1.2.2 Different Cell Types [A] Plant Tissues (a) Parenchyma : made up of parenchyma cells unspecialised cell with thin cell wall, possess only primary cell wall Tissue Living / dead Cell shape Main function Distribution Parenchyma Living Usually isodiametric , sometimes elongated 1. As ground tissue and packing tissue of plant body 2. Important in the maintenance of turgidity of the cell especially in herbaceous plant 3. Sites for metabolic activities, storage, deposition of waste products 4. Site of meristematic function 5. Large intercellular air spaces among cell for gaseous exchange 6. may be photosynthetic Cortex, pith, medullary rays and packing tissue in xylem and phloem C:\BIO\AL\NOTES\01CELL\1.2.2 cell types_PC(own) 1 Modified parenchyma: 1. Epidermis - Usually surface covered with cuticle made of cutin - Guard cells present in leaves or young stem, with unequal cell wall thickening - At the root tip, root hair may extend from these cells. How the structures of the epidermis are adapted to its functions Structure Functions it is a continuous, compact layer of cells Give mechanical support and protection against certain fungi on the surface of the plant body and bacteria it is one cell thick and transparent Allow light to penetrate into the mesophyll for photosynthesis epidermal cells of the stem and leaves Reduce water loss through transpiration from the leaf surface have a waxy cuticle on their outer walls Guard cells surrounding stomata Allow gaseous exchange between the leaf & atmosphere Guard cells with thicker inner wall and For the formation of stomata by the uneven stretching of the thinner outer wall cell wall as the cell becomes turgid. It can control the opening and closing of the stomata and thus control the gas exchange and water loss. Guards cells contain chloroplasts It can regulate the water potential of the guard cells and thus which allow the photosynthesis to carry control the stomatal opening. out. C:\BIO\AL\NOTES\01CELL\1.2.2 cell types_PC(own) 2 2. mesophyll - in green leaves, with abundant chloroplasts for photosynthesis Shape Amount of chloroplasts Intercellular air spaces Palisade Mesophyll Spongy Mesophyll Regular, columnar / cylindrical Irregular Abundant few Negligible and closely packed Large and very loosely arranged 3. endodermis - the innermost one-celled layer of cortex surrounding vascular tissue - with Casparian strips (made of suberin, water-proof fatty substance, surrounding the transverse & radial wall of the cell ) - At certain intervals, with passage cells (with NO Casparian strips) 4. pericycle - occurs only in root, one to several layers outside the central vascular tissue - gives rise to lateral root and takes part in secondary growth C:\BIO\AL\NOTES\01CELL\1.2.2 cell types_PC(own) 3 5. - (b) companion cell - found adjacent to sieve tube With dense cytoplasm & prominent nucleus, metabolically active - may be used to supply energy for active movement of substances into the companion cells and help the transport in sieve tube Collenchyma : Tissue made up of collenchyma cells Living /dead Collenchyma Living Cell shape Main function Distribution Elongated and polygonal Support (a mechanical Outer region of cortex e.g. angle of stems, function) with tapering ends midrib of leaves (Cell wall thickened at the corners) - cytoplasm is scarce, small lumen and small nucleus support is given entirely by the cell wall - its cell wall is usually thickened at the corners with deposition of extra cellulose, hemicellulose and pectin. - The cell walls are generally flexible and allow certain degree of stretching. - intercellular space absent or extremely scarce. C:\BIO\AL\NOTES\01CELL\1.2.2 cell types_PC(own) 4 (c) Sclerenchyma : Tissue Sclerenchyma made up of fibres and sclereids Living / dead Cell shape Dead 1. Fibres Elongated & polygonal with tapering interlocking ends 2. Sclereids Roughly isodiametric C:\BIO\AL\NOTES\01CELL\1.2.2 cell types_PC(own) Main function Support & mechanical protection Distribution Outer region of cortex, pericycle of stems, xylem & phloem Cortex, pith, phloem, fruits & seeds 5 - - Support is provided by the cell wall which is evenly thickened. This extra-thickening is mainly due to the deposition of lignin : a hard substance with great tensile (i.e. not easily break on stretching) and compressional strength. Mature cells are dead with empty lumen at the centre. Function : cannot carry out photosynthesis, not for food storage, solely for support 2 kinds: 1. fibres - elongated, with interlocking tapering ends - closely packed with little or no air-space - polygonal and closely packed leaving little or no air spaces between them. - usually found in the outline of the cortex of stem and root and frequently associated with vascular tissues. e.g. sclerenchyma sheath around the vascular bundles 2. (d) Xylem : sclereids or stone cells - short and spherical in shape - found in cortex, phloem and medulla of stem and in flesh of some fruits such as pear. made up of xylem tracheids and xylem vessels C:\BIO\AL\NOTES\01CELL\1.2.2 cell types_PC(own) 6 - 1. Tissue Living /dead Cell shape Main function Distribution Tracheids & vessels dead when mature Elongated & tubular Transport of water & mineral salts; Support Vascular system Xylem fibres and parenchyma (similar to that of sclerenchyma fibres and parenchyma) Xylem consists of non-living cells with supporting function includes tracheids and vessels. - Walls of tracheids and vessels are thickened with cellulose and lignin - they offer resistance to collapse especially during transpiration. Xylem tracheid 2. Elongated with fine tapering ends (L.S.), polygonal (X.S.) Wall is heavily lignified & pitted (secondary wall) when mature They have no perforation plates but they have pits connecting the tracheid to the adjacent one. No protoplast (when mature) Xylem vessel - more important in angiosperms for water transport and support. Structural adaptation of vessel : - made up of dead hollow cells (xylem vessel elements) joined end to end with no end wall minimum resistance - thick lignified wall prevent wall collapsing and give mechanical support C:\BIO\AL\NOTES\01CELL\1.2.2 cell types_PC(own) 7 - Secondary wall shows various patterns of thickening due to the deposition of lignin. Bordered pits in the longitudinal wall with adjacent cell perforated end wall Vessel has greater diameter and large lumen than tracheid C:\BIO\AL\NOTES\01CELL\1.2.2 cell types_PC(own) 8 Relationship between the featuresand functions of xylem vessels (e) Features Functions All the transverse walls are absent, the vessels effectively become continuous tubes. Unlignified area (pits) occur in the side wall of pitted vessels The wall is thickened with lignified cellulose To enable the continuous conduction of water and mineral salts through the plant. To facilitate the lateral movement of water since the lignified wall themselves are impervious. For mechanical support – the strength of their lignified walls enable xylem vessels to resist compression and tension : Phloem : made up of sieve (tube) elements and companion cells - Cell wall mainly made of cellulose. Tissue Living /dead Cell shape Main function Sieve tubes Living Elongated & tubular Translocation of organic solutes (food) Companion cells Living Elongated & narrow Work in association with sieve tubes Phloem fibres and parenchyma (similar to that of fibres and parenchyma described before) Distribution Vascular system 1. Sieve (tube) elements - During development from meristematic cell, nucleus degenerates. sieve element is a living cell with no nucleus. - Sieve elements are joined end to end forming a sieve tube, each element being separated from the next by a sieve plate. C:\BIO\AL\NOTES\01CELL\1.2.2 cell types_PC(own) 9 - When plasmodesmata of end walls enlarge greatly, sieve pores are formed on the sieve plate. Sieve tubes are tube-like structure with a wide lumen and a very narrow, indistinct, peripheral layer of living cytoplasm bounded by a plasma membrane. Sieve plates are regions of thick primary cell wall, with protoplasmic strands passing through. - Sieve areas are also present in the side wall for the lateral translocation of organic solutes to the neighbouring cells. - sieve tubes are living. contain obstructions (sieve plate and cytoplasm) to the flow of solution. C:\BIO\AL\NOTES\01CELL\1.2.2 cell types_PC(own) 10 2. Companion cells - are parenchyma cells derived from same parent cell as the neighbouring sieve element and closely associated with sieve element. - with prominent nucleus, dense cytoplasm, small vacuole, numerous mitochondria and ribosomes companion cells are very active metabolically. found in angiosperm only C:\BIO\AL\NOTES\01CELL\1.2.2 cell types_PC(own) 11 How the structural features of phloem related with their functions Features The sieve plate of sieve tubes with protoplasmic strands passing through Companion cell - has pitted wall for communication with the cytoplasm of the sieve tube by plasmodesmata - with dense cytoplasm, nucleus & is metabolic active. C:\BIO\AL\NOTES\01CELL\1.2.2 cell types_PC(own) Functions Enable the transport of solute through the tube. - It is functionally associated with sieve (tube) elements may be used to supply energy for active movement of substances into the companion cells and help the transport in sieve tube 12 [B] Animal Tissues (a) Epithelium : made up of epithelial cells - in single (simple epithelium) or multilayered sheets (compound epithelium) - covers the internal and external surfaces of the body of an organism for protection. - The bottom layer of cells rests on a basement membrane. - The free surface of the epithelium is often highly differentiated for various purposes: squamous epithelium lining of capillaries (endothelium) Alveoli in lungs Bowman’s capsule of kidney ciliated epithelium lining the oviducts, ventricles of brain, spinal cord, respiratory passages stratified epithelium skin oesophagus 1. squamous epithelium : with smooth lining, to provide a relatively friction-free passage of fluids through hollow structures such as blood vessels, heart chambers (endothelium) and the inside of the body cavity (peritoneum). The thinness of squamous epithelium facilitates diffusion across it. The smoothness facilitates the passage of fluid and lubricates the movement between adjacent surface. 2. Epithelium bears numerous cilia and always associated with mucus-secretory goblet cells (ciliated epithelium lining the oviducts, ventricles of brain, spinal cord, respiratory passages) serve to move materials along the passage. C:\BIO\AL\NOTES\01CELL\1.2.2 cell types_PC(own) 13 3. stratified epithelium : consists of several layers of cells making up a tough, thick and impervious lining of skin, oesophagus, vagina etc. protect underlying structures from injury through abrasion or pressure, and from infection. The cells of the outermost layers are dead and are continuously replaced from below. The innermost layer of cells, called generative layer, is in an active state of mitosis. Above the generative layer, the cells become progressively isolated from the oxygen and nutrients supplied by the blood. The cells die and eventually slough off. C:\BIO\AL\NOTES\01CELL\1.2.2 cell types_PC(own) 14 Special feature of stratified epithelium related to the function • The multilayered structure provides protection from abrasion. • e.g. A stratified epithelium occurs in the buccal cavity and in the vagina • e.g. in the skin (epidermis) additional protection is afforded by the dying cells, the proteins of which become modified into a hard waterproof layer. The protein of this layer (keratin) provides a tough surface referred to as “cornified”•The greater the friction that a given area of skin experiences, the thicker the cornified layer becomes. (b) Blood Cells 1. Red Blood Cell (Erythrocytes) Shape & structure biconcave discs Have a larger SA/V ratio for gaseous exchange. Thin permit efficient diffusion of gases from its surface inwards. Non-nucleated in human RBC Passive and no activity; only acting as a carrier; no nucleus to occupy any space or volume in cell. Permit more Hb to be packed in the cell to carry more O 2 (Whole interior filled with Hb : surrounded by a thin membrane. No nucleus (in human RBC) and cytoplasm containing haemoglobin (takes about 90% of the dry mass) – enable the RBC to carry O2 as oxyhaemoglobin and CO2 as carbamino haemoglobin formed in bone marrow Function : as O2 and CO2 (some) carriers 2. White Blood Cell (Leucocytes) Shape & structure All are nucleated very active. (may be) amoeboid shape formed in bone marrow & lymphatic system fewer number than RBC (1 WBC : 500 RBC) Function : for body defence. Types : 1. Granulocytes : granular cytoplasm and lobed nucleus (polymorphonuclear) - capable of amoeboid movement. - Types: (1) Neutrophils (phagocytes) - able to squeeze between the cells of capillary wall and enter intercellular spaces. - They can be stained by neutral dye. - Function : actively phagocytic : engulf and digest disease-causing bacteria in the intercellular spaces. (2) Eosinophils - Stained by eosin (an acid dye) - Function : have anti-histamine properties. - It increases in number with allergic conditions such as asthma, hayfever, parasitism etc. (3) Basophils - rare in blood - Granules stained by basic dye e.g. methylene blue - Polymorphic nuclei; - Function : produce heparin and histamine - They can leave the blood and form‘mast cells’. Mast cells cause blood vessel dilation at sites of inflammation such as those caused by insect bites. 2. Agranulocytes : non-granular cytoplasm; oval or bean-shaped nucleus - Types: (1) Monocytes C:\BIO\AL\NOTES\01CELL\1.2.2 cell types_PC(own) 15 - horseshoe-shaped / bean-shaped nucleus Function : actively phagocytic and ingest bacteria and other large particles. They are able to migrate from the bloodstream to inflamed area of the body and carry out phagocytosis (in the same manner as neutrophils) (2) Lymphocytes rounded nucleus, only a small quantity of cytoplasm 2 types: T cell, B cell Functions: To cause immune reactions e.g. T cell : cell-mediated response - phagocytic B cell : produce antibodies acting against antigens 3. Platelets (Thrombocytes) Shape & structure - Irregularly shaped membrane-bound cell fragments. - They are irregularly shaped membrane-bound cell fragments produced by large cells in the bone marrow. Frequently (Number is between 150 000 and 400 000 platelets in each cubic millimeter of blood.with no nucleus.) Function : for blood clotting. C:\BIO\AL\NOTES\01CELL\1.2.2 cell types_PC(own) 16 (c) Nervous tissue (Neurone) Structure 1. 2. 3. - Basic unit is neurone (about 1010 in the brain) All neurones have the following structures: Cell body containing a nucleus. Dendron – a fine cytoplasmic fibre which brings impulses in towards the cell body. Axon – a single fibre taking impulse away from the cell body. many fine cytoplasmic fibres connecting to the cell body are called dendrites) which transmit nerve impulses to the cell body. Impulses leave via the axon. Features The cytoplasm of a neurone cell body is densely packed with ribosomes, Golgi apparatus, rough endoplasmic recticulum and mitochondria Microtubules run from the cell body cut along the axons and tendons Functions Ribosomes, Golgi apparatus, rough ER are responsible for synthesizing neurotransmitters. Mitochnondria are responsible for the production of energy for the process to take place. Function of a neurone - responsible for initiating an electrical impulse and coordinating the activities of other tissues of the body. Neurone Receptors (receive stimuli, e.g. sensory cells) Effectors (response, e.g. muscle or glands) Some axons are covered by a fatty myelin sheath formed by Schwann cells. Different type of nerve fibres : (a) Myelinated nerve fibres Fibres surrounded by fatty myelin sheath. A tough inelastic membrane, neurilemma surround the myelin sheath. Myelin sheath is absent at nodes of Ranvier leaving the fibre unprotected. e.g. In cranial nerves and spinal nerves C:\BIO\AL\NOTES\01CELL\1.2.2 cell types_PC(own) 17 Functions of myelin sheath : for protection act as an electrically insulating sheath help in the increase in the speed of impulse transmission. (b) Non-myelinated nerve fibre no myelin sheath around nerve fibres Functional classification of neurones - Based on the direction in which they transmit impulse and on the positions they occupy in the nervous system. - Different types of neurone : (a) Afferent (sensory) neurone It conducts impulses along a dendron from a receptor organ to the central nervous system. Cell bodies of sensory neurones occur in clusters called ganglia next to the spinal cord. The axon carries the impulse into the central nervous tissues. (b) Efferent (motor) neurone They transmit impulses along axon from the central nervous system to the muscles and gland. (c) Intermediate (relay, association) neurone They occur within the central nervous system (brain or spinal cord) or in sympathetic ganaglia. They receive impulses from sensory neurones from other intermediary neurone to motor neurones or to other intermediary neurones. C:\BIO\AL\NOTES\01CELL\1.2.2 cell types_PC(own) 18 Nerve : a bundle of nerve fibres. C:\BIO\AL\NOTES\01CELL\1.2.2 cell types_PC(own) 19