Plant Repsonses & Hormones Nancy G. Morris Volunteer State Community College Campbell, 5th edition, Chapter 39 Plant Hormones Hormone = Gr. “to excite” 1) active in small amounts 2) produced in one part of plant & transported to another for action 3) action is specific for that site Each has a Multiplicity of Effects Depending on site of action Developmental stage of plant Concentration of hormone Auxins stimulate growth but too much inhibits growth functions: 1) root initiation, stem elongation 2) retard abscission (loss) of leaves & fruits (Figure 39.10) 3) stimulates cell differentiation 4) apical dominance (Figure 39.6) Apical dominance Gibberellins discovered due to a fungus Giberella which causes Japanese “foolish” rice seedling disease Figure 39.7 induces flowering stimulates growth by increasing cell size & numbers (Figure 39.8) “Foolish seedling disease” in rice 39.11 Effect of Gibberellin Dwarf pea plant treated with gibberellin Cytokinins 1) induces cell division (cytokinesis) 2) affects root growth & differentiation 3) stimulates germination 4) delays senescence (aging); the progression of irreversible change that eventually leads to death Abscisic Acid 1) stimulates abscission 2) converts vegetative buds (active) to dormant buds 3) inhibits growth 39.16 Abscission Ethylene 1) promotes fruit ripening 2) stimulates production of cellulase Photochromes Plant pigment Important in processes where light is a critical factor, such as, flowering in long day short night plants Figure 39.16 Plant Movements due to growth Phototrophism – movement toward (positive) or movement away from (negative) light Geotrophism – movement toward (positive) or away from (negative) center of the earth Thigmotrophism – directional growth responding to contact (tendrils) Photoperiodic control of Flowering Plant Movements not due to Growth Thigmonastic response – touch Photonastic response – shamrock stimulus, e.g. mimosa leaves Sleep response – caused by change in osmotic pressure Sleep movements of a bean plant: Movement of materials Path from roots: 1) epidermis of root & root hair cells 2) roots absorb water, minerals, gases 3) stems conduct Xylem & phloem are vertical conductors Parenchyma of cortex, pith, rays are lateral conductors Movement Root pressure –osmotic pressure & pressure created by active absorption & secretion by root cells; drives materials up xylem Transpiration – evaporation of water through leaf stomata; cohesion of water molecules exerts a pull on columns of water in xylem Guttation – release of water droplets at leaf tips; occurs when too much water is absorbed by plant & when humidity is high Translocation – movement of material from one location to another Guttation Fighting Back…. Response to herbivory (animals eating plants): 1) physical barriers – thorns & spines 2) chemical defenses – distasteful or toxic compounds 3) chemical signaling – salivary enzyme from herbivore triggers a pathway that produces a volatile attractant that recruits a parasitoid wasp. The wasp lays eggs within the caterpillar, and the larvae upon hatching destroy the caterpillar by consuming its tissues. (Figure 39.22)