Phylum Chordata the chordates Phylum Chordata chordates • Primary characters include: – Notochord Phylum Chordata Primary Characters • pharyngeal gill slits • dorsal nerve cord • ventral heart • post anal tail Phylum Chordata Secondary Characters • tailed larvae or direct development • dorsal and ventral blood vessels; closed circulatory system • endoskeleton Phylum Chordata • Includes two groups: – Protochordata or Acraniata – those without a cranium – Craniata – those with a cranium Phylum Chordata Group Protochordata (=Acraniata) (the first chordates) • Includes two subphyla: – Urochordata – tunicates or sea squirts – Cephalochordata- lancelets Subphylum Urochordata (tunicates or sea squirts) • about 2000 species • all marine, freeliving or sessile, solitary or colonial • adult has no coelom; no segmentation, no bones • have a test or tunic; made of cellulose Subphylum Urochordata (tunicates or sea squirts) • dorsal atrium, chamber used for filter feeding • there is a heart, with colorless blood • respiration is probably function of pharynx Subphylum Urochordata (tunicates or sea squirts) • notochord is restricted to tail, and is found only in larvae (except for one classthere are 3 classes) • larvae are called "tadpole larvae“ Subphylum Urochordata (tunicates or sea squirts) • most species are hermaphroditic, fertilization is external • some species reproduce asexually through budding Subphylum Cephalochordata (lancelets) • about 23 species • all marine, freeliving or sessile, solitary or colonial, live on bottom • no well developed head Subphylum Cephalochordata (lancelets) • many gill slits in pharyngeal basket, act in excretion and respiration • have a tail; projects behind anus • muscles arranged in blocks- myotomes • have a notochord made of muscle fibers Subphylum Cephalochordata (lancelets) • are microscopic suspension feeders • has closed circulatory system, with colorless blood • sexes are separate; fertilization is external Phylum Chordata Group Craniata (the vertebrates) • Includes one subphylum: – Subphylum Vertebrata Phylum Chordata Subphylum Vertebrata • Contains a wide variety of animals that include: – – – – – – – – Class Myxini – hagfishes Class Cephalaspidomorphi – lampreys Class Chondrichthyes – sharks and rays Class Osteichthyes- bony fishes Class Amphibia – amphibians Class Reptilia – reptiles Class Aves – birds Class Mammalia - mammals Subphylum Vertebrata • Divided into: – Without useable jaws • Agnatha – With useable jaws • Gnathostomata Subphylum Vertebrata • Earliest vertebrates were jawless: – Agnathans • Earliest of these were Ostracoderms – 500 million years ago Subphylum Vertebrata • Earliest jawed vertebrates or Gnathastomes • Earliest of these were Placoderms – 400 million years ago