Glossary: 1. air space 2. chlorophyll 3. compound leaf 4. compound leaf 5. entire 6. epidermis 7. guard cell 8. lamina 9. mesophyll 10. midrib 11 palisade mesophyll 12. petiole 13 photosynthesis 14 spongy mesophyll 15 stipule 16 stoma 17. vein (vascular bundle) Leaf is an above-ground plant organ specialized for the process of photosynthesis. Leaves are typically flat (laminar) and thin which evolved as a means to maximise the surface area directly exposed to light. The internal organisation of leaves has evolved to maximise exposure of the photosynthetic organelles, the chloroplasts, to light and to increase the absorption of carbon dioxide, all of which assist photosynthesis. These adaptations are at the expense of water loss and most leaves have stomata which regulate carbon dioxide, oxygen and water vapour exchange with the atmosphere. The shape and structure of leaves varies considerably depending on climate, primarily due to the availability of light and potential for water loss due to temperature and humidity. Leaves are also the primary site, in most plants, where transpiration and guttation take place. Leaves can also store food and water, and are modified in some plants for these purposes. The concentration of photosynthesis in leaves makes them rich in protein, minerals and sugars. Because of their nutritional value leaves are prominent in the diet of many animals, including humans as leaf vegetables. Plant Parts - Leaves Leaves are the food making factories of green plants. Leaves come in many different shapes and sizes. Leaves can be simple. They are made of a single leaf blade connected by a petiole to the stem. An oak leaf or a maple leaf are examples. A compound leaf is a leaf made up of separate leaflets attached by a petiole to the stem like an ash or a locust. Leaves are made to catch light and have openings to allow water and air to come and go. The outer surface of the leaf has a waxy coating called a cuticle which protects the leaf. Veins carry water and nutrients within the leaf. Leaves are the site of the food making process called photosynthesis. In this process, carbon dioxide and water in the presence of chlorophyll (the green pigment) and light energy are changed into glucose (a sugar). This energy rich sugar is the source of food used by most plants. Photosynthesis is unique to green plants! Photosynthesis supplies food for the plant and oxygen for other forms of life. A green plant helped make the oxygen you are breathing today. Leaves and Leaf Anatomy Leaf Function: Leaves are the powerhouse of plants. In most plants, leaves are the major site of food production for the plant. Structures within a leaf convert the energy in sunlight into chemical energy that the plant can use as food. Chlorophyll is the molecule in leaves that uses the energy in sunlight to turn water (H2O) and carbon dioxide gas (CO2) into sugar and oxygen gas (O2). This process is called photosynthesis. Leaf Structure: A leaf is made of many layers that are sandwiched between two layers of tough skin cells (epidermis). The epidermis also secretes a waxy substance called the cuticle. These layers protect the leaf from insects, bacteria, and other pests. Among the epidermal cells are pairs of sausage-shaped guard cells. Each pair of guard cells forms a pore (called stoma; the plural is stomata). Gases enter and exit the leaf through the stomata. Most food production takes place in elongated cells called palisade mesophyll. Gas exchange occurs in the air spaces between the oddly-shaped cells of the spongy mesophyll. Veins support the leaf and are filled with vessels that transport food, water, and minerals to the plant. Leaf Margins: Leaves come in many sizes and shapes; they are often used to help identify plants. Some leaves are flat and wide; others are spiky and thin. Plant spines (like cactus spines) are actually modified leaves. Leaf Margins