amino acid. Genetic code dictionary: universal for all organisms Translation: making protein 1. mRNA binds to a ribosome at its starting point and acts as a template for tRNA molecules Ribosomal RNA (rRNA) and ribosomal proteins act as an anchor for the mRNA 2. Transfer RNA (tRNA) with attached amino acids are attracted to complementary codons on mRNA Anticodon- a three base sequence on tRNA complementary to a codon 3. As each codon is read by a tRNA, the amino acid is released and attached to the previous amino acid to form a growing protein chain. The used tRNA returns to the cytoplasm to get another amino acid. 4. The process stops when the stop codon is reached by the ribosome and the protein is released into the cell. Gene expression- some genes are turned on, some turned off Eukaryotic gene architecture Exons- DNA containing coding information for amino acids Introns- ‘extra’ DNA that must be removed from the mRNA (90% of human genome) Mutations Point mutations (Single Nucleotide Polymorphism or SNP) Mutagens Examples: chemicals: cigarette tar, diesel exhaust, pesticides, stomach acid UV radiation (other radiation too) Chapter 10 Genetic engineering: moving genes from one organism to another Gene transfer- 4 steps 1. Cleaving DNA Restriction enzymes cuts DNA Gel electrophoresis separates cut fragments Sticky ends bind complementary base pairs of other DNA fragments 2. Recombining DNA Ligase enzymes joins cut fragments into the DNA vectors Vector is used to carry new gene into an organism Plasmid Viral DNA 3. Cloning is used to make copies of the inserted gene Clone library: a collection of an organism’s genome in fragment form inserted into vectors 4. Screening for a gene of interest Probe Polymerase chain reaction (PCR) Biology 1 106736530 11 The PCR cycle 1. Denature with heat to separate DNA strands 2. Primers added to start the sequence 3. Primers extended with heat stable DNA polymerase (Taq polymerase) DNA ‘fingerprint’: amplify DNA with PCR, cut with restriction enzyme, separate fragments with electric field Genetic engineering examples Glyphosate resistance gene for crop plants Pest resistance with the Bt gene Genetically modified (GM) ‘Golden rice’ with iron and Vit A enhancements Genetically engineered drugs Insulin to control blood sugar Anticoagulants to dissolve blood clots Factor VIII to clot blood Gene therapy example: Cystic fibrosis Reproductive Cloning: an asexual form of reproduction Chapter 11 Genomics Genome Human genome Human Genome Project DNA sequencing Genes code for proteins (30-40K) and sometimes can rearrange exon transcripts to form alternate proteins Noncoding regions of DNA make up 99% of human genome Variation within the genome Single nucleotide polymorphisms (SNPs) Markers of different alleles that can be used to find disease producing genes Gene exchanges have occurred laterally between very unlike species, ex. Humans and bacteria Plant hybrids can duplicate whole genomes to form a new species with double sets of chromosomes (polyploidy) Gene microarrays can measure all the SNPs of an individual: a profile of your inherited traits. EXAM 2 CHAPS 7, 8, 9, 10, 11 Biology 1 106736530 12 Unit 7 Evolution and Natural Selection Chapter 2 , 13 General Outcome: 7.0 The student should understand the basic mechanics of evolution and cite evidence that supports its existence. Specific Learning Outcomes: Upon successful completion of this unit, the students should be able to: 7.1 Chronicle the events of Darwin’s work that led to his published account of evolution “On the Origin of Species”. 7.2 Explain the evidence that supports the theory of evolution. 7.3 Describe the mechanism of how organisms evolve using natural selection. 7.4 Describe how populations evolve using microevolution, using the concepts of allele frequencies, HardyWeinberg equilibrium, and the 3 forms of natural selection (directional, stabilizing, disruptive). 7.5 Explain how the 2 examples, sickle cell anemia and industrial melanism, demonstrate how organisms have acquired adaptations that supports the concept of evolution in the world at large. 7.6 Explain how species may form during macroevolution using isolating mechanisms that reinforce differences. 7.7 Define the following terms: Chapter 2 Evolution Natural selection Charles Darwin HMS Beagle, Galapagos Islands and Darwin’s finches Thomas Malthus: Essay on the Principal of Population, 1798 On the Origin of Species, 1859 Darwin’s major evidence: Fossils: similar to living species, in strata with progressive change Geographic distribution of species: different organisms for each continent Oceanic island species: local species show strong relatedness and resemble nearest mainland species. Chapter 13 Fossil record Relative dating: position in rock strata Absolute dating: Radioisotopic dating (see Chapter 3, p 45) Half-life 50% 14C decays to 14N over 5,600 years 50% 40K decays to 40Ar over 1.3 billion years Fossil record shows progressive change Examples: whales, oysters, titanotheres Weaknesses: Gaps in the record, need a hard substance to mineralize Molecular record Molecular clock = the rate of DNA change due to mutation over time More distantly related species should have accumulated more genetic changes than more recently diverged species. Example: hemoglobin, Cytochrome c Molecular family tree Example: globin gene Structural evidence Embryo development shows previous structures Homologous structures show evolutionary paths Analogous structures results of parallel evolution Vestigial structures are structures no longer in use Macroevolution Microevolution Biology 1 106736530 13 Hardy-Weinberg equilibrium lets us predict allele frequencies for two alleles p and q p + q = 1, the sum of the 2 alleles must be 100%, and (p + q)2 = p2 (homozygous) + 2 pq (heterozygous) + q2 (homozygous) = 1 Allele frequencies Hardy-Weinberg rule predicts no change in allele frequency when: 1. The size of population is large. 2. Individuals mate at random. 3. All alleles are replaced equally (no natural selection) 4. No input of new alleles from any source (no mutation or immigration) 5 Agents of evolutionary change 1. Mutation 2. Migration 3. Genetic drift Founder effect Bottleneck effect 4. Nonrandom mating Inbreeding 5. Selection Natural Artificial These all cause the allele frequencies to change over time for a given trait. 3 Forms of selection 1. Directional 2. Stabilizing 3. Disruptive Adaptation (evolution at work) Sickle-cell anemia: people respond to deadly malaria infections Industrial melanism Example: Biston betularia,the peppered moth Species concept 4 steps in species formation 1. Local population adapts to environment. 2. Ecological races form. 3. Natural selection reinforces differences, successful hybrids are rare. Reproductive isolation mechanisms develop. Prezygotic Geographic isolation Ecological isolation Temporal isolation Behavioral isolation Mechanical isolation Prevention of gamete formation Postzygotic Hybrid embryo does not develop Hybrid adults do not survive or are infertile 4. Ecological races become incapable of interbreeding and are considered separate species. Biology 1 106736530 14 Unit 8. Our Living Environment 2, 31, 32, 34 General Outcome: 8.0 The students should be able to understand 1) how organisms interact with their environment and 2) the effects of human and natural events on the environment’s ability to support the diversity of life necessary for a stable ecosystem. Specific Learning Outcomes: Upon successful completion of this unit, the students should be able to: 8.1 Define the term ecosystem and explain how habitat and community form a self sustaining ecosystem. 8.2 Explain how energy drives the growth and interaction of organisms within ecosystems. 8.3 Discuss the flow of energy through the trophic (feeding) levels of an ecosystem and how disturbing a food web may alter the energy flow. 8.4 Explain how the major materials of life cycle in an ecosystem: water, carbon, nitrogen, and phosphorus. 8.5 Describe the major types of ecosystems and how the weather is responsible for the diversity of ecosystems on the earth. 8.6 Draw and label a population curve. 8.7 List the density dependent and independent factors that effect population growth. 8.8 Give examples of how coevolution affects ecosystems. 8.9 Explain aposematic coloration, cryptic coloration, Batesian mimicry, and Mullerian mimicry. 8.10 List the factors that promote biodiversity and stability in an ecosystem. 8.11 List and explain how humans disrupt ecosystems and ways humans can modify ecosystems successfully. 8.12 Discuss the global changes affecting the world ecosystem caused by chemical pollution, plastics, agricultural chemicals, water pollution, acid rain, ozone, and the greenhouse effect. 8.13 Discuss these four key areas of human intervention that will reduce global ecological changes: reduce pollution, develop other sources of energy, preserving nonreplaceable resources, and curbing population growth. 8.14 List five steps that can be successfully used to solve environmental problems. 8.15 Define the following terms: Chapter 2 Chapter 31 Ecology Ecosystem Community Habitat Physical habitat Biosphere Energy flow in ecosystems Sun energy: the source of all life on earth Trophic levels Producers = Level 1 Consumers = Level 2 Primary consumers (herbivores) Carnivores = Level 3 Detritivores (decomposers) = specialized class of consumers for organic waste and dead bodies Tertiary consumers = Level 4 Omnivores = Levels 2, 3, and 4 Food chain Food web Primary productivity Net primary productivity Biomass Trophic efficiency Energy stored in a trophic level is about 10% of the level below. Biology 1 106736530 15 Cycling of materials in ecosystems Water cycle Evaporation Transpiration Groundwater Carbon cycle Respiration Combustion Erosion Nitrogen cycle Nitrogen fixation Phosphorus cycle Eutrophication Major ecosystems Weather shapes ecosystems Latitude vs. rainfall Rain shadow Ocean currents Ocean ecosystems Shallow water Intertidal region Estuaries Surface water Plankton Deep water Freshwater ecosystems Lakes have three zones: shallow edge, open water surface, deep without light Thermal stratification Spring and fall overturn Eutrophic lakes Oligotrophic lakes Land ecosystems = biome Influenced by Climate Continental Drift Glaciation Types Tropical rainforest Savannas: dry tropical grasslands Deserts Temperate Grasslands (prairies) Deciduous forests (hardwood forest) Taiga (northern coniferous forest) Tundra (cold boggy plains) Other less widely distributed local types to Kern County Chaparral Semidesert Riparian Chapter 32, Chapter 2 Population dynamics Population Size Density Dispersion Random Clumped Biology 1 106736530 16 Uniform Population Growth Exponential growth model = growth without limits Logistic growth model = growth with limits, sigmoid curve Lag phase growth Exponential growth Carrying capacity limit steady state Death phase growth Biotic potential (population growth) = r r = (birth rate + immigrants) – (death rate + emigration) Carrying capacity (K) Population growth rate = r N ( K-N) N= current population K r selected vs K selected populations Example: r selected = aphids, cockroach, mice; k selected = whale, redwood tree Density dependent effects Density independent effects Mortality rate and survivorship Age distribution curve Survivorship curves: type I, II, III Coevolution and ecosystems Plants vs. herbivores Symbiotic relationships Commensalism Mutualism Parasitism Niche Fundamental Realized Niche division among sympatric species Competitive exclusion Resource partitioning Character displacement Predation-prey Cyclical pattern of populations Animal defenses Aposematic coloration- ‘I’m warning you’ Mullerian mimicry Batesian mimicry Cryptic coloration Ecosystems over time Disruption = beginning (ex., volcano) or change (ex., fire) Succession Primary Secondary Climax community Biodiversity Keystone species Ecosystem species richness Size Latitude Length of growing season Climatic stability Chapter 34 Pollution Chemical Biology 1 106736530 17 Plastics Agriculture Biological magnification of toxins Water Acid rain Ozone CFC’s Greenhouse effect: affects the entire biosphere Greenhouse gases: CO2, methane, nitrogen oxides, CFC’s Global warming Reducing pollution Antipollution laws Pollution taxes/credits Other sources of energy Nuclear Wind Solar Preserving nonreplaceable resources Topsoil Groundwater Biodiversity Disruption of ecosystems Physical habitat destruction Example: Lowering species diversity with clear-cutting Reducing competition Example: Exotic species introduction upsets balance Solution: Minimizing ecosystem damage by 1) limit physical disruption, 2) maintain biodiversity, 3) maintain competition Human population growth curve Population pyramid graphs (chapter 2) 6 billion in 1999 with 80 million more each year 5 Steps to solving environmental problems Scientific assessment and modeling Risk analysis- predict consequences action and inaction Public education- problem, solutions, costs Political education- public demand action from political process Follow-through (monitor effectiveness of solution, learn by doing) EXAM 3 CHAPS. 2, 30, 31, 32, 34 Biology 1 106736530 18 Unit 9. Single Celled Life and Simple Multicellular Life Chapter 14, 15, 16, 17 General Outcome: 9.0 The students should be able to understand how organisms are classified and the major characteristics of each major group (kingdom, phylum, etc). Specific Learning Outcomes: Upon successful completion of this unit, the students should be able to: 9.1 Explain the importance of Carolus Linnaeus’s development of the binomial classification system. 9.2 Cite the descending order of hierarchy taxons: kingdom, phylum, class, order, family, genus, and species. 9.3 Explain why the biological species concept works well for animals but not for other kingdoms. 9.4 Explain how ecological races develop. 9.5 Contrast the cladistic and traditional taxonomy approach to establishing phylogenies among groups of organisms. 9.6 Compare a prokaryotic to a eukaryotic cell. 9.7 Name the key differences between Archeabacteria and Eubacteria. 9.8 Explain the 5 steps of viral replication in a host cell. 9.9 Explain the theory of endosymbiosis and its relationship to the appearance of the eukaryotic cell. 9.10 Compare and contrast the 3 types of sexual life cycle among eukaryotes. 9.11 Be able to cite the characteristics of the major phyla of protists and give an example of each. 9.12 Explain the 2 main characteristics of complex multicellular organisms: cell specialization and intercell coordination. 9.13 Be able to cite the characteristics of the major phyla of fungi and give an example of each. 9.14 Explain the symbiotic relationship called mycorrhizae. 9.15 Define the following terms: Chapter 14 Levels of classification Polynomial system Binomial system Carolus Linnaeus (1750’s) Scientific name vs. common names Genus and species Taxonomy Taxons Higher categories Kingdom Phylum Class Order Family Genus Species Examples: Human being, honeybee, red oak A way to remember: ‘Kindly pay cash or furnish good security’ 3 Domains over the kingdoms Archeabacteria Eubacteria Eukarya The basic biological unit: the species The biological species concept This concept assumes that outcrossing occurs: works well for animals. Biology 1 106736530 19 Asexual reproduction dominates in bacteria and many protists, fungi, and plants. Hybrids can be common in plants and some animal (fish form fertile hybrids). Humans are putting genes from one organism into another. What are these? Ecological races or ecotypes Isolating mechanisms How many species are there? Measuring species diversity is difficult. 1.5 million named, 10 million total on earth? Phylogeny: the evolutionary history of life on earth. Constructing the phylogeny of a group of organisms: a family tree. Cladistics uses derived characteristics to evaluate relatedness and lineage Advantage: an objective exercise once the criteria are defined. Traditional taxonomy: weighting the characteristics by biological significance. Example: Flight gives birds (Aves) its own class vs. shared class with reptiles in cladistic approach. Chapter 15 Bacteria and viruses Bacteria: the oldest organisms on earth, 3.5 billion years old. Bacteria existed for 2 billion years before eukaryotes first appeared 1.5 billion years ago. Most abundant life form on earth (2.5 billion per tablespoon of rich earth). Role of bacteria Recycle minerals. Introduced oxygen into the earth’s atmosphere. Cause plant and animal diseases. Provide nutrients in animal digestive tracts. Bacterial structure Prokaryote: a simple design Single, circular DNA chromosome. No nucleus. Cell wall Capsule Flagella Pili Endospores Reproduction: binary fission Conjugation: passing plasmids Comparing prokaryotes and eukaryotes Characteristic Internal compartmentalization Cell Size Unicellularity Prokaryote Little 1x Single celled Chromosomes Single circular in cytoplasm Cell division Metabolic diversity Binary fission Diverse among species Eukaryote Many membrane compartments and organelles 10x Multicellular with integrated activities Many in a nucleus wound around protein, linear Mitosis with all its apparatus Similar among species Archeabacteria vs. eubacteria: the 2 domains of bacteria. Differences in cell wall, plasma membrane, gene translation, gene architecture. Archeabacteria: ancient bacteria Survivors live in extreme environments: strict anaerobes (methanogens), thermoacidophiles. Eubacteria: modern bacteria Autotrophs: Photosynthesis Example: Cyanobacteria Helped produce the oxygen atmosphere we depend on. Biology 1 106736530 20 Fix nitrogen in heterocysts. Only a few bacteria fix nitrogen. Heterotrophs: consume organic molecules. Decomposition: Breakdown organic molecules. Disease producers Mycobacterium tuberculosis- TB, tuberculosis Yersina pestis- the black plague Vibrio cholera- cholera Viruses: simpler than bacteria The ‘parasitic chemical’ concept Infective RNA or DNA Viral Structure Capsid: protective protein sheath in geometric shapes Envelope: some viruses have a membrane like covering. Viral replication 1. Attachment: virus attaches to a specific cell membrane receptor (protein, glycoprotein). 2. Penetration: Binding triggers endocytosis with release of viral nucleic acid. 3. Biosynthesis: virus nucleic acid directs synthesis of viral components by host cell. 4. Maturation: viral components assemble into virus particles. 5. Release: host cell lyses and new virus in released to infect surrounding cells. Example: HIV (human immunodeficiency virus), an RNA virus Attaches to CD4 receptor, reverse transcriptase makes DNA Latency Example: Oral and genital herpes simplex virus, chickenpox then shingles Human disease producer examples Influenza- changes protein coat often Hanta virus- acute respiratory disease, rodent vector, located in rural California. HPV (human papilloma virus)- sexually transmitted cause of cancer Prion (proteinaceous infectious particle) disease Transmissible spongiform encephalopathies (TSEs) Example: infectious protein in beef causing ‘mad cow disease’ Chapter 16 Endosymbiosis Theory of endosymbiosis as origin of eukaryotes Living example: Pelomyxa palustris, primitive amoeba-like organism that lacks mitochondria, chloroplasts, and mitosis. Resembles descendants of the non photosynthetic archaebacteria more than eubacteria. Membrane bound organelles of endosymbiotic origin Mitochondria: allows oxidative metabolism Cholorplasts: 3 types red algae, green algae (similar to plants), brown algae Both have a piece of circular DNA Evolution of sex in eukaryotes: sexual reproduction appears in eukaryotes Sexual reproduction Two parents, diploid (or multiploid) Haploid gametes made by meiosis combine to form diploid zygote Two copies of each chromosome in each individual Asexual reproduction Many eukaryotes use asexual reproduction for most of their life Parthenogenesis: unfertilized egg undergoes mitosis (without cell division) forming diploid cells Develops as zygote but same genes as parent Example: bees, females are fertilized, males are unfertilized Common in insects, some lizards, fish, amphibians Self-fertilization: a special form of sexual reproduction in a single individual Examples: plants (pea plant) and some fish Same genes as parent but with crossing over and independent assortment Biology 1 106736530 21 Sex evolved as a way to repair double stranded chromosome damage Sexual reproduction shuffles genes quickly to create genetically diverse populations Genetic diversity is the raw material of evolution Natural selection can defeat species that lack diversity Sexual life cycles: 3 major types 1. Zygotic meiosis. The zygote is the only diploid cell. Haploid cells occupy the major portion of the life cycle. Example: algae 2. Gametic meiosis. Diploid zygote occupies most of the life cycle with haploid gametes a small portion of organism. Example: animal (humans) 3. Sporic meiosis. A regular alternation of generation occurs between a haploid and diploid generation. Example: the primitive plants such as a fern Multicellularity on a simple scale when an organism is composed of many cells, permanently. Colonial organism: little or no integration of activities Example: Volvox Aggregations of organisms: cells come together transiently and then separate to live as single cells Example: slime molds Simple multicellular organisms: many cells that interact and coordination activities Example: Brown algae Kingdom Protista: a catchall kingdom of diverse eukaryotes that are not plants, animals, or fungi. Mostly single cells but some are simple multicellular forms No complex tissues or specialized organs Many have flagella for movement 15 separate phyla with diverse characteristics Evolutionary relationships are not clear. Grouped by shared characteristics, primarily mode of transportation, photosynthesis, and metabolism. 5 major groups of protists with selected examples of each group: 1. Heterotrophs with No Permanent Locomotor Apparatus, the Sarcodina. Amoebas and forams that move with pseudopods or cytoplasmic streaming Amoebas: phylum Rhizopoda Movement with pseudopodia Reproduction: asexual fission, no sex 2. Heterotrophs with Flagella Phylum Sarcomastigophora, the zoomastigotes, several thousand species Ancestor to all animals via the sponges. Heterotrophic, unicellular, movement with flagella Examples: Trypanosomes: insect to human blood pathogens causing sleeping sickness. Symbiotic gut flora in termites that digest wood. 3. Nonmotile Spore-formers Phylum Apicomplexa (Sporozoa), the sporozoans: nonmotile, spore forming, unicellular parasites of animals. 3900 species Complex life cycles of alternating generations Haploid cells with fast mitotic growth to increase infection. Some become gametes and join to become a diploid cell. Diploid cells that become spores, the oocyst that resists environmental challenges. Meiosis within the oocyst produces haploid spores that are infective. Example: Plasmodium = malaria. Spread person to person by mosquitoes. 4. Photosynthetic Protists Phylum Pyrrhophyta, 1000 species Mostly marine, may be luminous producing twinkling light Red tides are blooms of dinoflagellates that produce toxins Unusual shapes with a stiff cellulose coat often with silica (SiO2) Biology 1 106736530 22 Phylum Chrysophyta, 11,500 species Diatoms: photosynthetic unicellular protists with double shell of silica Both radial and bilateral symmetry Fossil diatoms = diatomaceous earth. Abrasive used in products, e.g. toothpaste. Algae: 3 kinds of photosynthetic protists Green algae: phylum Chlorophyta, 7000 species Ancestor to modern plants Usually mobile with flagella in aquatic environment but can be in soil or on trees trunks Usually unicellular but can have multicellular forms Example: Volvox Red algae: phylum Rhodaphyta, 4000 species Red pigment (phycobilins) Mostly multicellular with interwoven filaments and live in the sea. No flagella. Brown algae: phylum Phaeophyta, 1500 species Multicellular, photosynthetic, often large and fast growing, mostly in marine environment Giant kelp- large conspicuous seaweed with flat blades off California coast, up to 100 meters long. 5. Heterotrophs with Restricted Motility Molds: heterotrophs with restricted movement Phylum Oomycota: water molds called rusts and mildews, 580 species Parasitize living organisms or feed on dead organic matter Example: Phytophthora infestans that causes late potato blight. Irish potato famine of 1845-47. Chapter 17 Complex multicellular organisms Kingdom Fungi Kingdom Plantae Kingdom Animalia Characteristics of multicellular organisms 1. Cell specialization Development 2. Intercell coordination Kingdom Fungi Characteristics (and differences from plants) Heterotrophic Cell walls made of chitin Nuclear mitosis- a special type of mitosis that occurs within the nucleus Fungi have filamentous bodies Mycelium Hyphae with or without septa Cytoplasmic streaming between most cells All parts of the mycelia are metabolically active The mushroom structure found in some fungi are only a temporary reproductive structure Reproduction Primarily asexual by spore production Sexual: non motile gametes join between different mating types Nutrients absorbed from environment by external digestion Digestive enzymes secreted into local environment Cellulose can be digested (see on dead trees) Cause rot, decay, food spoilage, diseases of plants and animals. Some predators. Kinds of fungus: 73,000 species, 400 million years old, differ by sexual reproduction structures Phylum Zygomycota , 1050 species Biology 1 106736530 23 Sexual reproduction: an environmentally resistant zygosporangium forms similar to a zygote when two nuclei join. Examples: Rhizopus, black bread mold Phylum Ascomycetes, 32,000 species Sexual reproduction: 2 different hyphae fuse to form an ascus with diploid cells Examples: yeast, Dutch elm disease, morels, truffles Phylum Basidiomycota, 22,000 species Sexual reproduction: sexual reproduction is dominant, occurs in basidium on underside of mushroom cap Examples: mushrooms, puffballs, toadstools, shelf fungus Imperfect fungi, 17,000 species Sexual reproduction: unknown, probably mostly ascomycetes Example: dermatophytes (athlete’s foot, ringworm), Valley fever Fungal associations Lichens: a symbiotic association between a fungus (usually an ascomycete) and a photosynthetic partner (cyanobacteria or green algae) The photosynthesizer makes organic molecules and may fix nitrogen The fungus dissolves mineral in rock and provides nutrients. Lichens are extremely sensitive to pollution. Examples: rock lichen, tree lichen Mycorrhizae: a symbiotic relationship with plant roots Fungi release mineral nutrients (e.g., phosphorus) from the soil. Plant supplies organic compounds to fungus. Helped plants colonize the primitive infertile soils 2 types Endomycorrhizae: about 30 species of zygomycetes penetrates plant root cells of > 200,000 plant species Ectomycorrhizae: wrap around but do not penetrate root cells of about 10,000 Species of plants. Usually a one to one relationship with a plant and fungus species. Most are basidiomycetes with some ascomycetes. Unit 10. Plant Life Chapters 18, 19, 20, 21 General Outcome: 10.0 The students should be able to understand how organisms are classified and the major characteristics of each major group (kingdom, phylum, etc). 10.1 Describe the adaptations used by plants for terrestrial living. 10.2 Explain the phrase ‘alternation of generations’ in regards to plant reproduction. 10.3 Cite 4 reasons why seeds improved the adaptation of plants to terrestrial living. 10.4 Explain how water can move up a tall tree from the roots to the leaves. 10.5 Explain how carbohydrates move from the leaves to the roots of plants. 10.6 Explain how plants control water loss. 10.7 Define the following terms: Chapter 18 Life appeared on land about 440 MYA UV radiation prevented life on land until the O2, O3 atmosphere formed from photosynthetic organism emissions produced the oceans. Plants and fungi first invaded the land. Terrestrial autotrophs, most dominant life form on land surface, 288,700 species Biology 1 106736530 24 Moving from water to land (terrestrial living) Plants evolved from green algae The earliest plants formed symbiotic relationships with fungi = mycorrhizae to gain access to minerals of life from rocky soils. Nitrogen (protein), calcium (plant cell wall glue), potassium (regulate water loss/retention), phosphorus (nucleic acids/ATP), magnesium (chlorophyll), sulfur (amino acid cysteine) Plants needed to conserve water Cuticle Stomata Reproducing on land without a watery environment Spores Seeds Generalized plant life cycle Alternation of generations Sporophyte = diploid plant generation producing haploid spores by meiosis Gametophyte = haploid plant generation producing haploid spores by mitosis Early plants were largely gametophyte tissue, modern plants largely sporophyte tissue Evolution of vascular system in plants on land (about 410 MYA) Plants developed vascular plumbing Take advantage of land away from streamside Provided ability to grow tall A network of tubing from root tip to leaf tip Vascular plants are the most successful. 9 of 12 phyla. Types of plants Plants with no vascular systems: 2 phyla Phylum Hepaticophyta: Liverworts 8,500 species Phylum Anthocerophyta: Hornworts 100 species Both are inconspicuous and live in damp, shady places Plants with simple vascular systems: single phylum Phylum Bryophyta: Mosses, 12,000 species Strands of soft cells conduct water and carbohydrates. Inconspicuous plants found in moist places Plants with well developed vascular systems: 9 phyla with 250,000 species Vascular plant features Dominant sporophyte (diploid tissue) Specialized, reinforced bundles of water and nutrient conducting tissue Specialized body form: roots, stems (shoots), leaves Seedless vascular plants: 4 of 9 modern phyla Free-living sperm require water for fertilization Phylum Pterophyta: Ferns, the most abundant phyla with 12,000 species Can be small to tall (24 m) Life cycle of ferns Independent sporophyte (large) and gametophyte (small) Fronds: vertical leaves with spore forming structures on backside of frond Plants with seeds: a key evolutionary advance for land domination Seeds are plant embryos surrounded by a durable, watertight, cover that improves survival on land. 1. Dispersal of seeds to new habitats possible 2. Dormancy allows plants to postpone development until conditions are favorable. 3. Germination (the reinitiation of growth) permits development to be synchronized the seasons. 4. Nourishment is provided to the embryo by endosperm until the seedling is established Gymnosperms: the first seed plants, 4 phyla Biology 1 106736530 25 Example Phylum Coniferophyta: 600 species, seed produced in cones Examples: all in California Tallest tree = Sequoia sempervirens, coastal redwood Largest tree = Sequoidendron gigantea, mountian sequoia redwood Oldest tree = bristlecone pine All have cones, so called conifers Needle like leaves are adaptation to avoid water loss Life cycle Seed cones = eggs, pollen cones = sperm ( yellow pollen grains) Profuse amount of pollen produced with wind pollination Zygote forms new sporophyte that arrests as a seed Angiosperms: the most successful plants with 250,000 species Phylum Anthophyta: the flowering plants Half the calories humans consume originate in 3 species: rice, corn and wheat Angiosperms solved the problem of having efficient sex when anchored to the ground (to gain nutrients): induce insects and other animals to participate as a third party! Different kinds of flowers Yellow or blue sweet smelling = bee pollinators 20,000 bee species Long, slender floral tubes, nectar, with platform = butterfly with long proboscis White, heavily scented flowers = night time pollinators (moths) Red flowers, little smell = humming birds, sun birds (invisible to insects) Small greenish, odorless = wind pollinators (oak, birch, grasses) Seed dispersal by fruit Fruit = mature ripened ovary with seeds surrounded by a carpal Fruits taste good to animals so they will eat them and disperse the seeds to far away places. Chapter 18, 19, 20 Structure and function of vascular plant tissues Basic organization Root: below ground portion, penetrates ground and anchors plant, absorbs water and mineral ions Shoot: above ground portion Stems: framework for positioning leaves Leaves: most photosynthesis (food production) occurs here Most chlorophyll containing cells located here (green color) Combination of vascular bundle tissue and photosynthetic parenchyma Cells with large open areas for gas exchange Cuticle: waxy layer over epidermis toward sunny side Stomata: open close valves on leaf bottom All higher vascular plants share this theme: rose, pine, cactus Conducting tissue: elongated cells stacked end to end Phloem Conduct carbohydrates away from leaves to other parts of plant Xylem Conduct water and minerals up from the roots Water movement and regulation Transpiration: Water reaches leaves from roots and exits through the stomata (90%) Biology 1 106736530 26 Cohesion-Adhesion-Tension theory Polar water molecules are ‘sticky’ and cling to each other and capillary structure of vascular tissue Atmospheric pressure pushes water up the plant tubing 10.4 meters Evaporation from leaves creates vacuum and water rises above 10.4 m Regulation of stomata opening and closing (K+ important ion) Must be open for CO2 entry and O2 exit Must be closed when under water stress Turgid (high water content) guard cells are open Usually open in day and close at night Leaf size Carbohydrate transport Sugars (mainly sucrose) move from the high concentration (leaf) to low concentration (root) for storage. Water moves from high concentration (roots) to lower concentration (leaves) Translocation of sugar in phloem due to mass flow developed by osmosis Growth Primary growth Apical meristem: growth plate at tip of plant creates taller growth Early vascular plants (lycophyte trees, ferns) only had a primary meristem so were tall and slender (10 to 35 meters). Age of Coal- these plants formed the fossil fuels we use today Secondary growth Lateral meristem: cylinders of growth tissue around the plant to create wider shoots and roots Vascular cambium Wood with growth rings Large light rings = spring-summer growth Thin, Dark rings = winter-fall growth Cork cambium Secondary growth appeared about 380 MYA Thick-trunked and tall trees of today, 120 m high, 11 meters diameter (redwood) The flower of angiosperms Reproductive organs with sophisticated pollination structures arranged in whorls 1. Sepals: modified leaves to protect flower as a bud, usually green 2. Petals: often vividly colored, forming the corolla 3. Stamens, slender, threadlike filaments with swollen terminal anthers where pollen develops 4. Carpel: a case that enclosed the egg cells. Ovary Stigma Plant hormones Auxin: regulates plants cell growth 2,4,-D synthetic auxin , kills by growth acceleration in broadleaf dicots 2,4,5-T – ‘agent orange’ herbicide. Contaminated with dioxin, an endocrine disrupter. Ethylene- ripens fruit. Example: green tomatoes CO2 has opposite effect Biology 1 106736530 27 Unit 11 Kingdom Animalia: Evolution and History of Animal Life Chapter 21 General Outcome: 11.0 The students should be able to understand the basic evolutionary progression of animal characteristics among the major animal phyla, discuss the major events of terrestrial vertebrate natural history, and chronicle the major events of human evolution. Specific Learning Outcomes: Upon successful completion of this unit, the students should be able to: Describe Discuss Define Chronicle Cite Characterize Explain Give example 11.1 Cite the 9 major evolutionary stages (innovations) represented in the 9 animal phyla presented. 11.2 Define the following terms and concepts: Chapter 21 Animal characteristics Multicellular heterotrophs No cells walls so cells are flexible allowing for rapid movement Reproduce by sexual reproduction: the production of haploid gametes with diploid zygote formation No alternation of haploid and diploid generations as in plants. Nonreproductive animal tissues are diploid with few exceptions Kingdom Animalia: 36 phyla Subkingdom Parazoa: no definite symmetry or tissues, 1 phylum Subkingdom Eumetazoa: definite shape and symmetry, tissues organized into organs, 35 phyla 9 major evolutionary stages (innovations) in animals 1. Multicellularity 2. Symmetry and tissues 3. Internal organs and bilateral symmetry 4. Body cavity 5. Coelom 6. Segmentation 7. Jointed appendages and exoskeleton 8. Deuterostome development and endoskeleton 9. Notochord Phylum Porifera: the Sponges, 5000 species, only member of Parazoa Key evolutionary advance: multicellularity No symmetry or tissues Filter feeder Evolved from a unicellular protist = choanoflagellates Phylum Cnidaria : Cnidarians, the jellyfish, hydra, sea anemones, corals Key evolutionary advance: symmetry and tissues Radial symmetry: parts arranged around a central axis Tissues: specialized cells that act as a unit Extracellular digestion in a gut cavity 3 embryonic tissues found in eumetazoans Ectoderm: skin and nervous tissue Mesoderm: muscles and skeleton (not in Cnidarians, appears in Platyhelminthes) Endoderm: digestive organs and intestine Carnivores with stinging cnidoyctes with a nematocyst (harpoon) 2 body forms possible: Medusae: the jellyfish, a free-floating, gelatinous, umbrella-shaped body Polyp: a sessile, cylindrical, pipe-shaped body Phylum Platyhelminthes: solid worms, 20,000 species, mostly parasites of animals Biology 1 106736530 28 Key evolutionary advance: internal organs and bilateral symmetry with a head Bilateral symmetry: a right and left half that are mirror images Dorsal, ventral, anterior, posterior Cephalization: a distinct head with specialized sensory organs Designed for active, mobile life Head in front to sense the environment for food, danger, etc. Internal organs develop from mesoderm Acoelomate: no body cavity with organs imbedded in a solid body No circulation systems so body must be flat to allow for diffusion. Example: Planaria, the common flat worm; Schistosoma, the blood fluke Phylum Nematoda: roundworms, 12,000 species Key evolutionary advance: body cavity Pseudocoelomate: a cavity (pseudocoel) located between the endoderm and mesoderm that has 3 advantages: Circulation: better circulation, larger bodies possible Movement: allows muscle driven body movement Organ function: organs not deformed by movement Nematodes are abundant in soil: millions per square meter Important plant parasites Phylum Mollusca: Mollusks, 110,000 species, mostly marine Key evolutionary advance: coelom, a body cavity completely enclosed in mesoderm Primary induction: the interaction of embryonic tissues to form complex organs Example: stomach Circulation system of vessels and heart muscle carries nutrients and wastes Body plan: head, central section with organs, foot Examples: 3 major classes Gastropods: snails and slugs Bivalves: clams, oysters, scallops Cephalopods: octopuses, squids Phylum Annelida: segmented annelid worms, 11,100 species, mostly marine Key evolutionary advance: segmentation, the building of a body from similar segments Repeating segments can act independently Segments can evolve for different functions Connections between segments Circulatory system delivers nutrients and removes wastes Nervous system links all segments to a central brain for coordinated activity Example: earthworm Body plan is a tube within a tube, a digestive tract within a coelom, surrounded by repeated segments of muscle. Segmentation underlies body organization of all advanced coelomate animals Phylum Arthropoda: Arthropods, the most abundant eukaryotes on earth (2/3 of all named species) Key evolutionary advances: jointed appendages and exoskeleton Jointed appendages: used for movement (legs, arms, wings), chewing (mouth parts) Exoskeleton: rigid outer skeleton of chitin Advantages: Muscle attachment, protection, prevent moisture loss Disadvantage: too heavy to support a large body Most arthropods about 1 mm long Body plan: coelom, segmented body, jointed appendages Head with chewing appendages, thorax with legs (wings), and abdomen formed from fused segments Tracheae: breathing tubes with spiracles (outer openings) Special senses: antennae, compound eyes Examples Chelicerates: arthropods without jaws, horseshoe crabs and extinct trilobites, arachnids Class Arachnida: spiders, ticks, mites, scorpions, daddy long legs Biology 1 106736530 29 Mandibulates: arthropods with jaws (mandibles) Subphylum Crustacea, the crustaceans, ‘insects of the sea’, 35,000 species Many segments with appendages resembling their annelid ancestors Appendages on abdomen and thorax, 2 pair of antennae Crabs, shrimps, lobsters, crayfish, barnacles, pillbugs, copepods Subphylum Uniramia: insects, centipedes, millipedes Class Insecta: the insects, most abundant eukaryotes by numbers of species and individuals Appendages on thorax, Malpighian tubes eliminate wastes Occupy almost every conceivable habitat on land and fresh water Examples: Termites, fleas, bees, butterflies and moths, body lice Phylum Echinodermata: Echinoderms, marine, mostly bottom dwelling animals. 6000 species. Key evolutionary advances: deuterostome development and endoskeleton Deuterostome embryonic development Identical cells in early embryo differentiate under influence of DNA Protostomes develop by cell position in embryo Anus develops from blastopore, not mouth Protostome mouth develops from blastopore Endoskeleton: ‘spiny skin’ Calcium rich plates form within a delicate skin Plates fuse as tough spiny layer as animals becomes an adult Acts much the same as endoskeleton Body plan Bilateral symmetry as larvae but radial as adult Water vascular system for locomotion: tiny tube feet Extend by closed sac under water pressure Pull foot back by muscle contraction Regeneration of a whole individual from a single arm Phylum Chordata: the chordates, vertebrates, tunicates, lancets. 50,300 species Key evolutionary advance: notochord 3 Characteristics of chordates 1. Notochord: long stiff rod along back of early chordates Muscles attached to rod for swimming movement 2. Nerve cord: dorsal nerve cord that distributes nerves to all parts of the body 3. Pharyngeal slits: slits behind mouth in a muscular tube (pharynx) connected to the digestive tube. (Relics of aquatic ancestry) Body plan Segmented with distinct blocks of muscle Most with true endoskeleton and jointed appendages Bone and cartilage skeleton: strong (calcium carbonate) but flexible (collagen) Most are vertebrates Backbone: notochord replaced by bony vertebral column with nerve cord inside Head: well differentiated head with a brain encased in a skull Exam 4 Chapters 14 – 21 Biology 1 106736530 30