Global Patterns of Species Richness Global Amphibian Diversity Wallace’s hypotheses for higher species diversity in the tropics: 1. 2. 3. 4. 5. 6. 7. 8. 9. Productivity Spatial heterogeneity; morphological diversity High average rarity – no monocultures Climatic constancy – little seasonal variation Geological age Special adaptations required for “harsh” environments Greater competitive “balance” in tropics Alternating competitive advantages All niches filled – saturation of community Early Island Biogeography Island Habitats in Baltic Sea Reptile and Amphibian Species Area Curve for Caribbean Islands Habitat Islands In the Great Basin Top – Birds, Bottom - Mammals Species Richness related to Island Size and Island Isolation Species-Area Curves for Islands S = cAz where: S = number of species A = area of island c = constant for habitat type - forest, desert, grassland z = slope of the curve - predicted to be 0.263, but actually varies with dispersal ability An area of island will have fewer species than equal sized area of continent Ant species On New Guinea and Nearby Islands The slope of the line plotted for species-area curve is larger for species with poor dispersal abilities Oceanic islands birds - New Guinea islands beetles - West Indies land plants - Channel Islands Habitat islands birds - Andes paramo birds - mtns of Great Basin mammals - mtns of Great Basin inverts -caves in West Virginia z=0.22 z=0.34 z=0.37 z=0.29 z=0.165 z=0.326 z=0.72 Island Biogeography Species Dispersal to Islands Cape May Warbler on a Ship Krakatoa pre-eruption Looking at Volcano Rakata Krakatoa mid-eruption Map of Krakatoa pre-eruption Landsat Image of Anak Krakatau and Krakatau Dan Simberloff and Mangrove Islands Simberloff’s defaunation experiment on Mangroves Simberloff’s defaunation experiment on Mangroves Results from Simberloff’s Experiment Islands recovered to pre-defaunation number of species, but only 30% of original species resident on each island returned after defaunation. Bracken Fern Bracken fern stand Australia Number of species found on Bracken Fern Professor Axel Meyer – Cichlid Man Lake Malawi Cichlids Adaptive radiation within cichlid lineages