Appendix III
A list of the 201 morphological characters used in the cladistic analysis of this paper. Characters
and description are mainly from Norell and Clarke28 with slight changes, but are all unordered.
1.
2.
3.
4.
5.
6.
7.
8.
9.
10.
11.
12.
13.
14.
15.
16.
17.
18.
19.
20.
21.
22.
23.
Premaxillae: (0) unfused in adults; (1) fused anteriorly in adults; (2) fused posteriorly as
well as anteriorly.
Premaxillary teeth: (0) present; (1) absent.
Maxillary teeth: (0) present; (1) absent.
Dentary teeth: (0) present; (1) absent.
Tooth crown, ornamentation: (0) serrations; (1) serrations vestigial or absent.
Dentaries: (0) joined proximally by cartilage; (1) joined in osseous symphysis.
Mandibular symphysis, two strong grooves forming an anteriorly-opening “v” in ventral
view: (0) absent; (1) present.
Facial margin: (0) primarily formed by the maxilla, with the maxillary process of the
premaxilla restricted to the anterior-most extreme; (1) maxillary process of the premaxilla
extending 1/2 facial margin; (2) maxillary process of the premaxilla extending most of
facial margin.
Nasal process of the premaxilla: (0) short; (1) long, posteriorly approaching the frontals.
Nasal process of the maxilla, dorsal ramus: (0) prominent, exposed medially and laterally;
(1) absent or reduced to slight medial, and no lateral, exposure.
Nasal process of the maxilla, participation of ventral ramus in anterior margin of antorbital
fenestra in lateral view: (0) present; (1) absent.
Osseous external nares: (0) considerably smaller than the antorbital fenestrae; (1) larger.
Ectopterygoid: present (0); absent (1).
Articulation between the vomer and pterygoid: (0) present; (1) reduced, narrow process of
pterygoid passes dorsally over palatine to contact vomer; (2) absent, pterygoid and vomer
do not contact.
Palatine and pterygoid: (0) long anteroposteriorly overlapping contact; (1) short primarily
dorsoventral contact.
Palatine contacts: (0) maxillae only; (1) premaxillae and maxillae.
Vomer contacts premaxilla: (0) present; (1) absent.
Coronoid ossification: (0) present; (1) absent.
Basisphenoid articulation with the pterygoid developed as a process: (0) present; (1) absent.
Basipterygoid processes: (0) elongate; (1) short, articulation with pterygoid sub-equal to
longer than amount projected from the basisphenoid rostrum.
Basisphenoid, pterygoid articulations: (0) located basal on basisphenoid; (1) located
markedly anterior on basisphenoid (“parasphenoid”) such that the articulations are
subadjacent on narrow rostrum.
Basisphenoid/pterygoid articulation: (0) anteroventral; (1) mediolateral; (2) entirely
dorsoventral.
Pterygoid, articular surface for basisphenoid: (0) concave “socket,” or short groove
enclosed by dorsal and ventral flanges; (1) flat to convex; (2) flat to convex facet, stalked,
variably projected.
24.
25.
26.
27.
28.
29.
30.
31.
32.
33.
34.
35.
36.
37.
38.
39.
40.
41.
42.
43.
44.
45.
46.
47.
Pterygoid, kinked: (0) present, surface for basiphenoid articulation at a high angle to the
axis of the palatal process of the pterygoid; (1) absent, articulation in line with axis of
pterygoid.
Osseous interorbital septum (mesethemoid): (0) absent; (1) present.
Osseous interorbital septum (mesethemoid): (0) restricted to posterior or just surpassing
premaxillae/frontal contact in rostral extent, does not surpass posterior edge of external
nares in rostral extent; (1) extending rostral to posterior extent of the frontal processes of
the premaxillae and rostral to posterior edge of external nares.
Eustachian tubes: (0) paired and lateral; (1) paired, close to, cranial midline; (2) paired and
adjacent on midline or single anterior opening.
Eustachian tubes ossified: (0) absent; (1) present.
Squamosal, ventral or "zygomatic” process: (0) variably elongate dorsally enclosing the
dorsal process and extending anteroventrally along shaft of quadrate, dorsal head of
quadrate not visible in lateral view; (1) short, head of quadrate exposed in lateral view.
Orbital process of quadrate, pterygoid articulation: (0) pterygoid broadly overlapping
medial surface of orbital process; (1) restricted to anteromedial edge of process.
Quadrate, orbital process: (0) pterygoid articulates with anterior-most tip; (1) pterygoid
articulation does not reach tip; (2) pterygoid with no extent up orbital process.
Quadrate/ pterygoid contact: (0) as a facet, variably with slight anteromedial projection
cradling base; (1) condylar, with a well-projected tubercle on the quadrate.
Quadrate, well-developed tubercle on anterior surface of dorsal process: (0) absent; (1)
present.
Quadrate, quadratojugal articulation: (0) overlapping; (1) peg and socket articulation.
Quadrate, braincase articulation: (0) with squamosal only; (1) with squamosal and prootic.
Quadrate, dorsal process, development of intercotylar incisure between prootic and
squamosal cotylae: (0) absent, articular surfaces not differentiated; (1) two distinct
articular facets, incisure not developed; (2) incisure present, “double headed.”
Quadrate, ventral process: (0) bicondylar articulation with mandible; (1) tricondylar
articulation, additional posterior condyle or broad surface.
Quadrate, pneumaticity: (0) absent; (1) present.
Quadrate, cluster of pneumatic foramina on posterior surface of the tip of dorsal process: (0)
absent; (1) present.
Quadrate, large single pneumatic foramen: (0) absent; (1) posteromedial surface of dorsal
process.
Articular pneumaticity: (0) absent; (1) present.
Dentary, strongly forked posteriorly: (0) unforked; (1) forked.
Splenial, anterior extent: (0) splenial stops well posterior to mandibular symphysis; (1)
extending to mandibular symphysis, though non-contacting; (2) extending to proximal tip
of mandible, contacting on midline.
Mandibular symphysis, dorsal surface: (0) concave; (1) flat to convex.
Mandibular symphysis, symphysial foramina: (0) absent; (1) present.
Mandibular symphysis, symphysial foramina: (0) single; (1) paired.
Mandibular symphysis, symphysial foramina: (0) opening on posterior edge of symphysis;
(1) opening on dorsal surface of symphysis.
48.
49.
50.
51.
52.
53.
54.
55.
56.
57.
58.
59.
60.
61.
62.
63.
64.
65.
66.
67.
68.
69.
70.
71.
72.
73.
74.
75.
76.
77.
78.
Meckel’s groove: (0) open, conspicuous; (1) not exposed medially.
External mandibular fenestra: (0) absent; (1) present.
Jugal/postorbital contact: (0) present; (1) absent.
Frontoparietal suture (0) open; (1) closed.
Cervical vertebrae: (0) dorsoventrally compressed amphicoelous; (1) anterior surface
heterocoelous, posterior surface flat; (2) both anterior and posterior surfaces heterocoelous.
Thoracic vertebrae (with “true” ribs, articulating with the sternum), one or more with
prominent hypapophyses: (0) absent; (1) present.
Dorsal vertebrae (with free, ventrally projecting free ribs), count: (0) 12 or more; (1); 11; (2)
10; (3) 9 or less.
Thoracic vertebrae, at least part of series heterocoelic: (0) absent; (1) present.
Dorsal vertebrae, parapophyses: (0) rostral to transverse processes; (1) directly ventral to
transverse processes, close to midpoint of vertebrae.
Dorsal vertebrae, centra markedly longer than wide: (0) absent; (1) present.
Dorsal vertebrae, lateral excavations: (0) absent or slight indentations; (1) present, deep,
emarginate excavations.
Thoracic vertebrae with ossified connective tissue connecting transverse processes: (0)
absent; (1) present.
Notarium: (0) absent, (1) present.
Sacral vertebrae, number ankylosed: (0) less than 7; (1) 7; (2) 8; (3) 9; (4) 10; (5) 11 or
more; (6) 15 or more.
First sacral vertebra, anterior vertebral articulation: (0) approximately equal in height and
width; (1) wider than high.
Free caudal vertebrae, number: (0) more than 8; (1) 8 or less.
Caudal vertebrae, (chevron), fused on at least one anterior caudal: (0) present; (1) absent.
Anterior (free) caudals, length of transverse processes on caudals: (0) approximately the
vertebral width; (1) significantly shorter than vertebral width.
Anterior (free) caudal vertebrae: (0) well developed pre/post-zygopophyses; (1)
pre/post-zygopophyses short.
Distal caudals: (0) unfused; (1) fused into a pygostyle.
Pygostyle: (0) long, more than the length of 4 caudal vertebrae; (1) short less than the
length of 4 caudal vertebrae; (2) less then 2 caudal vertebrae in length.
Ossified uncinate processes: (0) absent; (1) present; (2) fused to ribs.
Gastralia: (0) present; (1) absent.
Ossified sternal plates: (0) unfused; (1) fused; (2) fused, with slightly raised midline ridge;
(3) fused with projected carina.
Carina: (0) restricted to posterior half of sternum; (1) approaches anterior limit of sternum.
Sternum, dorsal surface, central pneumatic foramen: (0) absent; (1) present.
Sternum, midline ridge from anterior edge: (0) absent; (1) present.
Sternum, deep coracoidal sulci: (0) present; (1) absent.
Sternum, coracoidal sulci spacing on anterior edge: (0) widely separated mediolaterally; (1)
adjacent; (2) crossed on midline.
Sternal costal processes: (0) three; (1) four; (2) five; (3) six; (4) seven; (5) eight.
Sternum: raised, paired intermuscular ridges parallel to sternal midline: (0) absent; (1)
79.
80.
81.
82.
83.
84.
85.
86.
87.
88.
89.
90.
91.
92.
93.
94.
95.
96.
97.
98.
99.
100.
101.
102.
103.
104.
105.
106.
107.
present.
Clavicles: (0) fused into a furcula; (1) unfused.
Interclavicular angle (clavicles elongate): (0) greater than 90; (1) less than 90 degrees.
Furcula, hypocleideum: (0) absent; (1) a tubercle; (2) an elongate process.
Furcula, laterally excavated: (0) absent; (1) present.
Scapula and coracoid: (0) sutured; (1) free.
Scapula and coracoid articulation: (0) pit shaped scapular cotyla developed on the coracoid,
and coracoidal tubercle developed on the scapula (“ball and socket” articulation); (1)
scapular cotyla of the coracoid convex; (2) flat facet.
Coracoid, procoracoid process, (0) absent; (1) present.
Coracoid: (0) height approximately equal mediolateral dimension; (1) height more than
twice the mediolateral, coracoid “strut like.”
Coracoid, lateral margin in ventral view: (0) straight to slightly concave; (1) convex.
Coracoid, dorsal surface: (0) strongly concave; (1) flat to convex throughout majority of
shaft.
Coracoid, dorsal surface, deep fossa: (0) absent; (1) present.
Coracoid: (0) solid; (1) hollow, pneumatized.
Coracoid: (0) pneumatic foramen proximal; (1) distal.
Coracoid, lateral process: (0) absent; (1) present.
Coracoid, ventral surface, distal fossa and lateral intermuscular line: (0) absent; (1) faint
impressions; (2) well developed depression and ridge.
Coracoid, glenoid facet: (0) dorsal to acrocoracoid process/ “biceps tubercle”; (1) ventral to
acrocoracoid process.
Coracoid, acrocoracoid: (0) straight; (1) hooked medially.
Coracoid, n. supracoracoideus passes through coracoid, foramen marking its passage: (0)
present; (1) absent.
Coracoid, passage of foramen n. supracoracoideus opening into a medial groove: (0) absent;
(1) present.
Coracoid, medial surface, area of exit of the foramen n. supracoracoideus (when developed):
(0) strongly depressed; (1) flat to convex.
Angle between coracoid and scapula at glenoid: (0) more than 90 degrees; (1) 90 degrees or
less.
Scapula, (0) posterior end as wide or wider than proximal dorsoventral width; (1) tapering
distally.
Scapular shaft: (0) straight; (1) dorsoventrally curved.
Scapular shaft: (0) mid-part of scapular blade expanded to approach width of anterior end;
(1) mid-shaft approximately constant width, narrow.
Scapula, acromium process: (0) blunt (1) tapering to a point.
Scapula, acromium process: (0) straight; (1) hooking laterally.
Humerus and ulna, length: (0) humerus longer; (1) ulna and humerus approximately the
same length (2) ulna significantly longer than humerus.
Humerus, proximal view, head in anterior or posterior aspect: (0) strap-like, articular
surface flat, no proximal midline convexity; (1) head domed proximally.
Humerus, (0) projected farther proximally dorsally; (1) midline projected farthest
108.
109.
110.
111.
112.
113.
114.
115.
117.
118.
119.
120.
121.
122.
123.
124.
125.
126.
127.
128.
129.
130.
131.
proximally.
Humerus, capital incisure: (0) absent; (1) present.
Humerus, capital incisure: (0) an open groove; (1) capital incisure closed by a muscular
tubercle associated with an insertion just distal to humeral head.
Humerus, anterior surface, well-developed fossa on midline making the proximal articular
surface appear v-shaped in proximal view: (0) absent, (1) present.
Humerus, ventral tubercle: (0) absent; (1) present.
Humerus, “transverse groove”: (0) absent; (1) present, developed as discreet depressed scar
on proximal surface of bicipital crest or slight transverse groove.
Humerus, deltopectoral crest: (0) projected dorsally (in line with long axis of humeral head;
(1) projected anteriorly (at an angle to long axis of the humeral head) .
Humerus, deltopectoral crest: (0) less than shaft width; (1) approximately the same measure
(2) dorsoventral width greater than shaft width.
Humerus, deltopectoral crest, proximal, posterior surface: (0) flat to convex, (1) concave.
116. Humerus, bicipital crest, scar/fossa for muscular attachment on cranial or distal
surface: (0) absent; (1) present.
Humerus, bicipital crest, pit shaped fossa for muscular attachment: (0) anterodistal on
bicipital crest; (1) directly ventrodistal at tip of bicipital crest; (2) posterodistal, variably
developed as a fossa.
Humerus, bicipital crest: (0) little or no cranial projection; (1) developed as a cranial
projection relative to shaft surface in ventral view; (2) hypertrophied, rounded tumescence.
Humerus, proximal end, one or more pneumatic foramina: (0) absent; (1) present.
Humerus, distal condyles: (0) developed distally; (1) developed on the anterior surface of
the humerus.
Humerus, long axis of dorsal condyle: angling proximoventrally at low angle to the
humeral axis; (0) overall more proximodistally oriented; (1) at high angle to humeral axis,
almost transversely oriented.
Humerus, distal condyles: (0) sub-round, bulbous; (1) weakly defined, “strap-like.”
Humerus, distal margin: (0) approximately perpendicular to long axis of humeral shaft; (1)
ventrodistal margin projected significantly distal to dorsodistal margin, distal margin
angling strongly ventrally, sometimes described as a well-projected flexor process.
Humerus, distal end, compressed anteroposteriorly and flared dorsoventrally at its distal
extreme: (0) absent; (1) present
Humerus, brachial fossa: (0) absent; (1) present, developed as a flat scar or as a
scar-impressed fossa.
Humerus, ventral condyle: (0) length of long axis of condyle less than the same measure of
the dorsal condyle; (1) same or greater.
Humerus, distal end, posterior surface, groove for passage of m. scapulotriceps: (0) absent;
(1) present.
Humerus, m. humerotricipitalis groove: (0) absent (1) present as a well developed ventral
depression contiguous with the olecranon fossa and demarcating a posteroventral ridge.
Ulna, cotylae: (0) dorsoventrally adjacent; (1) widely separated by a deep groove.
Ulna, dorsal cotyla convex: (0) absent; (1) present.
Ulna, distal end, dorsal condyle, dorsal trochlear surface developed as a semilunate ridge:
132.
133.
134.
135.
136.
137.
138.
139.
140.
141.
142.
143.
144.
145.
146.
147.
148.
149.
150.
151.
152.
153.
154.
155.
(0) absent; (1) present.
Ulna, distal end, dorsal condyle, dorsal trochlear surface, extent along posterior margin: (0)
less than transverse measure of dorsal trochlear surface; (1) equal; (2) greater than this
measure.
Ulna, bicipital scar: (0) absent; (1) present as a slightly raised scar (2) present as a
prominent tubercle.
Ulna, brachial scar: (0) absent; (1) present.
Radius, ventroposterior surface: (0) smooth; (1) with muscle impression along most of
surface; (2) deep longitudinal groove.
Ulnare: (0) absent; (1) present.
Ulnare: (0) bulky, “heart-shaped” little differentiation into short dorsal and ventral rami; (1)
V-shaped, well-developed dorsal and ventral rami.
Ulnare, ventral ramus ("crus longum," Baumel and Witmer, 1993): (0) shorter than dorsal
ramus ("crus breve"); (1) same length as dorsal ramus; (2) longer than dorsal ramus.
Semilunate carpal and metacarpals: (0) no fusion; (1) incomplete proximal fusion; (2)
complete proximal fusion; (3) complete proximal and distal fusion.
Semilunate carpal, position relative to metacarpal I: (0) over entire proximal surface; (1)
over less than 1/2 proximal surface.
Metacarpal III, anteroposterior diameter as a percent of same dimension of Metacarpal II:
(0) approximately equal or greater than 50%; (1) less than 50%.
Metacarpal I, extensor process: (0) absent, no anteroproximally projected muscular process;
(1) present.
Metacarpal I, anterior surface: (0) roughly hourglass shaped proximally at least moderately
expanded anteroposteriorly and constricted just before the flare of the articulation for
phalanx I; (1) anterior surface broadly convex.
Metacarpal I, distal articulation with phalanx I: (0) gynglimoid; (1) shelf.
Pisiform process: (0) absent; (1) present.
Carpometacarpus, ventral surface, supratrochlear fossa, deeply excavating proximal surface
of pisiform process: (0) absent; (1) present.
Intermetacarpal space (between metacarpals II and III); (0) reaches proximally as far as the
distal end of metacarpal I; (1) terminates distal to end of metacarpal I.
Carpometacarpus, distal end, metacarpals II and III, articular surfaces for digits: (0)
metacarpal II sub-equals or surpasses metacarpal III in distal extent; (1) metacarpal III
extends farther.
Intermetacarpal process or tubercle: (0) absent; present as (1) raised scar (2) tubercle or
flange.
Manual digit II, phalanx 1: (0) subcylindrical to subtriangular; (1) strongly dorsoventrally
flattened.
Manual digit II: (0) phalanx 1 shorter than phalanx 2; (1) longer.
Manual digit II, phalanx 2: (0) subcylindrical to subtriangular; (1) strongly depressed
posteriorly.
Ilium, ischium, pubis, proximal contact: (0) unfused; (1) fused in adult.
Ilium/ischium, distal fusion, closure of the “ilioishiatic fenestra”: (0) absent; (1) present.
Ischium: (0) forked; (1) straight, no dorsal process.
156.
157.
158.
159.
160.
161.
162.
163.
164.
165.
166.
167.
168.
169.
170.
171.
172.
173.
174.
175.
176.
177.
178.
179.
Ischium and pubis: (0) not subparallel; (1) subparallel and posteriorly directed; (2)
posteriorly directed and appressed.
Laterally projected process on ischiadic peduncle/antitrochanter: (0) directly posterior to
acetabulum; (1) posterodorsal to acetabulum.
Preacetabular tubercle, “pectineal process”: (0) absent; (1) present as a small tubercle or
flange; (2) extremely well developed.
reacetabular ilii: (0) approach on midline, open, or cartilaginous connection; (1) coossified,
dorsal closure of “iliosynsacral canals”.
Preacetabular ilii extend cranial to first sacral vertebrae: (0) no free ribs overlapped, no
parapophyseal facets on sacral ribs; (1) one or more ribs overlapped.
Postacetabular ilii: (0) dorsoventrally oriented; (1) mediolaterally oriented.
Postacetabular ilii, ventral surface, renal fossa developed: (0) absent; (1) present.
Ilium, m. cuppedicus fossa as broad mediolaterally oriented surface directly anteroventral
to acetabulum: (0) present; (1) surface absent, insertion variably marked by a lateral fossa
anterior to acetabulum.
Ischium, distinct proximal anteroventral process: (0) absent; (1) present developed as a
small flange or raised scar contacting /fused with pubis and demarcating the obdurator
foramen distally.
Pubis: (0) sub-oval in cross section; (1) compressed mediolaterally.
Pubes, distal contact: (0) contacting, variably coossified into a symphysis: (1)
non-contacting.
Distal end of pubes: (0) expanded, flared; (1) straight, subequal in proportions with rest of
pubis.
Femur, fossa for insertion of lig. capitis femoris: (0) absent; (1) present.
Femur, posterior trochanter: (0) present; (1) absent.
Femur, posterior trochanter and lateral ridge (Norell and Makovicky, 1999): (0) weakly
projected muscular ridges; (1) hypertrophied “shelf -like” conformation; (2) not present.
Femur, lesser and greater trochanters: (0) separated by a notch; (1) developed as a single
trochanteric crest.
Femur, intercondylar groove (“patellar groove” of other authors, but see Baumel and
Witmer, 1993): (0) developed only distally; (1) extending around distal end and onto
posterodorsal surface.
Femur: (0) ectocondylar tubercle and lateral condyle separated by deep notch; (1)
ectocondylar tubercle and lateral condyle form single trochlear surface.
Femur, posterior projection of the lateral border of the distal end, continuous with lateral
condyle: (0) absent; (1) present.
Laterally projected fibular trochlea: (0) absent; (1) shelf-like projection.
Femur, popliteal fossa: (0) a groove open distally and bounded medially and laterally by
narrow condyles; (1) closed distally by expansion of both condyles (primarily the medial).
Calcaneum and astragalus: (0) unfused to each other or tibia in adult; (1) fused to each
other, unfused to tibia; (2) complete fused to each other and tibia.
Tibiotarsus, cnemial crest(s): (0) one; (1) two.
Tibia: (0) medial condyle projecting farther anteriorly than lateral; (1) equal in anterior
projection.
180. Tibia, extensor canal: (0) absent; (1) an emarginate groove; (2) groove bridged by an
ossified supratendinal bridge.
181. Tibia/tarsal formed condyles, tuberositas retinaculi extensoris (Baumel and Witmer, 1993)
indicated by short medial ridge or tubercle proximal to the condyles close to the midline
and a more proximal second ridge on the medial edge (0) absent; (1) present.
182. Tibia/tarsal formed condyles, mediolateral widths: (0) medial condyle wider; (1)
approximately equal; (2) lateral condyle wider.
183. Tibia/tarsal formed condyles: (0) gradual sloping medial constriction of condyles; (1) no
medial tapering of either condyle.
184. Tibia/tarsal formed condyles, intercondylar groove: (0) mediolaterally broad,
approximately 1/3 of the width of anterior surface; (1) less than 1/3, of the total anterior
surface.
185. Tibiotarsus, extension of the articular surface for distal tarsals/tarsometatarsus: (0) no
posterior extension of trochlear surface or restricted to distal-most edge of the posterior
surface; (1) well-developed posterior extension, sulcus cartilaginis tibialis of Aves
(Baumel and Witmer, 1993); distinct surface extending up the posterior surface of the
tibiotarsus; (2) with well-developed, posteriorly projecting medial and lateral crests.
186. Tibia-and proximal tarsals, distal-most mediolateral width: (0) wider than mid-point of
shaft giving distal profile a weakly developed triangular form; (1) approximately equal to
shaft width, no distal expansion of whole shaft though condyles may be variably splayed
mediolaterally.
187. Fibula: (0) reaches tarsal joint articulating into distinct socket formed between the proximal
tarsals and the tibia; (1) reduced in length, does not reach tarsal joint.
188. Distal tarsals and metatarsals, fusion: (0) distal tarsals fuse to metatarsals; (1) distal tarsals
fuse to metatarsals and proximal metatarsals coossify; (2) distal tarsals as well as proximal
and distal coossification of metatarsals (3) extreme distal fusion, distal vascular foramen
closed.
189. Metatarsal V: (0) present; (1) absent.
190. Metatarsal III: (0) proximally in plane with II and IV; (1) proximally pinched between II
and IV.
191. Tarsometatarsus, intercondylar eminence: (0) absent or very weakly protuberant; (1) well
developed, globose.
192. Tarsometatarsus, hypotarsus: (0) absent; (1) developed as a posterior projection with a flat
posterior surface; (2) projection, with distinct crests and grooves; (3) at least one groove
enclosed by bone posteriorly.
193. Tarsometatarsus, proximal vascular foramen(a); (0) absent; (1) one, between metatarsals III
and IV; (2) two.
194. Metatarsal I: (0) straight; (1) “J shaped” (Chiappe, 1993).
195. Metatarsal I: (0) curved or distally deflected but not twisted, ventral surface convex; (1)
deflected and twisted such that medioventral surface is concave proximal to trochlear
surface for phalanx I.
196. Metatarsal II, dorsal surface, tubercle: (0) absent; (1) present.
197. Metatarsal II, distal plantar surface, “fossa for metatarsal I”: (0) absent; (1) present.
198. Metatarsal II, articular surface for first phalanx: (0) gynglimoid; (1) rounded.
199. Metatarsals, comparative mediolateral width: (0) metatarsal IV approximately the same
width as metatarsals II and III; (1) metatarsal IV narrower than MII and MIII.
200. Metatarsals, comparative trochlear width: (0) II approximately the same size as III and IV;
(1) II broader than III and IV
201. Distal vascular foramen: (0) simple, passing dorsoplantarly; (1) forked, two exits, plantar
and distally between metatarsals III and IV.