WILD HORSE AND BURRO DESCRIPTIVE FACT SHEET EXPLAINING THEIR REAL PLACE IN NORTH AMERICA with references (January 26, 2011) By Craig C. Downer, Wildlife Ecologist, P.O. Box 456, Minden, NV 89423-0456 ccdowner@yahoo.com 1. Origin: The entire horse family Equidae, including the three major branches: zebras, asses, and caballine horses, have their origin and long-standing evolution in North America. This includes the many branches and side branches over the past 60-million years with no significant absence up to the present. The genus Equus, including the modern horse Equus caballus, originated upon this continent, the latter nearly two million years ago. The burro also has its roots in North America and some paleontologists consider it to be the same species that originally migrated over the Bering Straits from North America to Asia and thence to Central Asia and Africa. 2. Equids’ Ecological Contribution: Given the fact that equids have played such a major role in North America for so long a time, they have developed many mutualistic relationships with the plants and animals upon which they depend. The reason is simple: members of the horse family are a different type of herbivore from the prevalent ruminants in North America, be they cattle, bison, sheep, or the several members of the deer family Cervidae. Unlike these mammalian ruminant digesters, equids are post-gastric, or caecal, digesters, who do not decompose their plant food nearly as much as does a ruminant. Because of this, equids are great fertilizers of the soil wherever they roam and spread their droppings. And for the same reason, equids also disperse many more intact seeds of a greater variety of species when compared with ruminant herbivores. Given the wide-spread and frequent movements of equids, this seed dispersal is very broad for a given horse or band of horses covering hundreds, even thousands of square miles. In other words, there is an augmenting of natural seed banks because of them. Another important factor here concerns the bolstering of the food chain or web that incurs due to the input of nutrient-rich droppings. These feed anyone from tiny beetles and microorganisms to birds, lizards, rodents, and larger predators such as bobcat that feed on the latter. Our challenge as humans is to learn to appreciate a wild-horse/burro-containing ecosystem and to grant it sufficient protection in terms of habitat space and generational time so that it can establish its own internal harmony. This is bound to ensue judging from all the amazing past evolutionary history on our planet – but we must allow it to do so, and we must learn to be a part of this wonderful inter-species harmony by living in a non-damaging way with it and allowing it the freedom to establish itself. The benefits of this just wild-equid-containing ecosystem would help restore the living Earth in so many known and yet-to-be-discovered ways. 3. Wild Horse and Burro Tours: It has been shown time and time again that many people both in America and abroad greatly enjoy observing wild horses and burros who are free-living in their natural habitat. Successful eco-tours have been mounted in several states throughout the West and proven themselves profitable. If these equid ecotours were to be promoted in a way that is sensitive to the wild horses and burros and the ecosystem they inhabit, a great mutual benefit could result between humans and equids, uplifting to both. But these tours should be 1 rest-rotated among a great variety of herds so as not to overwhelm any one herd or its habitat. This could be accomplished by proper organization among the tour operators in conjunction with conscientious oversight from government agencies. It is doable and would be a great boon to all involved. 4. Equids as Moderators of Global Warming: Along with zebras, horses and burros are perfect species to reduce dry flammable vegetation that is becoming increasingly prevalent throughout the West due to Global Warming. Again, their post-gastric digestive system makes them perfect for this role. This is due to their not having to as thoroughly decompose the vegetation they consume as a ruminant would. Hence this drier food would not require so much metabolic energy as a ruminant cow for example – who would literally burn itself up trying to subsist on this dry diet. Also, the semi-nomadic nature of the wide-ranging horses and burros lends itself well to this role. They practice a natural rest rotation where fences do not impede this -- fences that are all too prevalent and really illegal within their legal herd areas on BLM and USFS lands. Again the question is whether we people will allow the equids to realize their very healing life style by removing fences and other impediments to their natural freedom. We should learn to appreciate the benign process that is taking place with the restoration of wild horses and burros within their very cradle of evolution: North America. – Remember too that through their droppings, the wild equids will recycle the dry vegetation they consume converting it into vital humus for the rebuilding of soils. This will redound in more moisture-retaining soils, higher water tables, restored watersheds, and more nutrient-rich soils capable of fostering a greater abundance and diversity of plant and animal species, naturally balanced and healing to the living Earth as a whole. Then our human challenge will be a most welcome one: to discover a way to live harmoniously with this wild-horseor-burro-containing ecosystem, even if this is to be just as a visitor, an appreciator of such. –Surely we can do this with God’s help. This is the right way to proceed. Bibliography with notes A Alison, Robert M. (Ph. D. biologist) 2000 (August 19). Canada’s Last Wild Horses. http://members.shaw.ca/save-wild-horses/Research%20Paper%20%20R.%20Alison.htm Substantiates fitting place of wild horses in North America through use of evolutionary evidence and declaims against their insensitive and rash elimination in Canada. B Bell, R.H.V. 1970. The use of the herb layer by grazing ungulates in the Serengeti. IN Adam Watson (ed.) Animal Populations in Relation to Their Food Source. British Ecological Society Symposium. Blackwell Science Publications, Oxford, UK. P.11-125. This elaborate study shows how another equid, the Zebra, complements a variety of grazers, including the Thompson’s Gazelle and the Wildebeest by eating coarser, drier grasses. The removal of these allows other types of vegetation to grow. The study describes the movement patterns of the zebras in relation to the other herbivores and how this relates to an elaborate natural system that has evolved over thousands of generations in Africa. Undoubtedly, a similar system existed between equids in North America and 2 other sympatric species of herbivores. Many of the observations of this elegant study apply to the wild horses and burros in North America, including the West, for the horse, far from being a misfit, restores and enhances the native North American ecosystem, and given the right setting, much the same can be said of the burro. Benton, Michael. 1991. The Rise of the Mammals. Crescent Books, New York. P. 135. Berger, Joel. 1986. Wild Horses of the Great Basin: Social Competition and Population Size. University of Chicago Press, Chicago. Brungardt, Kurt. 2006 (Nov.). Galloping Scared. Vanity Fair. Pages 224-234. This exposes many irregularities in America’s wild horse and burro program with interviews with some of the key players. C Center for Biological Diversity. 2002. Assessing the Full Cost of the Federal Grazing Program. Tucson, Arizona. Cloud Foundation, The. www.thecloudfoundation.org Observation of Ginger Kathrens on effects of PZP on wild horses. D Dallas, Peter. 1987. Wild Horses of the Nevada Desert. A film by Peter Dallas of Sacramento, CA, produced for Nevada Commission for the Preservation of Wild Horses. Ca. ½ hour. Author guided filmmaker and wrote narrative for much of this film. deHaan, C., Steinfeld, H., Rosales, M., Gerber, P., Wassenaar, T. & Castel, V. 2006. Livestock’s Long Shadow: Environmental Issues and Options. Food and Agriculture Organization of the United Nations. Rome. 390 pages. Donlow, J. et al. 2005. Rewilding North America. Nature 436(7053):913-914. Downer, Craig C. 1977. Wild Horses: Living Symbols of Freedom. Western Printers and Publishers, Sparks, NV. 73 pages. Illustrated account by author based on studies undertaken while earning M.S. at University of Nevada-Reno and including many first hand observations in the Pine Nut Range of western Nevada. Photos by author. Later reprinted in 2007 with some new digital photos of wild horses from Western Nevada. Forward by Wild Horse Annie, i.e. Velma Bronn Johnston. Downer, Craig C. 2005 (Dec.). Wild and Free-Roaming Horses and Burros of North America: Factual and Sensitive Statement – How They Help the Ecosystem. Natural Horse 7(3):10-11. Downer, Craig C. 2008 (July-August)). Conservationist with a Heart: A Brief Sketch of the True Lady Wild Horse Annie, Velma B. Johnston. Natural Horse 10(4):22-24. Also presented at convention of Nevada Women’s History Project in Carson City, Nevada and presented in altered version on their website: www.nevadawomen.org Downer, Craig C. 2009, April. Return to Calico Mountain. Published in April entry of www.humaneobserver.blogspot.com page of Elyse Gardner of Novato, CA. Duncan, Patrick. 1992. Zebras, Asses, and Horses: An Action Plan for the Conservation of Wild Equids. IUCN Species Survival Commission, Equid Specialist Group. IUCN, Gland, Switzerland. E F Fazio, Patricia Mabee (Ph.D.). 1995. The Fight to Save a Memory: Creation of the Pryor Mountain Wild Horse Range (1968) and Evolving Federal Wild Horse Protection through 3 1971.” Ph.D. Dissertation. Texas A & M University. Excellent and thorough-going treatment of the wild horse preservation/restoration movement, factually based. Ferguson, D. & N. Ferguson. 1983. Sacred Cows at the Public Trough. Maverick Publications, Bend, Oregon. 250 pages. Excellent documentation and discussion of livestock monopolization of public lands. Forsten, Ann. 1992. Mitochondrial-DNA timetable and the evolution of Equus: comparison of molecular and paleontological evidence. Ann. Zool. Fennici 28:301-309. Fuller, Alexandra. 2009 (Feb.) Spirit of the Shrinking West: Mustangs. National Geographic. Pp. 100-117. Reveals many discrepancies between the law and the actual treatment of returned native wild horses in North America. G General Accounting Office. 1990 (August). Rangeland Management: Improvements Needed in Federal Wild Horse Program. Doc. #RCED-90-110. See page 55. General Accounting Office. 2005 (Sept.) Livestock Grazing: Federal Expenditures and Receipts Vary Depending on the Agency and the Purpose of the Fee Charged. Document # GAO-05-869. Washington, D.C. Grzimek, Bernhard. 2004. Grizimek’s Animal Life Encyclopedia – 2nd Ed. See sections on horse feeding ecology on pages 141, 220, 228 & surrounding pages. To quote from page 141: “The ruminant is faced with being more selective in its feeding. In contrast, the hindgut fermentors, such as some perissodactyls (e.g. modern horses) are able to increase the rate that food passes through their gut, so they extract only the most readily digestible fraction of the food and excrete the indigestible material. As a result, although they must feed almost continuously, they can be much less selective in what they eat. This allows horses to survive on poorer-quality food than artiodactyls are able to do” [i.e. dry flammable vegetation such as cheat grass]. To quote from page 228, Feeding Ecology and Diet: “Equids are primarily grazers and have dental adaptation for feeding on grasses. Their high-crowned molars with complex ridges allow them to effectively grind grasses with higher-fiber content. Though individuals will select the most nutritious and low-fiber forage, they can process senescent [dying] and higher fiber grasses. Equids also have a single stomach and hindgut fermentation. This allows them to digest and assimilate larger amounts of forage during a 24-hour period. By contrast, ruminants with a four-chambered stomach are limited in the volume of forage that can be digested in a 24-hour period. Equids are more effective in assimilating forage and can tolerate and survive on a greater breadth of diet in terms of relative forage qualities/nutrition.” Groves, Colin P. 1974. Horses, asses and zebras in the wild. Newton Abbot Publishers, London. 192 pages. H Haile, J. et al. 2009, Dec. 29. Ancient DNA reveals late survival of mammoth and horse in interior Alaska. Proceedings of the National Academy of Sciences. Vol. 106. No. 52. Pp. 22352-22357. www.pnas.org/cgi/doi/10.1073/pnas.0912510106 Handwerg, Katherine. 1980 (July). Grazing Fees and Fair Market Value. Cascade Holistic Economic Consultants. Eugene, Oregon. 20 pages. Very penetrating analysis. Henderson, Claire. 1991 (February 1). Statement of Claire Henderson in Support of Senate Bill 2278 (North Dakota). Anthropologist. Brings out little recognized facts. 4 High Plains Films. 2001. El Caballo: Wild Horses of North America. The Fund for Animals, New York. 30 minute film. Excellent expose of wild horses’ unfair treatment. Horsetalk. 2009 (April 5). http://www.horsetalk.co.nz/news/2009/08/054.shtml News item regarding BLM’s obstinant stance on zeroing out of West Douglas wild horse herd. Hudak, Mike. 2008. Western Turf Wars: The Politics of Public Lands Ranching. A very timely expose of the livestock industry’s monopoly of the U.S. public lands. Hubert, Marie-Luce. 2007. Mustangs: Wild Horses of the West. Excellent photos and description of day-to-day life of wild horses based on first hand observations over all seasons of the year. I J Janis, C.M. 1976. The evolutionary strategy of the Equidae and the origins of rumen and cecal digestion. Evolution 30:757-774. This explains in eloquent terms how equids are capable of thriving in what would be to other herbivores very marginal ecosystems, such as the desert valleys and ranges of Nevada, especially compared with ruminants evolved in moister, more lush and verdant biomes, such as Hereford or Black Angus cattle. K Kent News. ? date. www.kentnews.co.uk/kent-news/Rare-wildlife-helped-by-wild-horses --newsinkent27563.axpx Explains many beneficial effects of wild horses for ecosystem. Kirkpatrick, J.F. & Fazio, P.M. 2005 (Jan. 25). Wild Horses as Native North American Wildlife: Statement for the 109th Congress (1st Session) in Support of H.R. 297: A Bill in the House of Representatives. www.saponline.org/wild-horses-native.htm Kirkpatrick, J.F. &. Fazio, P.M. 2008 (May). Ecce Equus. Natural History. American Museum of Natural History. New York. Page 30. KLAS TV. 2009, Nov. Stampede to Oblivion. Excellent expose on mistreatment of America’s last wild horses by award winning producer George Knapp. Aired on CBS Channel 8 TV, Las Vegas. Recognized by Edward R. Murrow top award for region. Kleinert, James. 2008. Saving the American Wild Horse. A film by Moving Cloud Productions. www.THEAMERICANWILDHORSE.com Kunzig, Robert. 2008 (February). Drying of the West. National Geographic. Pp. 90-113. L M MacDonald, David. 2001. The New Encyclopedia of Mammals. Oxford University Press. See pages 456-458 & 471-472. To quote: “Food is fermented after passing through the stomach, so the passage rate of vegetation is not as limited as it is in the ruminant grazers, enabling equids to consume large quantities of low-quality forage and to sustain themselves on more marginal habitats and on diets of lower quality than ruminants. Even when vegetation grows rapidly, equids forage for about 60% of the day and up to 80% when conditions worsen.” MacDonald, Cindy R. 2007 (February). Wild Burros of the American West: A Critical Analysis of the National Status of Wild Burros on Public Lands 2006. See www.americanherds.com (Also under American Herds.) MacFadden, Bruce J. 1992. Fossil horses: systematics, paleobiology, and evolution of the family Equidae. Cambridge University Press, Cambridge, U.K. 369 pages. Very valuable compendium & analysis of information. Among other sections, be sure to check 5 out Chapter 12: Population Dynamics, Behavioral Ecology, and “Paleoethology” and also page 112. Martin, P.S. 2005. Twilight of the mammoths: ice age extinctions and the rewilding of America. University of California Press, Berkeley. Martin, P.S. & Wright, H.E. 1967. Pleistocene Extinctions. Yale University Press, New Haven, CT. Meeker, Jo. O. 1979 (May). Interactions Between Pronghorn Antelope and Feral Horses in Northwestern Nevada. Masters of Science Wildlife Manag.Thesis. Un. Nevada-Reno. Morrison, J.C., Sechrest, W., Dinerstein, E., Wilcove, D.S., & Lamoreux, J.F. 2007. Persistence of Large Mammal Fauna as Indicators of Global Human Impacts. Journal of Mammalogy 88(6):1363-1380 Mullen, F.X. Jr. 2010, March 21. Wild horses: Managed wisely or to extinction?. Reno Gazette Journal. Page 1 & following pages. Illustrated. Brings up unfairness issue of relative resource allocations, zeroed out herd areas, favoritism to ranchers, low population numbers, etc. N National Research Council (U.S.). 1980 (Dec.). Wild and Free-Roaming Horses and Burros: Current Knowledge and Recommended Research. Phase I: Final Report of the Committee on Wild and Free-Roaming Horses and Burros. National Technical Information Service, Washington, D.C.. 382 pages. Nock, Bruce (PhD). 2010 (April 19). Wild Horses – the Stress of Captivity. In www.wildhorsepreservation.com/pdf/death-report.pdf Erudite account of suffering, harm and death incurred due to helicopter chase roundups of wild horses by a veterinarian and professor of vet. Medicine. Excellent account easy to read. O Oxley, Ralph & Downer, Craig. 1994. Deserts IN Hare, Tony, Editor. Nature Worlds. MacMillan Reference. Duncan Baird Publishers, London. See especially page 116. P Pelligrini, Steven W. 1971. Home range, territoriality and movement patterns of wild horses in the Wassuk Range of Western Nevada. Master’s thesis. Univ. of Nevada, Reno. Explains how wild horses naturally establish individual band home ranges and even defended territories in some seasons. The latter space their groups within the mountains and adjacent valleys they inhabit. Public Employees for Environmental Responsibility (PEER). 1997. Horses to Slaughter – Anatomy of a Coverup within the BLM (1997-04-01). See www.peer.org Q R Ricklefs, R.E. 1979. Ecology, 2nd Edition. Chiron Pres, New York. See pages 382-384. Rifkin, Jeremy. 1992. Beyond Beef: The Rise and Fall of the Cattle Culture. Dutton, NY. 353 pages. Excellent and thorough-going expose of humanity’s blind promotion of livestock as a way of life and the devastating consequences this is having for life on Earth. Rogers, P., & LaFleur, J. 1999 (Nov. 7). Cash Cows: Taxes support a Wild West holdover that enriches ranchers and degrades the land. San Jose Mercury News, San Jose, California. Another excellent expose with many facts and figures. 6 Ryden, Hope. 1999. America’s Last Wild Horses. 30th Anniversary Edition. The Lyons Press, New York. 345 pages. Excellent discussion of wild horses. This is the book that helped pass the Wild Horse Act in 1971. A must read. S Salter, R.E., Hudson, R.J. 1979. Feeding ecology of feral horses in Western Alberta. Journal of Range Management 32:221-225. Simpson, George Gaylord. 1951. The story of the horse family in the modern world and through sixty million years of history. Oxford University Press, Oxford, U.K. Simpson, George Gaylord. 1965. The Geography of Evolution. See Figure 24. Stolzenburg, William. 2006 (Jan.-Mar.). Where the Wild Things Were. Conservation in Practice 7(1):28-34. Stillman, Deanne. 2008. Mustang: the Saga of the Wild Horse in the American West. Houghton Mifflin Co., Boston. See part III: Last Stand. 345 pages. Excellent. Sussman, Karen. 2008. Various articles in journal of the International Society for the Protection of Mustangs and Burros on pages 4-8. Vol. 48(1):4-8. ISPBM, Lantry, SD. T Trippett, Frank. 1974. The First Horsemen. IN The Emergence of Man series. Time-Life Books, New York. U University of Wyoming. 1979. Proceedings of the Symposium on the Ecology and Behavior of Wild and Feral Equids. Laramie, WY, Sept 6-8, 1974. 236 pages. V Vila, Carles et al. 2001. Widespread origins of domestic horse lineages. Science 291: 474-477. Confirms Forsten’s findings re: origin of modern horse in North America. W Webpages, various. References for Chapter IV regarding bills, etc., concerning wild horses: www.thecloudfoundation.org, www.wildhorsespirit.com, www.wildhorsepreservation.com, www.wildmustangcoalition.org, www.wildhorsesummit.org, www.fund4horses.org, www.naturalhorsesense.com, www.americanherds.com, http://homepage.mac.com/john.brian , www.naturalhorseecology.net, www.naturalhorsesense.com , www.ecology.info/horses.htm Wuerthner, George and Matteson, Mollie (Editors). 2002. Welfare Ranching: The Subsidized Destruction of the American West. Island Press, Washington, D.C. 346 pp. Excellent documentation and illustrations proving livestock’s very damaging effect. Wynne-Tyson, Jon. 1989. The Extended Circle: A Commonplace Book of Animal Rights. Paragon House, New York. 436 pages. X Y Z Zimov, S.A. 2005. Pleistocene park: return of the mammoths’ ecosystem. Science 308:796-98. Indicates great importance of horses to establishing and maintaining tundra grassland that has a cooling effect on the Earth’s atmosphere. Their reintroduction in parts of Siberia is proposed as a way of preventing further Global Warming, especially given accelerating melting of the permafrost. This would apply to northern North America. 7 8