Physical Anthropology / M. Waters FAMILY: Hominidae GENUS: Australopithecus, 4.2 to 1.6 mya Relatively small brains (as compared to Homo, average cranial capacity: 420 cm3) Shape of jaw long, and narrow with more prognathism than Homo Large molar teeth (as compared to Homo), adaptation to diet based heavily on vegetation Relative limb length (i.e. forelimbs to hindlimbs) intermediate between apes and Homo Early species had stiff wrist joints like great apes Sexual dimorphism Examples of species: 1) afarensis (discovered 1974, Hadar, Ethiopia/E. Africa) 2) africanus (discovered 1925, Taung, South Africa) GENUS: Paranthropus (a.k.a. “Robust” Australopithecus), 2.4 to 1 mya Relatively small brains (as compared to Homo, average cranial capacity: 520 cm3) Adaptations to diet consisting of hard, fibrous vegetation (e.g. roots, tubers, etc.): - large jaw bone with massive, square molars and thick tooth enamel - sagittal crest on braincase - large, flaring zygomatic arches, resulting in large cheekbones and facial “dishing” (cheekbones protrude beyond nasal cavity) Examples of species: 1) aethiopicus (a.k.a. the “Black Skull”, discovered 1985, Lake Turkana, Kenya/E. Africa) 2) boisei (discovered 1959, Olduvai Gorge, Tanzania/E. Africa) GENUS: Homo, 2.5 mya to present Larger cranial capacities than previous hominids (ranging from 500 to 2000 cm3) Rounder cranium than earlier hominids with less prognathism (smaller delicate facial bones, though some had heavy brow ridges) Smaller molar teeth and jaws More “advanced” behaviors: stone tool cultures, migration out of Africa (H. erectus), cultural traditions, art (H. sapiens), etc. Examples of species on the next page… GENUS: Homo SPECIES: habilus/rudolfensis (2.4 to 1.6 mya) Discovered 1961, Tanzania/E. Africa (Olduvai Gorge); remains also found in S. Africa Larger cranial capacity (average 630 cm3) than Australopithecus/Paranthropus Smaller molars and less prognathism than Australopithecus/Paranthropus Behavioral features: Oldowan tool culture (stone core and flake technology); stone tool cutmarks found on animal remains hint at scavenging as a strategy for obtaining meat SPECIES: erectus/ergaster (1.8 mya to 200,000ya) Discovered 1891 Java, Indonesia; remains also found in China, Europe and Africa Average cranial capacity: 900 cm3 Large, robust body build Large browridges, sloping forehead Other distinctive cranial features in those found outside Africa (erectus): nuchal torus (neck muscle attachment site) and sagittal ridge (ridge along top of braincase); ergaster specimens (Africa) lack these features and have thinner cranial bones, smaller browridges Behavioral features: Acheulian tool culture (bifacial hand axes) as early as 1.4 mya in E. Africa; first to migrate out of Africa?; contradicting evidence exists regarding hunting behavior (scavenging for meat was more likely) SPECIES: neanderthalensis (125,000 ya to 28,000 ya) Discovered 1856, Germany; remains also found in other parts of Europe, and Middle East Range of cranial capacity: 1300cm3 to 2000 cm3 Distinctive cranial features: large browridge, sloping forehead, occipital bun, lack of chin Distinctive skeletal features: “Stocky”/compact build, thick/robust bones with heavy muscle markings (more so than modern H. sapiens) Behavioral features: Mousterian tool culture (including spearheads for hunting large game); burials, care of the sick/elderly, language (?) SPECIES: floresiensis (95,000 ya to 18,000 ya) Discovered 2004, Flores, Indonesia Extremely small bodied (approx. 3 feet tall) and small brained (380 cm3) Cranial bones and dentition mostly resemble H. erectus Highly unusual because small hominids were thought to have died out long ago; along with tiny hominid remains, dwarf elephants were also discovered on Flores (perhaps small body size was an evolutionary adaptation to limited resources available on the island) SPECIES: sapiens (160,000 ya to present) Earliest fossil evidence is from Ethiopia/East Africa Cranial features: distinctly round cranium, average cranial capacity of 1400 cm3, smaller facial bones and browridges than Neanderthals, more pronounced chin Skeletal features: body less muscular than earlier members of genus Homo Early behavioral features: Upper Paleolithic tool culture (wider variety of materials used: bone, antler, ivory, horns, etc. to make more complex tools and artifacts such as spear throwers, bone needles, barbed hooks, etc.); first to create art (cave paintings, carvings, sculptures, etc.); inventors of plant and animal domestication, civilizations, writing, etc.