Chapter 35 Plant Structure, Growth, and Development Student Guided Notes Concept 35.1 Plants have a hierarchical organization consisting of organs, tissues, and cells. o A tissue is a _____________________________________________, consisting of one or more types, which perform a specific function. o An _______________________ consists of several types of tissues that work together to carry out particular functions. MONOCOTS EUDICOTS (Previously called DICOTS) One cotyledon __________ cotyledons (Single seed leaf) (Double Seed Leaf) Leaf Venation Veins are __________________ Veins are __________________ Stems Vascular bundles usually complexly Vascular bundles usually arranged arranged in a ___________________ Roots Fibrous _______________________ Flowers Flowers in multiples of ____ Multiples of ____ or ____ Embryos **Remember, Monocots and Eudicots (Dicots) are Phylum ______________________________ and each group is monophyletic. Vascular plants have three basic organs: ___________________, _______________________, and __________________________________. o Vascular plants obtain __________________________ and _____________________ from the soil. o Vascular plants obtain ______________ and ______________________ above ground. To obtain the resources they need, vascular plants have two systems: a subterranean ________________ system and an aerial ________________________ system of stems and _________________________. Roots provide anchorage, absorption, and storage. A root is ___________________________________________________________________________ ____________________________________________________________, absorbs minerals and water, and often stores carbohydrates. They have epidermal tissue, but no waxy cuticle (like leaves do). 35-1 Most eudicots and gymnosperms have a _____________________________ system, consisting of one large vertical root (the taproot.) o The taproot produces many small _________________________________ roots, also called branch roots. o Taproot systems generally penetrate deeply and are well adapted to __________________________. In most monocots, such as grasses, many small (adventitious) roots grow from the stem. o A fibrous root system is usually _______________________________ than a taproot system and is best adapted to shallow soils with light __________________________________. In both taproot and fibrous root systems, absorption of water and minerals occurs near the root tips, where vast numbers of tiny ____________________________ enormously increase the ____________ ___________________________. Stems consist of alternating nodes and internodes. A ___________________ is an organ that raises or separates leaves, exposing them to _____________. A stem consists of alternating nodes, _____________________________________________________ _______________________________________, and internodes, the __________________________ __________________________________________________________________________________. o In the angle formed by each leaf and the stem is an _______________________________________ with the potential to form a lateral shoot or branch. The growth of a young shoot is usually concentrated at its ___________________, where there is an ____________________________ bud, or _____________________________ bud. The presence of a terminal bud is partly responsible for _______________________________ ____________________________, a phenomenon called ______________________________. o If the ________________________________ bud is removed, the axillary buds break dormancy and give rise to lateral shoots complete with their own apical buds, leaves, and axillary buds. o This is why _______________________________ trees and shrubs makes them bushier. Modified shoots with diverse functions have evolved in many plants (examples: tubers and bulbs.) ____________________________ are the main photosynthetic organs of most plants. The leaf is the primary site of ____________________________________ organs in most plants. o They consist of a flattened _________________ and a stalk, the ________________, which joins the leaf to a stem node. They are also covered by a waxy cuticle that helps ______________________. 35-2 Plant organs are composed of three tissue systems: _______________________, _______________________, and ___________________________. The _____________________________________ system is the plant’s outer protective covering (aka epidermis or “skin”.) o The epidermis of leaves and most stems secretes a waxy ________________________, which helps the plant _________________________________. The _____________________________________ system is involved in _______________________ __________________________________________________________________________________. o Xylem conducts _______________________ and ___________________________________ upward from __________________________ into the __________________________. o Phloem transports _____________________, the products of ______________________________, to the roots and sites of growth, such as developing ______________________ and _____________ (up AND down.) The vascular tissue of a root or stem is called the __________________________. The ______________________________________ system is tissue that is neither dermal nor vascular. o Ground tissue is divided into ____________________, internal to vascular tissue, and ______________________, external to the ____________________________ tissue. o The functions of ground tissue include _______________________________, _________________________________, and _______________________________. Plant tissues are composed of three basic cell types: parenchyma, collenchyma, and sclerenchyma. Plant cells are differentiated, with each type of plant cell possessing structural adaptations that make specific functions possible. Parenchyma Have primary walls that are _________________________________________________________; most lack __________________________________________. Often depicted as “typical” plant cells because they generally are the _________________________. Perform most of the ______________________________________________ of the plant, synthesizing and storing various organic products. Examples: photosynthesis, contain plastids, fleshy tissue of fruit 35-3 Most parenchyma cells retain the ability to _____________________ and _____________________ into other cell types under special conditions, such as the repair and replacement of organs after injury to the plant. Thus, these cells can de-differentiate for plant tissue cultures. Collenchyma Have ___________________________________________________ than parenchyma cells, although the walls are _______________________________. They lack _________________ walls. Grouped into strands or cylinders, collenchyma cells help support ___________________________ ___________________________________________________. Sclerenchyma Have thick secondary walls usually strengthened by _____________________________; they function as supporting elements of the plant. Sclerenchyma cells are much more _________________________ than collenchyma cells. Unlike parenchyma cells, sclerenchyma cells cannot _______________________________. o Sclerenchyma cells occur in plant regions that have ________________________________. More on Xylem and Phloem Xylem: The water-conducting elements of xylem, ______________________ and ______________________________, are elongated cells that are ____________ at functional maturity. Tracheids are ____________________________________________________________________, hardened with lignin. Vessel elements are generally _______________________________________________________, and less ___________________________ than tracheids, and are aligned to form long micropipes. Phloem: In the phloem, sucrose, other organic compounds, and some mineral ions move through tubes formed by chains of cells called ______________________________________________. Sieve-tube elements are ___________________________ at functional maturity, although a sievetube element lacks a ____________________, __________________________, and a distinct ______________________________. 35-4 o The end walls, the ________________________________, have pores that facilitate the flow of fluid between cells. Each sieve-tube element has a non-conducting nucleated _________________________________, connected to the sieve-tube element by ___________________________________. Sieve tube elements move the sucrose solution by a process called ___________________________. Concept 35.2 Different meristems generate new cells for primary and secondary growth. Unlike growth of many animals, plant growth is not limited to an ____________________________ or ______________________________ period. o Most plants demonstrate ________________________________________, growing as long as the plant lives. A plant is capable of indeterminate growth because it has perpetually undifferentiated tissues called ________________________________. (Meristems Lifelong growth!) o There are two main types: ___________________ meristems and __________________ meristems. o The pattern of growth depends on the ______________________________ of the meristems APICAL MERISTEMS, located at the tips of ____________________ and in axillary buds of _____________________, supply cells for the plant to grow in length. (primary growth length) o This elongation enables roots to ___________________________________________ and shoots to ________________________________________________________________________________. _________________________ plants also show secondary growth, progressive thickening of roots and shoots where primary growth has ceased. (secondary growth ___________________________) o Secondary growth is produced by __________________________ MERISTEMS, cylinders of dividing cells that extend along the lengths of roots and shoots (“sideways” rather than “up and down”). There are two lateral meristems: o The _____________________________________________ adds layers of vascular tissue called secondary xylem (_________________________) and secondary phloem. o Plants with vascular cambium with lignified cell walls are called woody plants (not herbaceous) o The _____________________________________ replaces the __________________________ with thicker, tougher periderm. 35-5 __________________________ complete their life cycle—from germination to flowering to seed production to death—in a single year or less. The lives of ______________________________ span two years, with _______________________ _______________________________________ in the second year. Plants such as trees, shrubs, and some grasses that live many years are ________________________. Concept 35.3 Primary growth lengthens roots and shoots. Roots show primary growth. The root tip is covered by a thimble-like __________________________, which protects the ________________________________. Growth occurs just behind the root tip in three overlapping zones of cells. The _________________________________________________ includes the root apical meristem. The zone of cell division blends into the ______________________________________________, where cells elongate and is responsible for _________________________________________, including the meristem, into the ________________________. In the zone of differentiation (aka zone of maturation), cells ______________________________ ____________________________________ and become distinct cell types. The primary growth of roots produces the ____________________________, ____________________ tissue, and ______________________________ tissue. Water and minerals absorbed from the soil must enter the plant through the ______________________, a single layer of cells covering the root. o Root hairs greatly increase the __________________________________ of the epidermis. In angiosperm roots, the _________________________ is a vascular cylinder with a solid core of __________________________ and ____________________________. The ground tissue of roots consists of _______________________________ cells that fill the cortex, the region between the vascular cylinder and the epidermis (the innermost layer of cortex is _____________________________________.) When plant stems are cut, roots will develop at the cut end which is opposite the _________________. 35-6 3 Primary Meristems: _________________________________ dermal tissue (epidermis) _________________________________ vascular tissue (stele = vascular bundle in the center of roots; contains xylem and phloem) _________________________________ ground tissue Concept 35.4 Secondary growth increases the diameter of stems and roots in woody plants. Secondary growth, the growth in ______________________________ produced by _________________________ meristems, occurs in stems and roots of ________________________ plants, but rarely in leaves. Secondary growth consists of the tissues produced by the ______________________________ and ______________________________________. o The vascular cambium adds secondary __________________________ (wood) and secondary phloem, increasing vascular flow and support for the _______________________________ system. o The cork cambium produces a tough, thick covering consisting of wax-impregnated cells that protect the stem from water loss and invasion by insects, bacteria, and fungal spores. Bark is ___________________________________________________________________________, including secondary phloem (produced by the vascular cambium), the most recent periderm, and all the outer layers of periderm. Concept 35.5 Growth, morphogenesis, and cell differentiation produce the plant body. The specific series of changes by which cells form ______________________, __________________, and _________________________________ is called development. There are three overlapping processes in development: o Growth is an irreversible ____________________________________________________. o _______________________________________ is the process that gives a tissue, organ, or organism its shape and determines the positions of cell types. (pattern formation = specific structure in specific location) o Cell differentiation is the process by which cells with the same genes _____________________ ______________________________________________________________________________. 35-7 o RECALL: plants have developmental plasticity (totipotency) o When a new plant is grown in culture, the parenchyma must first DE-DIFFERENTIATE and then it can differentiate again. o Cuttings reflect the period in which they were formed (example: to get plant features from the juvenile form, must take cuttings from areas formed in that period) ALSO REMEMBER Plants need adaptations to respond to heat and water loss including: - Stomates that can open and close - A fibrous root system - Mycorrhizae (symbiotic fungus on the roots to help get water) - Waxy cuticle 35-8