Chapter 19: The History of Life 19–1 The Fossil Record A. The Fossil Record B. How Fossils Form C. Dating Fossils Relative Dating Radioactive Dating D. Geologic Time Scale Eras (Paleozoic, Mesozoic, Cenozoic) Periods The Fossil Record • Provides evidence • • about history of life on Earth Shows how different groups of organisms change over time More than 99% of all species that have ever lived on Earth have become extinct. Archaeopteryx How Fossils Form • How Fossils Form: Active Art http://www.phschool.com/webcodes10/index.cfm?fuseaction=home.gotoWebCode&wcprefix=cbe&wcsuffix=5171 • Lucy’s fossil: Water carries small rock particles to lakes and seas. http://www.pbs.org/wgbh/evolution/library/04/3/l_043_01.html Dead organisms are buried by layers of sediment, which forms new rock. The preserved remains may later be discovered and studied. Dating Fossils Relative Dating Radioactive Dating How old is this Archaeopteryx fossil? 150 million years old Relative Dating • estimate age of a fossil by comparing to other fossils • oldest fossils are usually in deepest layers • index fossils (trilobites) used to compare the ages of rocks and other fossils • cannot determine exact numerical age Radioactive (“absolute”) Dating • calculate numerical age of a • sample based on amount of radioactive isotopes Half-life = length of time required for half of the radioactive atoms to decay • Carbon-14 half-life = 5770 yrs. used to date younger fossils • Potassium-40 half-life = 1.26 billion yrs. used to date older fossils • http://www.pbs.org/wgbh/evolution/library/03/3/l_033_01.html Relative vs. Radioactive Dating Comparing Relative and Radioactive Dating of Fossils Relative Dating Can determine Is performed by Drawbacks Radioactive Dating Age of fossil with respect to another rock or fossil (that is, older or younger) Age of a fossil in years Comparing depth of a fossil’s source stratum to the position of a reference fossil or rock Determining the relative amounts of a radioactive isotope and nonradioactive isotope in a specimen Imprecision and limitations of age data Difficulty of radioassay laboratory methods Geologic Time Scale • represents evolutionary time • major changes in animal/plant fossils at specific rock layers divided geologic time according to major fossil changes Geologic Time Scale Geologic Time Scale First life = 4mya 19–2 Earth’s Early History A. Formation of Earth B. The First Organic Molecules (Miller & Urey) C. How Did Life Begin? Formation of Microspheres Evolution of RNA and DNA D. Free Oxygen E. Origin of Eukaryotic Cells F. Sexual Reproduction and Multicellularity Formation of the Earth • Earth formed about 4.6 • billion years ago. Early earth’s atmosphere contained: Hydrogen cyanide, carbon dioxide, carbon monoxide, nitrogen, hydrogen sulfide, and water vapor. • Little or no oxygen • Hot! No oceans or liquid water, just water vapor in air. Early earth Approx. 4 billion years ago Miller-Urey Experiment • What were the first organic molecules on earth? Mixture of gases simulating atmospheres of early Earth Spark simulating lightning storms • (1950’s) Miller & Urey made amino acids by passing sparks through a mixture of hydrogen, methane, ammonia, and water vapor. • (1995) one of Miller’s experiments produced cytosine & uracil (found in RNA) • experiments suggest how simple compounds found could combine to form organic compounds needed for life Condensation chamber Water vapor Cold water cools chamber, causing droplets to form Liquid containing amino acids and other organic compounds How Did Life Begin? • Under certain conditions, large organic molecules can sometimes form tiny bubbles called proteinoid microspheres. • These are NOT cells, but have some of the characteristics of living things. Selectively permeable membrane Can store and release energy Free Oxygen • No oxygen in early Earth’s • • atmosphere. (3.5 bya) Ancient photosynthetic organisms (cyanobacteria) produced a rise in oxygen in the Earth’s atmosphere (2.2 bya) Cyanobacteria produce stromatolite formation in the ocean Endosymbiotic Theory Chloroplast Aerobic bacteria Ancient Prokaryotes Nuclear envelope evolving Plants and plantlike protists Photosynthetic bacteria Mitochondrion Primitive Photosynthetic Eukaryote Animals, fungi, and non-plantlike protists Ancient Anaerobic Prokaryote Primitive Aerobic Eukaryote Eukaryotic cells arose from living communities formed by prokaryotic organisms. Endosymbiotic Theory Video The Evolution of Cells Sexual Reproduction & Multicellularity • Eukaryotic cells began to reproduce sexually, producing more genetic variation, leading to evolution of multicellular Ancient jellyfish fossil organisms. multicellular organism Evolution of Life Concept Map Evolution of Evolution Life ofConcept Map Life Early Earth was hot; atmosphere contained poisonous gases. Earth cooled and oceans condensed. Simple organic molecules may have formed in the oceans.. Small sequences of RNA may have formed and replicated. First prokaryotes may have formed when RNA or DNA was enclosed in microspheres. Later prokaryotes were photosynthetic and produced oxygen. An oxygenated atmosphere capped by the ozone layer protected Earth. First eukaryotes may have been communities of prokaryotes. Multicellular eukaryotes evolved. Sexual reproduction increased genetic variability, hastening evolution. Geologic Time Websites 1. Go to Biology Page Website. 2. Click on Unit 5 Evolution 3. Look under WebLinks column. 4. Explore the following websites: 1. Geologic Time 2. A Brief History of Life 19–4 Patterns of Evolution Macroevolution: large scale evolutionary patterns that occur over long periods of time. The six types are: 1. 2. 3. 4. 5. 6. Extinction Adaptive Radiation Convergent Evolution Coevolution Punctuated Equilibrium Developmental Genes and Body Plans Extinction • More than 99% of all species • • that have ever lived are now extinct. Mass extinctions lead to bursts of evolution that produce many new species. Ex. the extinction of the dinosaurs cleared the way for the evolution of mammals and birds. • http://www.pbs.org/wgbh/evolution/library/03/2/l_032_02.html • http://www.pbs.org/wgbh/evolution/extinction/inde x.html K-T event (65 mya) 70% of species died in a mass extinction Adaptive Radiation • Example = Birds on the Hawaiian islands evolved from a common ancestor over millions of years. • The birds that survived were most adapted to their environment. Birds on Hawaiian Islands Convergent Evolution • Unrelated organisms resemble one another. • Ex. all organisms above have wings, adapted for flying, however they evolved from different ancestors. Coevolution • Two species evolve in response to changes in each other over • time. Ex. hawk moth evolved and has long feeding tube to suck nectar from orchid Plants and pollinators co-evolve Punctuated Equillibrium • Long, stable periods interrupted by brief periods of more rapid • change. Ex. peppered moths evolved rapidly after industrial revolution. Developmental Genes & Body Plans • All animals have master control genes that code for • the a basic body plan. Mutations cause variations in those body plans. • http://www.pbs.org/wgbh/evolution/library/03/4/l_034_04.html Macroevolution Flowchart Species that are Unrelated form Related in under under in in Interrelationshiops Similar environments Intense environmental pressure Small populations Different environments can undergo can undergo can undergo can undergo can undergo Coevolution Convergent evolution Extinction Punctuated equilibrium Adaptive radiation