Curriculum Vitae
Jeff R. Havig
1
Postdoctoral Researcher
Department of Geology
University of Cincinnati
500 Geology-Physics Building
Cincinnati, OH 45221-0013 jeffhavig@gmail.com http://www.jeffhavig.com/
509-637-6375
Education:
 Doctor of Philosophy in Geology, Emphasis in Biogeochemistry
Arizona State University, 2009
Dissertation Title: Geochemistry of hydrothermal biofilms: Compositions of biofilms in a siliceous sinter-depositing hot spring.
 Master of Science in Geology, Emphasis in Groundwater Studies
Washington State University, 2002
Thesis Title: Effects of perturbation on nutrient pools and loss from an experimental forest mesocosm.
 Bachelor of Science in Chemistry, Emphasis in Environmental Chemistry
Washington State University, 1999
Employment History:
 Postdoctoral Researcher, Dept. of Geology, University of Cincinnati (Start:
August, 2015)
 Visiting Scientist, Dept. of Geology, University of Cincinnati (March 2015 –
August 2015)
 Postdoctoral Researcher, PSARC, PSU (July 2012 – January 2015)
 Postdoctoral Research Faculty, SESE, ASU (January 2010 – December 2011)
 Astrobiology Postdoctoral Research Faculty, SESE, ASU (June – December,
2009)

Curriculum Vitae
Jeff R. Havig
2
Publications: (listed in chronological order from most current)
Havig, J. R., McCormick, M. L., Hamilton, T. L., and Kump, L. R. (2015) The behavior of biologically important trace elements across the oxic/euxinic transition of meromictic
Fayetteville Green Lake, New York, USA. Geochimica et Cosmochimica Acta 165,
389-406. DOI: 10.1016/j.gca.2015.06.024
Boyd, E.S., Hamilton, T.L., Havig, J.R., Skidmore, M., Shock, E.S. (2014)
Chemolithotrophic primary production in a subglacial ecosystem. Applied and Environ.
Microbiology 80: 6146-6132. DOI:10.1128/AEM.01956-14
Hamilton, T. L., Koonce, E., Howells, A., Havig, J. R., Jewell, T., de la Torre, J. R.,
Peters, J. P., and Boyd, E. S. (2014) Competition for ammonia influences the structure of chemotrophic communities in geothermal springs. Applied and Environ. Microb.
80(2), 653-661. DOI: 10.1128/AEM.02577-13
Paukert, A. N., Matter, J. M., Kelemen, P. B., Shock, E. L., and Havig, J. R. (2012)
Reaction path modeling of enhanced in situ CO
2
mineralization for carbon sequestration in the peridotite of the Samail Ophiolite, Sultanate of Oman. Chemical Geology 330-
331:86-100. DOI: 10.1016/j.chemgeo.2012.08.013
Boyd, E. S., Fecteau, K. M., Havig, J. R., Shock, E. L., and Peters, J.W. (2012) Modeling the Habitat Range of Phototrophs in Yellowstone National Park: Toward the
Development of a Comprehensive Fitness Landscape. Front. Microbio.
3:221. DOI:
10.3389/fmicb.2012.00221
Miller-Coleman, R.L., Dodsworth, J.A., Ross, C.A., Shock, E.L., Williams, A.J.,
Hartnett, H. H., McDonald, A. I., Havig, J. R., and Hedlund, B. P. (2012) Korarchaeota
Diversity, Biogeography, and Abundance in Yellowstone and Great Basin Hot Springs and Ecological Niche Modeling Based on Machine Learning. PLoS ONE 7(5): e35964.
DOI:10.1371/journal.pone.0035964
Swingley, W. D., Meyer-Dombard, D. R., Shock, E. L., Alsop, E. B., Falenski, H. D.,
Havig, J. R., and Raymond, J. (2012) Coordinating Environmental Genomics and
Geochemistry Reveals Metabolic Transitions in a Hot Spring Ecosystem. PLoS ONE
7(6): e38108. DOI:10.1371/journal.pone.0038108
Loiacono, S. T., Meyer-Dombard, D. R., Havig, J. R., Poret-Peterson, A. T., Hartnett, H.
E. and Shock, E. L. (2012), Evidence for High-Temperature in situ nifH Transcription in an Alkaline Hot Spring of Lower Geyser Basin, Yellowstone National Park.
Environmental Microbiology , 14: 1272–1283. DOI: 10.1111/j.1462-2920.2012.02710.x

Curriculum Vitae
Jeff R. Havig
Havig, J. R., Meyer-Dombard, D. R., Raymond, J., and Shock, E. L. (2011) Merging
Isotopes and Metagenomics: Coupling Biofilm C and N Isotopes and Metagenomics in a
Siliceous Sinter-Depositing Hot Spring. JGR-Biogeosciences 116: G01005. DOI:
10.1029/2010JG001415
Cox, A., Shock, E. L., and Havig, J. R. (2011) The Transition to Microbial
Photosynthesis in Hot Spring Ecosystems. Chemical Geology 280(3-4): 344-351. DOI:
10.1016/j.chemgeo.2010.11.022
Meyer-Dombard, D. R., Swingley, W., Raymond, J., Havig, J., Shock, E. L., and
Summons, R. E. (2011) Hydrothermal Ecotones and Streamer Biofilm Communities in the Lower Geyser Basin, Yellowstone National Park. Environmental Microbiology
13(5): 1-16. DOI: 10.111/j.1462-2920.2011.02476.x
Boyd, E. S., Lange, R. K., Mitchell, A. C., Havig, J. R., Hamilton, T. L., Shock E. L.,
Peters, J. W., Skidmore, M. (2011) Diversity, Abundance, and Potential Activity of
Nitrifying and Denitrifying Microbial Assemblages in a Subglacial Ecosystem. Appl.
Environ. Microbiol.
July 2011 vol. 77 no. 14 4778-4787. DOI: 10.1128/AEM.00376-11
Keller, C. K., R. O'Brien, J. R. Havig, J. L. Smith, B. T. Bormann, and D. Wang. (2006)
Tree Harvest in an Experimental Sand Ecosystem: Plant Effects on Nutrient Dynamics and Solute Generation. Ecosystems 9(4): 634-646. DOI: 10.1007/s10021-006-0162-6
Mount, G. H., Rumburg, B., Havig, J., Lamb, B., Westberg, H., Yonge, D., Johnson, K., and Kincaid, R. (2002) Measurement of Atmospheric Ammonia at a Dairy Using
Differential Optical Absorption Spectroscopy in the Mid-Ultraviolet. Atmospheric
Environment 36(11): 1799-1810.
Manuscripts:
Havig, J. R., Hamilton, T. L., Bachan, A. V., and Kump, L. R. (est. 2015) Interpreting the sulfur isotope record through a molecular lens. (Invited Review, Chemical Geology)
Havig, J. R., Hamilton, T. L., Boyd, E. S., Meyer-Dombard, D. R., and Shock, E. L. (est.
2015) Effects of Geochemical Change on Microbial Community Structure in Hot Spring
Ecosystems.
Havig, J. R., Hamilton, T. L., Boyd, E. S., Zolotova, N., and Shock, E. L. (est. 2015)
Influence of Subsurface Hydrothermal Processes on the Availability and Use of Fixed
Nitrogen for Microbial Metabolism.
In Prep:
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Curriculum Vitae
Jeff R. Havig
Havig, J. R., McCormick, M. L., Bachan, A. V., Hamilton, T. L., McClure, B. E.,
Sowers, T., and Kump, L. R. (2014) High resolution geochemical sampling coupled to input modelling at meromictic Fayetteville Green Lake, N.Y.
Havig, J. R., McCormick, M. L., Bachan, A. V., and Kump, L. R. (2014) High resolution sampling reveals a complex geochemical redoxcline in a meromictic lake.
Havig, J. R., Moore, G., Zolotova, N., and Shock, E. L. (2014) Geochemistry of hydrothermal biofilms: Uptake and sequestration of major and trace elements in hot spring biofilms.
Meeting Abstracts: (listed in chronological order from most current)
Havig, J. R., McCormick, M. L., Bachan, A., and Kump, L. R. (2014) High resolution sampling reveals a complex geochemical environment at Fayetteville Green Lake, N.Y.
Goldschmidt Conference, Sacramento, CA, June, 2014.
Havig, J. R., Hamilton, T. L., Bachan, A. (2014) Interpreting the ancient sulfur-isotope signal from a metabolic pathway perspective. Goldschmidt Conference, Sacramento, CA,
June, 2014.
Havig, J. R., McCormick, M. L., Bachan, A., and Kump, L. R. (2014) Using high resolution sampling to reveal the geochemical environment across a chemocline at
Fayetteville Green Lake, N.Y. Northeastern Geobiology Symposium, Yale University,
New Haven, CT, March 29, 2014.
Havig, J. R., Hamilton, T. L., Boyd, E. S., Meyer-Dombard, D. R., and Shock, E. L.
(2012) Geochemical and physical drivers of microbial community structure in hot spring ecosystems. Abstract, presented at 2012 Fall Meeting, AGU, San Francisco, Calif., 2-7
Dec.
Havig, J. R., Hamilton, T. L., Boyd, E. S., Meyer-Dombard, D. R., Peters, J. W., and
Shock, E. L. (2012) Effects of geochemical environmental drivers on microbial community size and structure in a hot spring ecosystem. Abstract, presented at AbSciCon
2012
Havig, J. R., Hamilton, T. L., Boyd, E. S., Meyer-Dombard, D. R., and Shock, E. L.
(2011) Effects of geochemical changes on microbial community structure in a hot spring ecosystem. Abstract B31K-01, presented at 2011 Fall Meeting, AGU, San Francisco,
Calif., 5-9 Dec.
Havig, J. R. and Shock, E. L.. (2011) The geochemical composition of hydrothermal microbial biofilms. Thermophiles. (Sept. 11-16, 2011).
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Curriculum Vitae
Jeff R. Havig
Havig, J. R., and Shock, E. L. (2010) Environmental consequences of geochemical change in hot spring ecosystems. Abstract B51A-0343, presented at 2011 Fall Meeting,
AGU, San Francisco, Calif., 13-17 Dec.
Havig, J. R., and Shock, E. L. (2009) Using hydrothermal biofilm geochemical signatures to generate predictions of elemental behavior with implications for gene hunting, biogeochemical rate measurements, and novel biosignatures. Eos. Trans. AGU 89(53):
Fall Meet. Suppl., Abstract B23D-0408.
Havig Jeff R., Jason Raymond, D’Arcy R. Meyer-Dombard, Natalya Zolotova, and
Everett L. Shock. (2009) Merging Geochemical and Metagenomic Data to Predict C and
N-fixation Pathways in Hydrothermal Microbial Communities. Geochimica et
Cosmochimica Acta 73(13): A505-A505 Suppl. S, Abstract 505.
Havig, Jeff R., Panjai Prapaipong, Natalya Zolotova, Gordon Moore, and Everett L.
Shock. (2008) Biogeochemistry of hot spring biofilms: Major and trace element behavior.
Eos. Trans. AGU 89(53): Fall Meet. Suppl., Abstract B51D-0425.
Havig, Jeff R., D'Arcy R. Meyer-Dombard, Jason Raymond, Natalya Zolotova, and
Everett L. Shock. (2008) Merging isotopic and metagenomic data to predict Carbon and
Nitrogen-fixation in hydrothermal biofilms. Geochimica et Cosmochimica Acta 72(12):
A358-A358 Suppl. 1, Abstract 358.
Havig, Jeff R., D'Arcy R. Meyer-Dombard, Jason Raymond, Natalya Zolotova, and
Everett L. Shock. (2008) Combining Isotopic and Metagenomic Data from Hydrothermal
Biofilms. NSF Research Coordination Network Geothermal Biology and
Geochemistry in Yellowstone National Park, 2008 Workshop, January 10-13, 2008,
Poster #24.
Havig, Jeff R., Everett L. Shock, and Gordon Moore. (2007) Major Element
Geochemistry of Biofilms in a Silica-Precipitating Hot Spring. Eos. Trans. AGU 88(52):
Fall Meet. Suppl., Abstract B31D-0621.
Havig, Jeff R., Everett L. Shock, Panjai Prapaipong, Gordon Moore, and Anthony
Michaud. (2006) Geochemistry of Hot Spring Biofilms. Eos. Trans. AGU 87(52): Fall
Meet. Suppl., Abstract B11A-1011.
Meyer-Dombard, D. M., A. S. Bradley, J. R. Havig, J. Raymond, J. P. Amend, E. L.
Shock, and R. E. Summons (2006) Biogeochemistry of Siliceous Biofilms in Geothermal
Ecosystems. Geochimica et Cosmochimica Acta 70(18): A418.
Havig, J. R., P. Prapaipong, and E. L. Shock. (2005) Geochemical Differences in
Hydrothermal Microbial Community Biomass. GSA Abstracts with Programs 37(7):
444.
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Curriculum Vitae
Jeff R. Havig
O'Brien, R., C. K. Keller, J. R. Havig, T. White, T. A. Coe, G. Hawley, and D. Wang
(2003) Using Field Lysimeters in Watershed Research: Results from the Hubbard Brook
Sandbox Study. GSA Abstracts with Programs 35(4): 615.
Havig, Jeff R. and C. Kent Keller. (2001) Nutrient Storage and Release Following
Perturbation of an Experimental Forest Ecosystem.
Eos. Trans. AGU 82(47): Fall Meet.
Suppl., Abstract H11C-0252.
Havig, Jeff R. and C. Kent Keller. (2000) Effect of Cessation of Photosynthetic Input on
Subsurface Nutrient Storage and Loss in an Experimental Ecosystem. Eos. Trans. AGU
81(48): Fall Meet. Suppl., Abstract H21B-04.
Research Interests:
 Geochemistry – gaining better and more complete geochemical understanding of natural systems, and exploring how it can act as a driver of environment creation and ecosystem development
 Biosignatures – searching for novel means of determining the presence of life in the past through connecting extant communities with the rock record
 Collaboration and interdisciplinary connections – exploring new ways of connecting various fields of research in order to maximize the utilization of different perspectives, skill sets, and strengths of research in order to advance science in nonconventional ways
 Biogeochemical system dynamics – the interaction of life with the environment as interconnections with geochemical processes, and the expression of this through molecular genetics and molecular systematics
 Nutrient uptake and cycling – how living systems uptake and sequester elements
 Field research and applications – understanding the best methods and means for collecting comprehensive sample sets from natural systems as well as types of contextual samples that provide the best knowledge base for interpreting and understanding primary sample targets
 Geobiology – studying geochemistry and geomicrobiology in order to better quantify elemental uptake and sequestration by microbial systems
 Biogeography – coupling geochemistry and microbiology to understand the effects of geography on microbial adaptation and evolution
 Astrobiology – exploring dynamic systems as a means for better understanding early-earth and extraterrestrial habitats
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Curriculum Vitae
Jeff R. Havig
 Planetary Geology/Geochemistry – connecting geochemistry of known terrestrial systems with those found on Mars and icy moons
Research Field Sites:
Yellowstone National Park, WY.
Research conducted at fourteen separate hydrothermal areas within the park to study the geochemistry and microbiology of hydrothermal systems across a wide range of temperature (up to boiling), pH (from less than 2 to almost 10), and element concentrations (as much as over six orders of magnitude). Sampling trips have been at least once a year since 2003, providing a powerful time-series dataset for several dozen hot springs. Chemical analyses conducted in situ on water samples for redox and temperature sensitive chemical compounds.
Samples collected for later laboratory analysis include water, biofilm, sediment, rock, and other contextual samples for determining major elements, trace elements, carbon and nitrogen content and isotopes, oxygen and hydrogen isotopes (water only), as well as molecular analysis of biofilm samples. Collaborators include researchers from Montana
State University, University of Illinois, Chicago, Massachusetts Institute of Technology,
University of Nevada, Las Vegas, University of Colorado, and MacMaster University.
Geochemistry of Fayetteville Green Lake, New York. Research conducted on a meromictic lake in coordination with collaborators from Hamilton College on the effects of distinct geochemical gradients formed between the oxygenated upper 20 m, the lower anoxic/euxinic 30 m, and the boundary layer between the two that is inhabited by a photosynthetic purple-sulfur bacterial community.
Hydrothermal Biogeochemistry of Mafic and Rhyolitic Systems, Iceland.
Research conducted in basalt and rhyolite-hosted hydrothermal systems, studying the geochemistry and molecular microbiology, including hot springs with higher temperatures and pH values than seen at Yellowstone. The goal of this work is to build a collaborative effort for a comprehensive study of hot springs in Iceland similar to the work that has been done in Yellowstone, with an eye towards looking at the similarities between rhyolitehosted systems at Yellowstone and those of Iceland, and contrasting the rhyolite-hosted and basalt-hosted systems found in Iceland. Working in conjunction with Dr. Andri
Stefánsson at the University of Iceland in Reykjavik and Dr. Everett Shock at Arizona
State University.
Subglacial Biogeochemistry of Robertson Glacier, Alberta, Canada.
Geochemical and microbiological research conducted in coordination with collaborators from Montana
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Curriculum Vitae
Jeff R. Havig
State University on the ice-liquid water-sediment/rock interface with the goal of better understanding the geochemical environment and inhabitant microbial communities of this actively changing system.
Active Serpentinization in the Semail Ophiolite Complex, Oman.
Coordinated research focusing on geochemical, microbiological, and geological sampling in conjunction with researchers from Columbia University on areas where highly alkaline
(pH of 11+) springs issue from ultramafic rocks, triggering carbonate precipitation upon contact with the atmosphere. Active serpentinization drives hydrogen production, and is connected with methane production.
Grants/Awards:
Spring, 2013 – Pennsylvania Space Grant Consortium Seed Grant Award:
Geochemical alteration and biological colonization of volcanic substrates:
Utilizing the Penn State Steam Plant; $3,400; PI
Spring, 2010 – Arizona State University Astrobiology Minigrant Award: Development of an in-lab hydrothermal artificial mesocosm; $5,172.94; PI
Special Activities:
Session Chair, AGU 2015 Fall Meeting

Session Title: Signs of Life: Exploring new frontiers in the search for biosignatures in the rock record

Co-chair: Dr. Andrew D. Czaja (UC)
Invited Talk, School for the Environment, University of Massachusetts, Boston, March
27, 2014

A tale of two hot springs: Exploring geochemical change and biological response in dynamic systems.
Shake, Rattle, and Rocks (Jan., 2013) – Outreach program through the Department of
Geosciences for 5 th
graders in all schools in the greater State College area.

Co-instructor and curriculum developer, ‘Geochemistry – Life in Extreme
Environments’
Session Chair, AGU 2012 Fall Meeting

Session Title: Integrating Geochemical and Biological Datasets to Predict the
Response of Microbial Communities to a Changing Environment

Co-chair: Dr. Eric S. Boyd (MSU)
Session Chair, AGU 2011 Fall Meeting

Session Title: Fun with Isotopes: A lighthearted look at a powerful tool.

Co-chairs: Dr. Laura Wasylenki (IU) and Dr. Lynda Williams (ASU)
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Curriculum Vitae
Jeff R. Havig
Invited Talk, Thermal Biology Institute, Montana State University, April 5, 2011

The Geochemical Composition of Hydrothermal Microbial Biofilms
Professional Associations:
Geochemical Society of America/European Geochemical Society

Member since 2008
American Geophysical Union

Member since 2000
Courses Taught/been Teaching Assistant for:
Arizona State University
Introductory Geology Lecture, Introductory Geology Lab, Geochemistry,
Geological Hazards Lab, Astrobiology, Historical Geology, Colloquium
Washington State University
Introductory Geology Lab, Organic Chemistry Lab
Professional Contacts:
Dr. Christopher H. House
Department of Geosciences
218 Deike Building
The Pennsylvania State University
University Park, PA 16802 chrishouse@psu.edu
(814) 865-8802
Dr. Lee R. Kump
Department of Geosciences
535 Deike Building
The Pennsylvania State University
University Park, PA 16802 lrk4@psu.edu
(814) 863-1274
Dr. Eric S. Boyd
Department of Chemistry and Biochemistry
224 Chemistry and Biochemistry Building
Montana State University
Bozeman, MT 59717
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eboyd@montana.edu
(406) 994-7211
Dr. Everett L. Shock
Department of Chemistry and Biochemistry
PO Box 871604
Arizona State University
Tempe, AZ 85287-1604
School of Earth and Space Exploration
PO Box 871404
Arizona State University
Tempe, AZ 85287-1404 eshock@asu.edu
(480) 965-0631
Curriculum Vitae
Jeff R. Havig
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