CURRICULUM VITAE
Katherine E. Willmore
ADDRESS
Anthropology Department
409 Carpenter Building
Pennsylvania State University
University Park
State College, PA, 16802
E mail: kew20@psu.edu
Phone: 814-865-2006
CURRENT POSITION
Postdoctoral Scholar in Dr. Joan Richtsmeier’s laboratory. Currently working on a
collaborative project aimed at identifying genes that are involved in craniofacial
development and their role in the evolution of modern human craniofacial morphology.
ACADEMIC BACKGROUND
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BSc. (honours) University of Guelph, Guelph, Ontario. Human Kinetics (1999)
PhD University of Calgary, Calgary, Alberta. Title: Factors affecting variability
in the mammalian skull (2006)
AWARDS AND DISTINCTIONS
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2005 William H. Davies Scholarship, University of Calgary
2005 Province of Alberta Graduate Scholarship, University of Calgary
2004-2005 Alexa W. Church Graduate Scholarship in Medical Sciences,
University of Calgary
2004 William H. Davies Scholarship, University of Calgary
2003-2004 Medical Science Graduate Education Scholarship, University of
Calgary
2003 William H. Davies Scholarship, University of Calgary
2002 Alberta Heritage Foundation, Graduate Student Scholarship, University of
Calgary
2002-2003 Faculty of Graduate Studies Scholarship, University of Calgary
2003 CMSA Letter of Excellence, Teaching Award for Faculty of Medicine
Anatomy
PROFESSIONAL ACTIVITIES
Professional Service
 Participated in a faculty hiring committee for the Bachelor of Health Sciences
Program at the University of Calgary
 Peer Review for Evolutionary Biology, Journal of Anatomy, American Journal of
Physical Anthropology
 Co-organizer of American Association of Physical Anthropology symposium on
Morphological Integration (Columbus, OH)
 Presented at the 2007 Postdoctoral Society Meeting at the Pennsylvania State
University
Academic memberships
 American Association of Physical Anthropology
 Society for Comparative and Integrative Biology
 Society for the Study of Evolution
TEACHING
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MDSC 402 (Full Year): Organismal Biology, Bachelor of Health Sciences
Program, University of Calgary (Teaching Assistant: Fall and Winter, 20052006) - Undergraduate Students
Renal Course- Anatomy, University of Calgary, Faculty of Medicine
(Teaching Assistant: Spring, 2001, 2002, 2003, 2004, 2005) - Medical
Students
Musculoskeletal Course- Anatomy, University of Calgary Faculty of
Medicine (Teaching Assistant: Fall, 2000, 2001, 2002, 2003, 2004) - Medical
Students
Gastrointestinal System Course- Anatomy, University of Calgary Faculty of
Medicine (Teaching Assistant: Fall, 2000, 2001, 2002, 2003, 2004) - Medical
Students
Cardiovascular System Course- Anatomy, University of Calgary Faculty of
Medicine (Teaching Assistant: Fall-Winter, 2000, 2001, 2002, 2003, 2004,
2005) - Medical Students
Reproductive System Course- Anatomy, University of Calgary Faculty of
Medicine Teaching Assistant: (Fall, 2000, 2001, 2002, 2003, 2004) - Medical
Students
Structure and Development Integrated Gross Anatomy, Embryology and
Histology. University of Puerto Rico Medical School (Co-taught Fall, 2002). Medical Students
Human Gross Anatomy. Mount Royal College, Calgary, AB (Full instructor:
Fall and Winter, 2001-2002) - Undergraduate Students
PEER REVIEWED MANUSCRIPTS
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Willmore K.E., Young N.M., and Richtsmeier J.T. (2007) Phenotypic
Variability: Its Components, Measurement, and Underlying Developmental
Mechanisms. Evolutionary Biology. 34(3-4): 99-120.
Willmore K.E., Leamy L., and Hallgrímsson B. (2006) The Effects of
Developmental and Functional Interactions on Mouse Cranial Variability Through
Late Ontogeny. Evolution and Development. 8:550-567.
Willmore, K.E., Zelditch, M.L., Lozanoff, S., Young, N., and Hallgrímsson, B.
(2006) Canalization and Developmental Stability in the Brachyrrhine Mouse.
Journal of Anatomy. 208:361-372.
Willmore, K.E., Klingenberg, C.P., and Hallgrímsson, B. (2005) Congruence of
Fluctuating Asymmetry and Environmental Variance in Rhesus Macaque Skulls.
Evolution. 59:898-909.
Hallgrímsson, B., Ó Donnabhain, D.E., Blom, M.C., Lozada, J.F., and Willmore,
K. (2005) Why Are Rare Traits Unilaterally Expressed? Trait Frequency and
Unilateral Expression for Cranial Non-Metric Traits in Humans. American
Journal of Physical Anthropology. 128:14-25.
Hallgrímsson, B., Willmore, K., Dorval, C., and Cooper, D.M.L. (2004)
Craniofacial Variability and Modularity in Macaques and Mice. Journal of
Experimental Zoology (MDE) 302B:207-225.
Hallgrímsson, B., Miyake, T., Willmore, K., and Hall, B.K.H. (2003) The
Embryological Origins of Developmental Stability: Size, Shape and Fluctuating
Asymmetry in Prenatal Random Bred Mice. Journal of Experimental Zoology
(MDE) 296B:40-57.
Hallgrímsson, B., Willmore, K., and Hall, B.K. (2002) Canalization,
Developmental Stability, and Morphological Integration in Primate Limbs.
Yearbook of Physical Anthropology 45:131-158
Sealy, J., Pfeiffer, S., Yates, R., Willmore, K., Manhire, A., Maggs, T., and
Lanham, J. (2000) Hunter-gatherer child burials from the Pakhuis mountains,
Western Cape: growth, diet and burial practices in the late Holocene. South
African Archaeological Bulletin 55:32-43.
MANUSCRIPTS SUBMITTED FOR PUBLICATION AND IN PREPARATION
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Willmore K.E., Roseman C.C., Rogers J., Cheverud J.M., and Richtsmeier J.T.
Genetic Variation in Baboon Craniofacial Sexual Dimorphism. (In Preparation)
Willmore K.E., Chung J., Liu W., and Hallgrímsson B. Developmental Stability
and Canalization in Mice with Deficient Type I Collagen (In Preparation)
Willmore K.E., Lawson H.A., Roseman C.C., Cheverud J.M., and Richtsmeier
J.T. Comparison of Phenotypic and Genetic Integration Between Mouse and
Baboon. (In Preparation).
Willmore K.E., and Richtsmeier J.T. Patterns of Craniofacial Phenotypic
Integration in Male and Female Baboons (In Preparation).
BOOK CHAPTERS
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Willmore, K., and Hallgrímsson, B. (2005) Within Individual Variation:
Developmental Noise versus Developmental Stability. In Variation: A Central
Concept in Biology. Hallgrímsson B. and Hall B.K.eds. New York: Academic
Press.
 Stock, J., and Willmore, K. (2003) Body Size, Bone Mass and Biomechanical
Analyses of the Adult Post-cranial Remains. In Bones of the Ancestors: The
Archaeology and Osteobiography of the Moatfield Ossuary. Williamson R.F.
and Pfeiffer S. eds. Canadian Museum of Civilization, Quebec.
PAPERS PRESENTED AT PROFESSIONAL MEETINGS
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Willmore, K.E., Lawson, H.A., Roseman, C.C., Cheverud, and Richtsmeier, J.T.
(2008). Comparison of phenotypic and genetic integration between mouse and
baboon. Journal of Physical Anthropology. April, Columbus OH.
Frazier, B.C., Willmore, K.E., and Richtsmeier, J.T. (2008). Characterizing
neurocranial shape in microcephalic children. Journal of Physical Anthropology.
April, Columbus OH.
Starbuck J., Willmore, K., Reeves, R., and Richtsmeier, J. (2008). Quantitative
comparison of midface morphology in Down syndrome individuals and
unaffected siblings. Journal of Physical Anthropology. April, Columbus OH.
Richtsmeier, J., Hill, C., Willmore, K., Fisher, E.M., Tybulewicz, V., Sharpe, P.,
and Reeves, R. (2007). Skull shape in Down syndrome mouse models. Expert
Workshop on the Biology of Chromosome 21 Genes: Towards Gene-phenotype
Correlations in Down syndrome. September, Washington DC.
Willmore, K.E., Roseman, C.C., Cheverud, J.M., Frazier, B.C., Rogers, J.,
Weiss, K.M., Walker, A., Hildebolt, C.F., and Richtsmeier, J.T. (2007). Genetic
and phenotypic variation in craniofacial dimorphism in baboons. The American
Journal of Physical Anthropology 132(S44):251. March, Philadelphia PA.
Richtsmeier, J.T., Weiss, K.M., Buchanan, A., Walker, A., Jablonski, N., Frazier,
B.C., Willmore, K.E., Lawson, H., Yan, P., Hill, C.A., Roseman, C.C.,
Cheverud, J.M., Falk, D., Hildebolt, C.F., and Rogers, J. (2007). Developmental
genetic basis of primate craniofacial variation and human origins. The American
Journal of Physical Anthropology 132(S44):198. March, Philadelphia PA.
Roseman, C.C., Willmore, K.E., Richtsmeier, J.T., Rogers, J., Weiss, K.M.,
Walker, A., Hildebolt, C.F., and Cheverud, J.M. (2007). The genetic basis of
phenotypic integration in baboon and mouse craniomandibular morphology with
implications for human cranial evolution. The American Journal of Physical
Anthropology 132(S44):202. March, Philadelphia PA.
Willmore, K., and Hallgrimsson, B. (2006). The effects of developmental and
functional interactions on mouse cranial variability at different ages. American
Association of Physical Anthropologists. The American Journal of Physical
Anthropology 129(S42):188. March, Anchorage AK.
 Willmore, K., Klingenberg, C.P., and Hallgrímsson, B. (2005). Congruence
between canalization and developmental stability in Macaca mulatta crania.
American Association of Physical Anthropologists. The American Journal of
Physical Anthropology 126(S40):224. April, Milwaukee WS.
 Willmore, K., Lozanoff, S., Zelditch, M., and Hallgrímsson, B. (2005) Direct and
Indirect Developmental Effects of a Craniofacial Mutation in a Mouse Model.
Society for Integrative and Comparative Biology. San Diego, January.
 Hallgrímsson, B., Willmore, K., and Dymond, M. (2004). Ontogeny of variance
and developmental noise. Journal of Morphology 260 (3):296. July Boca Raton,
Florida.
 Willmore, K., and Hallgrímsson, B. (2003). Variance Components and
Developmental Stability in Rhesus Macaque Skull. European Society for
Evolutionary Biology. August, Leeds, UK.
 Hallgrímsson, B., Willmore, K., and Dymond, M. (2003). The use of mouse
mutant models for understanding canalization and developmental stability.
European Society for Evolutionary Biology. August, Leeds, U.K.
 Willmore, K., Lozanoff, S., and Hallgrímsson, B. (2002) Canalization and
developmental stability in craniofacial development in the Brachyrrhine mouse.
The American Association of Physical Anthropologists. April, Buffalo, NY.
American Journal of Physical Anthropology 117(S34):166.
 Hallgrímsson, B., and Willmore, K. (2002) Mutant models for Canalization and
Developmental Instability in Mice. International Congress of Systematic and
Evolutionary Biology. August, Patras, Greece