Chapter 2 Plant Structure and Function ARBORICULTURE INTEGRATED MANAGEMENT OF LANDSCAPE TREES, SHRUBS, AND VINES 4TH EDITION Kristina Salzman Mechanical design of trees y Evenly distribute stress along surfaces y No overloaded or under loaded points over time y Active cambium growth in response to stress y Internal anatomy directly related to strength y Tree growth reveals stress pattern Crown Leaf cross-section Cross-section of a pine needle Tree Crown Excurrent vs Decurrent Sweet gum White ash Crown y Shoots y Buds { Definition { Arrangement y Shoot elongation { Spurs { Watersprouts vs suckers { Patterns of shoot elongation Periodicity Ù Apical dominance Ù Tree roots Mycorrhizal fungi y Myco = fungus; rhiza = root y More than 90% of plants in nature have a mycorrhizal symbiont y Ectomycorrhizae = fungus forms a sheath outside the root y Endomycorrhizae = no sheath; inside the root Ecotomycorrhizal root tips Tree Trunk cross-section y Xylem { { { Wood Four functions Gymnosperms (nonporous) vs angiosperms (porous) y Phloem { { { Two functions Bark Cork cambium Trunk y Taper y Movement y Unidirectional shading of tree y Reaction wood Tension wood vs compression wood { Dry vs in live state { Compartmentalization y Barriers or compartments formed around decay y CODIT Wall 1 { Wall 2 { Wall 3 { Wall 4 { Decay y Variable y Stages { Incipient { Early { Intermediate { Advanced y Two ways { White rots { Brown rots Branch attachment y Biological (transport) vs mechanical (growth) y Branch collar y Included bark Palms y Monocots y No secondary tissue y Single growing point y Two measurable heights Angiosperm flowers y Complete y Perfect y Staminate vs pistillate y Fruitfulness y Seed viability {Seed dormancy {Seed coat imperability Gymnosperm flowers y Pollen cones (smaller and lower) vs seed cones Tree death y Mortality spiral y Biological and environmental y Mechanical Photosynthesis y Using CO2, water, and light to produce O2 and sugars y Temperature y CO2—limiting factor when light is sufficient y Water y Nutrients y 6 CO2 + 6 H2O → C6H12O6 + 6 O2 Carbon dioxide + Water + Light energy → Glucose + Oxygen Photosynthesis Temperature y Range from near freezing to 104°F y High temperatures also increase respiration Carbon Dioxide Water yEssential for photosynthesis yDeficiency yExcess Nutrients y Needed to increase leaf area y Nitrogen (even at low levels) y Iron y Pesticides, air pollution, insects and diseases directly and indirectly reduce the rate of photosynthesis Sooty mold Respiration y Using photosynthates as fuel y Needs oxygen y Releases water and CO2 also y Effects of temperature y Effects of soil water y Effects of soil oxygen y Needed for compartmentalization Transpiration y Dependent upon leaf structures y Environmental effects Genetic growth mechanisms y Anatomy and structure y Pest and environmental tolerances y Life history patterns y Life span y Compartmentalization response y Production of allelopathic chemicals Genetic Blue Prints Environmental growth mechanisms y Any environmental factor y Adequate chilling y Day length Time as a growth mechanism y Juvenility and maturity y Existing structure and vigor of the tree y Maturity vs Aging Mt St Helens erupted 1980 Changes slow over time Plant Growth Regulators y Auxins y Gibberellins y Cytokinin y Abscisic acid y Ethylene Auxin y Used in plant propagation y Used in high concentrations = 2,4-D (Herbicide) Ethylene y Used in to ripen fruit and as a defoliant Root-shoot ratio y Lower on high-quality sites y Top usually 5-6 times heavier than the roots y Usually a negative situation in containers No tree part is independent of any other part. Questions?