Phylum Porifera The Sponges 1 Phylum Porifera Phylum Porifera Overview Most primitive of the multicellular animals – Sponges – – 2 There is some debate if sponges are complex colonial protozans and not metazoans. Over 7,000 species, approximately 40 species that occur in local waters 2% of all sponges are freshwater, none are terrestrial Phylum Porifera Phylum Porifera Overview 3 Sponges occur in shallow water habitats and vary widely in size (up to 1m. high) and shape – Unlike most metazoans they lack: All sponges are sessile filter feeders Phylum Porifera Sponge Diversity Erect Rope Sponge Black-ball sponge Yellow Tube Sponge 4 Phylum Porifera Porifera Anatomy 5 Spongocoel Ostia Oscules Phylum Porifera Collar Cells Choanocytes 6 Choanocytes: (collar cells) act as a pump to bring water into the sponge Phylum Porifera Sponge Support Collagen is found between the inner canals and chambers – Ameboid cells located in the mesohyl, have different roles – – 7 Mesohyl Archeocytes Sclerocytes Phylum Porifera Sponge Support 8 Phylum Porifera Spicules Collagen is stiffened by adding microscopic mineral accretions or additional protein fibers (spongin) or both. – 9 Spicules: skeleton structures, made of calcium carbonate (CaCO3) or silicon dioxide (SiO2). Spicules Phylum Porifera Sponge Types Porifera Reproduction Asexual budding Regeneration: can regenerate from broken pieces Sexual – 11 Usually hermaphroditic with male and female cells scattered throughout the connective tissue. Phylum Porifera Porifera Larvae Neighboring sponges are fertilized by sperm entering through the ostia – 12 Ciliated mouth less larvae (parenchymella) is released. Parenchymella Phylum Porifera Porifera Classification Phylum Porifera – – – – 14 Class Calcarea Class Demospongiae Class Hexactinellida Sclerospongiae is no longer considered a class Phylum Porifera Class Calcarea 15 Have spicules made of calcium carbonate Mostly small in size (<15 cm.), and form irregular masses Never contain spongin, restricted to shallow water, and strictly marine Phylum Porifera Class Demospongiae (Most sponges) Have spicules made of silicon dioxide (SiO2) or spongin or a combination of both Most sponges belong to this class (90%) – 16 Nearly all are leuconoid body type Mostly found on the continental shelf Spongia spp. (Bath sponge) Phylum Porifera Class Hexactinellida (Glass sponges) 17 Spicules are made of silica Usually found in deep water on soft substrates in the tropics 200-1,000m. Spicules are six pointed and have a lattice-like structure Cup, vase or urn shape Euplectella (Deep sea Glass sponge) Phylum Porifera