What is a Protist? How are Protists related to other eukaryotes? Does everyone agree how to classify protists? • No, at present, biologists do not agree how to classify protists • The amount of diversity among the protists, is much greater than within or between the other three eukaryotic kingdoms The Protist Dilemma • Protists are grouped together solely because they are not fungi, plants or animals • Furthermore, many protists are more closely related to members of other eukaryotic kingdoms than they are to other protists. Current Protist classification • It has been proposed that the protista kingdom be divided into six groups or clades • Today, while we still use the term Protist, this is not a single kingdom, but a collection of organisms in six clades What is a Protist? • A protist is a eukaryote (has a nucleus) • A protist is any eukaryote that is not a plant, animal or fungus Evolution of Protista Endosymbiont Hypothesis Are all protists unicellular? • No, although most are unicellular, some protists are colonial, and some like the giant kelp are multicellular. Unicellular Colonial Multicellular How do Protists Move? Some move with flagella • Long whip-like projections • One to two per cell • Examples – Trypanosoma – Euglena Trypanosoma Euglena Two flagella No cell wall Chloroplasts How do Protists move? Some move with cilia Cilia can be used for feeding and movement Cilia are short and used like oars on a boat Example Paramecium Ciliates - Paramecium Lysosomes Trichocysts Oral groove Gullet Anal pore Contractile vacuole Micronucleus Macronucleus Go to Section: Food vacuoles Cilia Some do not move Those that do not move produce spores and live as parasites Plasmodium causes malaria Cryptosporidium spreads through contaminated drinking water and caused intestinal disease Excavates: feeding groove, flagella • Diplomonads – Giardia is an intestinal parasite that causes cramping and diarrhea • Discicristates – Euglena is free living and can use its chloroplast for photosynthesis or can live as a heterotroph – Trypansoma causes African sleeping sickness; carried by tsetse flies Euglena Chloroplast Carbohydrate storage bodies Gullet Pellicle Flagella Go to Section: Eyespot Nucleus Contractile vacuole Chromalveolates: very diverse group; most are photosynthetic • Phaeophytes = multicellular brown algae • Chrysophytes = unicellular golden algae • Diatoms = unicellular algae with intricate silicon dioxide (silica) shells • Ciliates = paramecium are not photosynthetic • Dinoflagellates = half are photosynthetic, half are heterotrophs; some are luminescent • Apicomplexans = parasitic Plasmodium Brown algae Phaeophytes • Photosynthetic • Chlorophylls a and c • Brown accessory pigment fucoxanthin • Multicellular • Giant kelp, Fucus Photosynthetic protists • Chrysophytes “Golden plants” Gold-colored chloroplasts Cell walls contain pectin instead of cellulose Store food as oil rather than starch Can form thread like colonies Photosynthetic protists Diatoms Glass like cell walls Cell walls contain silicon (Si) Cell walls like petri dish Photosynthetic protists Dinoflagellates Luminescent “Fire plants” Half photosynthetic Half heterotrophs Two flagella Apicomplexan • Plasmodium • Mosquito borne parasites like the species that causes malaria Cercozoa, Foraminiferan, Radiolarian • Have pseudopods • Many produce protective shells Heliozoan Foraminiferans Rhodophytes Red Algae • Chlorophyll a • Red accessory pigment – phycobilin • Absorbs blue light • Grows very deep • Multicellular • Nori Ecology of photosynthetic protists • Base of the food chain • Half of the photosynthesis on earth is carried out by phytoplankton Ecology of photosynthetic protists • Algal blooms Caused by too much pollution or nutrients Deplete water of oxygen Kill fish and invertebrates Dinoflagellates cause “red tides” Red tides produce toxins which can be taken in by shellfish. Eating these shellfish can cause illness, paralysis and death Green algae • Phylum Chlorophyta • Same chlorophyll and cell wall composition as green plants • Chlorophyll a and b • Store food as starch • Found in fresh and salt water and on land • Unicellular, colonial and multicellular • Now classified with plants Unicellular green algae • Chlamydomonas Lives in ponds, ditches and wet soil Egg shaped Two flagella Large, cup-shaped chloroplast Colonial green algae • Spirogyra Filamentous Forms threadlike colonies Spiral chloroplasts • Volvox Hollow spheres 500 – 50,000 identical cells Some cell specialization Human uses of algae • Oxygen • Food (nori; thickening agent (carrageenan) in ice cream, egg nog, chocolate, salad dressing) • Industry (plastics, waxes, paints, lubricants) • Science labs (agar) • Alternation of generation – alternating between diploid and haploid organisms • Diploid – having two copies of each chromosome • Haploid – having one copy of each chromosome • Gametophyte – haploid gamete producing organism • Sporophyte – diploid spore producing organism Heterotrophic protists • Amoebozoa = Amoebas use pseudopods for movement and feeding • Ciliates = Paramecia use cilia to move food to gullet; food vacuoles and lysosomes digest the food; waste is released through the anal pore • Slime Molds and Water Molds absorb food through their cell walls from dead or decaying matter; decomposers Section 20-2 An Amoeba Contractile vacuole Pseudopods Nucleus Food vacuole Go to Section: Slime molds Slime molds Water molds • Cells are multinucleate • Cell walls of cellulose • White fuzz on dead fish in water • Plant parasites on land Cause potato blight responsible for potato famine Reproduction in water molds • Can produce sexually and asexually • Motile (swimming) spores • Antheridium produces sperm • Oogonium produces eggs Mutualistic relationships • Zooxanthellae – live inside coral and provide food through photosynthesis • Trychonympha – live in the gut of termites and digest cellulose