Evolution Chapter 16, 17, and 19 in textbook Darwin’s Theory of Evolution Evolution: the process by which all living organisms have developed and diversified from earlier forms over time. Evolution is Just a Theory – Scientific theory: well-substantiated explanation of some aspect of the natural world that is confirmed through observation and experimentation – A scientific theory can be supported or disproven, but never proven. – Other scientific theories: Cell Theory, Theory of Plate Tectonics, Collision Theory Darwin’s Theory of Evolution Darwin set sail on a voyage around the world and made many observations. • Organism in similar environments around the world had similar characteristics – Australia has different plants and animals than Africa – Australia and Africa have similar grassland environments. – The plants and animals in both grasslands have similar characteristics even though they are different. Darwin’s Theory of Evolution • Species always seemed well-suited for their environment – At the Galapagos the environment on each island is very different. – The animals found on each island seemed to had changed to match the environment. – What can you infer about the environments on Hood and Isabella island by observing the tortoises? Darwin’s Theory of Evolution • Ancient organisms resemble modern organisms. – Fossils: preserved remains or traces of ancient organisms – By studying fossils you can infer why the ancient organism no longer exist. – What modern organism Australopithecus and Neanderthal resemble. Why did they go extinct? Idea’s That Shaped Darwin’s Thinking Geology • In Darwin’s time, most people believed the Earth was a few thousand years old. • Geologists began to gather evidence that the continents have been forming for millions of years. • How did this information help Darwin form his theory on evolution? Idea’s That Shaped Darwin’s Thinking Selective Breeding • How could a farmer use selective breeding to get cow with larger horns? – In order for selective breeding to work you need natural variation. – Natural Variation: individual organisms of the same species can have different characteristics. – Darwin called this process artificial Selection – Artificial Selection : nature provides the variation and humans select those they find useful. – How did this information help Darwin form his theory of evolution? (Darwin did not know about heredity) Idea’s That Shaped Darwin’s Thinking Population Growth • Economist Thomas Malthus realized that the human population was growing. – If the population continued to grow, we would run out of resources. – Once we run out of resources many people would die. • How did this information help Darwin develop his theory on evolution? Darwin’s Theory of Evolution Darwin proposed natural selection as the mechanism of evolution. – Natural selection: the process by which organisms with variations most suited to their environment survive and produce more offspring. 3 criteria for natural selection to occur: 1. Natural Variation 2. Struggle for Existence 3. Survival of the Fittest Darwin’s Theory of Evolution 1. Natural Variation – There must be diversity among the individuals of a population (the have different characteristics). – The traits must be heritable 2. Struggle for Existence – Species must compete for limited resources and avoid predators. – Fitness: the organism’s ability to survive and reproduce. Darwin’s Theory of Evolution • Survival of the Fittest – Only the most “fit” individuals will be able to survive and reproduce to pass on their adaptations. – Adaptation: any inherited characteristic that increases an organism’s fitness (ability to survive and reproduce) – Which insect has the highest fitness? Lamarck’s Theory of Evolution Lamarck proposed a theory of evolution 50 years before Darwin’s theory. Lamarck’s Two Ideas 1. The inheritance of acquired traits. • Lamarck thought that traits acquired during an organisms life could be passed on to the next generation. • How would Lamarck’s explain the evolution of a giraffe’s long neck? Lamarck’s Theory of Evolution 2. Use and Disuse – Lamarck thought that body parts that were not being used were gradually disappearing. – How would Lamarck explain the evolution of a cave fish’s lack of eyes? Pattern’s of Evolution Adaptive Radiation • Adaptive Radiation: When a single species evolves into several different species over time. • All organisms share a common descent: – All living organism evolved from a single common ancestor that lived billions of years ago. – Over time, that organism evolve in different ways to give to rise to all of the modern living organisms. – Organisms are closely related if they share a recent common ancestor. Pattern’s of Evolution How long ago did human and chimpanzee’s share a common ancestor? What is the chimpanzee’s closest living relative? Pattern’s of Evolution How are new species formed during adaptive radiation? Speciation: The formation of a new species through evolution. – Populations must be reproductively isolated from each other – Populations can be isolated by geography, behavior or time. Pattern’s of Evolution 2. Convergent Evolution – – – Unrelated organisms evolve similar adaptations because they live in similar environments Sharks and dolphins do not share a recent common ancestor. They have similar characteristics because they share a similar environment. Pattern’s of Evolution 3. CoEvolution – – – The evolution of two or more species, each adapting to changes in the other Occurs when species interact closely with each other. Ex: Cheetah and Gazelles have co-evolve to become the fastest and 2nd fastest land animals Pattern’s of Evolution 4. Punctuated Equilibrium: – Species do not evolve for long periods of time and then go through short period of rapid evolution. – What causes the period of rapid evolution? • Mass Extinctions: A large number of species die out in a short time. • Isolation: A group of individual gets isolated from the main population. Pattern’s of Evolution 5. Gradualism – Organisms have evolve at a slow and steady rate. – This theory is in contrast with punctuated equilibrium. Evidence of Evolution Homologous Body Structures – Body structures shared by related species that have a similar structure but different functions. – Supports the theory that organisms evolved from a common ancestor. Evidence of Evolution Analogous Body Structures – Body parts in different species that is similar in function but not in structure – Structures evolved in response to a similar environmental challenge. – Supports convergent evolution Evidence of Evolution Embryology – Closely related organisms look more similar in the early stages of embryonic development. Genetics – All living things have the exact same genetic code. – A group of 3 DNA nucleotides will code for the same amino acid in all organisms. – Closely related organisms have a similar DNA sequence. Evidence of Evolution Geologic Time Scale: a timeline of earth’s history. – Major changes in fossils separate segments of time (periods). – The end of each “Era” is usually a mass extinction. – 99% of all species that ever lived on Earth are extinct Evidence of Evolution Scientists must determine the age of the fossil to determine where the organism fits in the geologic time scale. 2 Methods of Determining Age of Fossils 1. Relative Dating – Where is it in the rock layer? – Deeper layer = older fossil – Can’t tell you exactly how old the fossil is, just that it is older or newer than another fossil Put the fossils in order from most modern to oldest. Evidence of Evolution 2. Radioactive Dating – Calculate the age of the sample based on the amount of radioactive isotopes it contains. – half life: the time required for half of the radioactive isotopes to decay. – Half life of carbon-14 is 5,715 years. How old is a fossil if 25% of the carbon 14 is remaining? Origin of Life How did life begin on Earth? • Miller and Urey conducted an experiment to determine if organic molecules could be formed under the conditions of early Earth? – They placed water in a flask and they added the gases that made up the atmosphere at that time. – They boiled and electrocuted it many time. – Eventually this process produced amino acids. Origin of Life • The earliest forms of life had to be anaerobic. Why? • Most living organisms need oxygen for cellular respiration. • What is the main purpose of cellular respiration? • Anaerobic organisms used fermentation produced energy without oxygen. • Cyanobacteria: the first organism to do photosynthesis. • How did photosynthesis change the atmosphere? Origin of Life Endosymbiotic Theory: The mitochondria and chloroplasts of eukaryotic cells were once independent prokaryotic cells. • The prokaryotic cells were engulf by larger cells: – One engulfed cell became the mitochondria – One engulfed cell became the chloroplast • Evidence: The mitochondria and chloroplasts have their own membranes and DNA. Evolution as Genetic Change Evolution, in genetic terms, is a change in the allele frequency a population over time. – Gene Pool: the combined genetic info. of all the members of a population – Relative Frequency: the number of times that an allele occurs in a gene pool compare to the total number of alleles What is the allele frequency of the black and brown alleles? Evolution as Genetic Change Natural selection affects polygenic traits differently than single gene traits. • Single-Gene Trait • Polygenic Trait – Controlled by one gene that has two alleles – Two distinct phenotypes – Ex: Tall or Short Pea Plants – Controlled by more than one gene with many alleles – Many possible genotypes and phenotypes – Ex: Human Height Evolution as Genetic Change Natural selection can affect the distributions of polygenic phenotypes in any of 3 ways: 1. Directional Selection – When individuals at one end of the curve have higher fitness than individuals in the middle or other end. – Entire curve shifts – Favors one of the extreme variations Evolution as Genetic Change 2. Stabilizing Selection – Individual in the center of the curve have higher fitness than those at the ends of the curve. – Center remains stable but narrows entire graph – Favors average individuals – Ex: weight of human infants Evolution as Genetic Change 3. Disruptive Selection: – When individuals at each end of the curve have higher fitness than those in the middle – Favors both extreme variations – Creates two distinct phenotypes (can lead to speciation) Evolution as Genetic Change • Genetic Drift: random change in allele frequency of a small population due to chance. – Does not increase the overall fitness of the population – More likely to occur in small populations. Evolution as Genetic Change • Genetic Equilibrium: when the allele frequencies in a population remain constant over time. – The population is not evolving Hardy-Weinberg Principle A population will not evolve if the following conditions are met: 1. Random Mating 2. Large Population 3. No Migration 4. No mutations 5. No natural selection Evolution as Genetic Change Lamarck’s Theory of Evolution Evaluating Lamarck’s Theory • Lamarck was one of the first to recognize that life changes over time • He also realized that organisms were adapt to their environment • He didn’t know how traits were inherited (acquired traits cannot be inherited) Coevolution Darwin vs. Lamarck Lamarck vs. Darwin