Santa Catalina Island Biogeography “Biogeography is the study of the facts and patterns of species distribution. It’s the science concerned with where animals are, where plants are, and where they are not” (Song of the Dodo, David Quammen). “Islands have been especially instructive because their limited area and their inherent isolation combine to make patterns of evolution stand out starkly. It’s such an important truth I’ll repeat it: Islands give clarity to evolution. On an island you might find those enormous tortoises, those flightless birds weighing half a ton, those pygmy chameleons and hippos. Generally you will also find fewer species, and therefore fewer relationships among species, as well as more cases of species extinction. All these factors result in a simplified ecosystem, almost a caricature of nature’s full complexity. Consequently islands have served as the Dick-and-Jane primers of evolutionary biology, helping scientists master enough vocabulary and grammar to begin to comprehend the more complex prose of mainlands” (Quammen, pg. 19). Lying roughly 20 miles off the coast of Southern California, Catalina Island has had many organisms float and fly right to it. Where as other organisms may have brushed the shores and proceeded to a different destination. How Catalina acquired its different species of organisms is quite amazing and dependent on many different processes and chains of events. First of all the type of island determines how many different species will inhabit it. Islands can be broken into the following categories: Continental islands once part of the mainland, varied in their geology, usually inhabited by land mammals. * Immigration of species is faster than evolution Oceanic islands “built” over an oceanic plate, remote and have never been connected to the mainland, inhabited by few land mammals. * Evolution is faster than immigration Catalina is an oceanic island. However, it is relatively close to the mainland for an oceanic island and is fairly large in size. This has had a profound influence on the organisms that inhabit it. The closer an island to the mainland the higher the probability of dispersed organisms to travel the short distance. The larger the island the chances of organisms landing on the island increase tremendously. This chance rather than logic theory is referred to as Sweepstakes Dispersal. There are hundreds of thousands of organisms dispersed from the mainland each year, and very few actually land in a hospitable environment. The odds of an organism landing in a hospitable environment are equivalent to winning the lottery. Millions of people play and there is only one winner, possibly two. Like dispersal to islands it is chance not logic in determining the winner. The three main modes of dispersal of organisms to islands are: Wind seeds, insects, spiders, birds and bats may disperse this way. Because Catalina Island is relatively close to the mainland many seeds and small creatures may have dispersed to the island in this manner. Wing seeds may get caught in the feathers of a bird or travel to the island within the bird’s digestive system and be deposited on the island in a nice fertilized package. Water larger animals such as squirrels, fox, snakes and frogs may have rafted over during periods of flooding on the mainland. Creatures may take refuge on floating logs and other debris, which is then swept out to sea (also known as rafting). Seeds may also use this means of dispersal. Bladder pod, a plant found on Catalina probably traveled this way across the channel, it can float in seawater for up to 4 weeks. Once organisms colonize an island they begin to adapt to their new surroundings immediately. There are common adaptations and patterns that occur on most islands. Rarely are large grazing animals native to oceanic islands. Therefore, plants tend to lose their thorns and other means of protection from grazing animals. Due to the difference in climate, soil, less competition and grazing, some plants that are bushes on the mainland become treelike on islands through a process known as arborescence. Examples of arborescence occurring on Catalina are Toyon and the Island Scrub Oak. Niche shifts are also common on islands. These occur because of fewer numbers of species and absence of competition therefore, creating empty niches. An animal can then fill a new niche that is different from the one it occupied on the mainland. Animals may also fill two niches like the Beechy Ground Squirrel, which occupies both the arboreal and terrestrial niches on Catalina. Some animals on islands evolve over time to become smaller or larger creating either a dwarf or gigantic species gigantism and dwarfism. On Catalina we have both dwarf and gigantic species. The Catalina Island Fox is a dwarf species while the quail and squirrel are gigantic species. There is no one answer as to why this occurs. One theory on gigantism is that on islands generally there is reduced predation and competition within the species that would normally draw on available food and shelter. This may allow an animal to get much larger than it could on the mainland. A larger animal may also store water and fat better, survive hungry and cold times, and finally give birth to larger babies who will also have a better chance at surviving. Dwarfism may occur because larger animals with no population control, often the case on islands, result in over population, leading to malnutrition of the group, which may cause stunting of growth in individuals, and over time may lead to dwarfism. Smaller animals may also be more successful on an island that has fewer food sources. Therefore producing smaller offspring that would have a better chance of survival. Some plant and animal species have evolved and adapted enough to become a new species or subspecies; these new species are endemic to Catalina Island. Species can be endemic to islands, regions or continents. It simply means that species is unique to a specific area. When looking in a guidebook make sure to look closely whether a plant is endemic to Catalina or all of the Channel islands. Catalina Island’s 8 endemic plants include: Ironwood, St. Catherines Lace, Catalina Island Live Forever, Island Mahogany, Yerba Santa, Manzanita, Wallace’s Night Shade, Catalina Island Bed Straw. **Remember scientific theories are not wrought in stone, please try to keep current on changing theories and ideas and teach them to your students as such.