Exam 2 • Next Thursday • Topics: – Development, Trophic relationships – Hoeinghaus et al. large-river food webs – Illinois macroinvertebrate paper Trichoptera • Large group ~ 1350 spp. North America • Closely related to Lepidoptera • Holometabolous Trichoptera in most freshwater habitats • Spring and seepage streams, rivers • Lakes • Marshes, temporary ponds Trichoptera diet • Larvae eat algae, decaying vascular plants, and microorganisms on plants • Few larvae are predators • Best known for nets, retreats, cases = silk from labrum • Several forms Net Spinners Net Spinners Net Spinners Purse Case Builders Saddle Case Builders Tube Case Builders (portable) Other cases Free Living Adults • Eggs in gelatinous case attached to submerged vegetation or log Morphology • Anal prolegs have anal claw • Mandibles curved, often crossing each other ID of Trichoptera: 12 families • Anal claw comb-shaped, case is snailshaped • Helicopsychidae Hook-shaped anal claw • Dorsum of all three thoracic segments covered in sclerotized plates • Abdomen with ventral rows of branched gills • Hydropsychidae • Common netspinners • Dorsum of all three thoracic segments covered in sclerotized plates • No gills on abdomen, portable cases • Hydroptilidae = micro caddisflies (2-6 mm) Thoracic segment 3 membranous • Thoracic segment 2 membranous or small plates • Abdominal segment 9 membranous • Labrum T-shaped • Philopotamidae Thoracic segment 3 membranous • Thoracic segment 2 membranous or small plates • Abdominal segment 9 membranous • Labrum rounded, sclerotized • Trochantin pointed • Polycentropodidae – No cases Thoracic segment 3 membranous • Thoracic segment 2 membranous or small plates • Abdominal segment 9 sclerotized • Abdominal segment 1 with hump, prosternal horn • Phryganeidae – Cases of plant fragments Thoracic segment 3 membranous • Thoracic segment 2 membranous or small plates • Abdominal segment 9 sclerotized • Abdominal segment 1 w/o hump, no prosternal horn • No case • Rhyacophilidae • Thoracic segment 2 membranous or small plates • Abdominal segment 9 sclerotized • Abdominal segment 1 w/o hump, no prosternal horn • Tortoise-like case • Glossosomatidae Thoracic segment 3 membranous • Thoracic segment 2 sclerotized • Antennae long, hook-shaped anal claws • Leptoceridae = long-horned caddisflies Thoracic segment 3 membranous • Thoracic segment 2 sclerotized • Antennae short • Tarsal claw on hind leg smaller than others • Molannidae = hood-cased caddisflies Thoracic segment 3 membranous • • • • Thoracic segment 2 sclerotized Antennae short Tarsal claws similar on all legs Abdominal segment 1 no hump, pronotum divided by crease • Brachycentridae Thoracic segment 3 membranous • • • • • Thoracic segment 2 sclerotized Antennae short Tarsal claws similar on all legs Abdominal segment 1 hump Anterior margin of mesonotum notched on sides of midline, head elongate • Uenoidae Thoracic segment 3 membranous • • • • • Thoracic segment 2 sclerotized Antennae short Tarsal claws similar on all legs Abdominal segment 1 with hump Anterior margin of mesonotum not notched on sides of midline, head round • Limnephilidae