RICHARD P. READING
PERSONAL
Home Address: 1452 Hudson St., Denver, CO 80220
Work Address: Denver Zoological Foundation, 2300 Steele Street, Denver, CO 80205
E-Mail: rreading@denverzoo.org; rpreading@gmail.com; rich_reading@hotmail.com
Phone/Fax: 720-337-1545 (W); 303-653-8436 (C); 303-376-4806 (Fax)
EDUCATION
Yale Graduate School, New Haven, CT. M.S., M.Phil., & Ph.D. in Wildlife Ecology.
Dissertation: Toward an Endangered Species Reintroduction Paradigm: A Case Study of the Black-footed Ferret.
Yale School of Forestry and Environmental Studies, New Haven, CT. Master of Environmental Studies.
Master's Project: Conservation & Management of the Endangered Species of the Ryukyu Islands, Japan.
Trinity College, Hartford, CT. B.S. Biology.
Duke University Marine Laboratory, Beaufort, NC. Research & Courses in Marine Biology.
EXPERIENCE
2015 – Present Consultant in Conservation Biology.
Providing consultation services for a wide variety of private and government organizations on a broad range of
conservation topics, including field research into biodiversity conservation, protected areas management, animal
translocations, establishing high performance teams to address conservation issues, developing conservation
policies, developing conservation education and outreach programs, conducting data analyses, and more. Current
(summer, 2015) clients include the Southern Plains Land Trust (assisting with developing and implementing a
bison translocation and conservation program) and the Butterfly Pavilion (assisting with establishing a
conservation biology department and conservation programming).
1996 - 2015
Vice President for Conservation. Denver Zoological Foundation.
Founder and Vice President of the Department of Conservation Biology (now the Department of Conservation
and Research). The Department is responsible for the conservation and research missions of the zoo and
therefore develops, conducts, and facilitates research and conservation activities, including dozens of projects
within the zoo, in the West, and throughout the world. Position includes setting overall direction for the
conservation initiatives of the organization; overseeing and conducting research projects in the zoo, the U.S., and
overseas; initiating and implementing conservation projects and programs; developing, overseeing, and fund
raising for the departmental budget; teaching at area universities, and overseeing graduate students at a number
of universities in the U.S. and in countries in which we work; post-doctoral position, and an international staff),
an unpaid staff of several Research Associates, and the conservation and research work of staff from other
Denver Zoo departments; assisting the zoo with conservation message development and integrating education
and conservation initiatives; serving on the boards of other non-profit, conservation organizations; serving as the
conservation and research liaison with the Board of Trustees; and forging cooperative relationships with other
organizations in the region and throughout the world.
1997 - Present Adjunct Professor. Biology Department, University of Denver.
Conduct research, advise graduate and undergraduate students, and teach classes in Conservation Biology,
Wildlife Biology, Field Research Methods, Population Ecology, and various graduate seminars.
2012 - Present Associate Fellow. Center for Great Plains Studies. University of Nebraska-Lincoln.
Collaborate on research and conservation activities in the Great Plains.
1988 - Present Research Associate. Northern Rockies Conservation Cooperative.
Conduct research in a several disciplines, including ecology of black-tailed prairie dog colonies; social, political,
organizational, policy, and biological aspects of black-footed ferret reintroduction; carnivore conservation
policy; attitudes toward ecosystem management; and large felid biology and conservation in Mexico.
R. P. Reading, Ph.D. — Page 2
2002 - Present Research Associate. Denver Museum of Nature and Science.
Conduct research and coordinate conservation efforts between the Denver Zoo and the Museum.
2008 - 2012
Adjoint Senior Research Professor. Biology Department. University of Colorado-Denver.
Conduct research, advise graduate and undergraduate students, and teach classes in Conservation Biology and
various graduate seminars.
1994 - 2011
Research Associate. Nature Conservation International (Berlin, Germany).
Conduct research on wildlife and national parks in Mongolia, including population monitoring of argali, ibex,
khulan, wild camels, Mongolian gazelle, and goitered gazelle, assessments of Mongolian national park and
conservation initiatives, and analysis of hunting and commercial harvests.
1999 - 2010
Research Associate. the Wilds (Cumberland, Ohio).
Conduct research in collaboration staff from the Wilds on wildlife and national parks in Mongolia, especially
wild Bactrian camels.
1999 - 2002; 2005 – 2009
Affiliate Faculty. Animal Sciences Department, Colorado State University.
Conduct research and advise students in animal physiology, wildlife ecology, nutrition, and related disciplines.
2001 - 2006
Faculty Affiliate. School of Forestry, University of Montana.
Conduct research and advise student in wildlife ecology.
1998 – 2000
Affiliate Faculty. Biology Department, Colorado State University.
Conduct research and advise students in conservation biology, wildlife ecology, and related disciplines.
1995 – 1997
Consultant. German Technical Advisory Group (GTZ).
Led teams of international and national experts in biological assessments of Three Beauties of the Gobi National
Conservation Park, Mongolia. Conducted social assessment of local people’s knowledge, values, and attitudes
with respect to wildlife, the park, and park management and provided management recommendations.
1994 - 1995
Consultant. UNDP/Global Environ. Fund & Mongolia Ministry for Nature & the Environment.
Co-Leader, with a Mongolian counterpart, of the Species and Park Management component of the Mongolian
Biodiversity Project. Assessed the adequacy of species management and protected areas programs; developed
training programs; held conferences; advised officials, and worked to improve research and management.
1991 - 1994
Consultant. Victoria Department of Conservation and Natural Resources.
Critically reviewed and provided recommendations on eastern barred bandicoot (a small Australian marsupial)
conservation efforts in Victoria, Australia. Helped improve population monitoring, ecological research, captive
propagation and reintroduction, program structuring, public support, and internal cooperation of recovery
actions.
1991 - 1994
Wildlife Management Biologist. Bureau of Land Management.
Threatened and endangered species specialist focusing on black-footed ferret conservation. Conducted and
supervised research on black-tailed prairie dogs and associated specie; engaged in community outreach; and
worked with other organizations to promote ferret recovery.
1988 - 1992
Teaching Assistant. Yale University. (School Year)
Assisted teaching Species & Ecosystem Conservation: Developing & Applying a Policy Orientation; Wildlife
Ecology; Natural Resource Science & Politics; and Science & Politics of Environmental Regulation.
1986 - 1988
Menagerie Keeper. New York Zoological Society.
Keeper throughout the Mammal Department of the Bronx Zoo. Performed independent research on captive pudu
deer ingestion and nutrition.
1985 (Summer) Independent Researcher. Trinity College Department of Biology.
R. P. Reading, Ph.D. — Page 3
Taxonomic laboratory research on Rhodophyta (red algae), including description of a new species and several
new records from North Carolina.
STUDENTS
Main Advisor or Field Supervisor
Tuya, Tserenbataa. M.S. Biology. Department of Biology, University of Denver. March 2003.
Population genetic study of argali sheep (Ovis ammon) in Mongolia: Implications for conservation.
Murdoch, Jed. M.S. Biology. Department of Biology, University of Denver. March 2004.
Scent marking behavior of the San Joaquin kit fox (Vulpes macrotis mutica).
Kristin Salamack. M.S. Biology. Department of Biology, University of Denver. March 2008
The influence of a highly interactive species on bird species in urban habitat fragments.
Murdoch, James. Ph.D. Zoology. Department of Zoology, Oxford University. April 2009.
The ecology and conservation of small carnivores in Mongolia. (Field Supervisor).
Glyn Maude. Ph.D. Candidate. University of Bristol, Bristol, England, U.K. August 2010.
Ecology and conservation of brown hyaenas in Botswana. (Field Supervisor)
Tshimologo, Botilo. M.S. Candidate. University of Bostwana, Gabarone. Oct 2014.
A comparison of African wild dog ecology in the Central Kalahari and Okavando Delta. (Field Supervisor).
Jadamsuren Adiya. Ph.D. Candidate. Mongolian National University, Ulaanbaatar, Mongolia. Planned completion:
December 2016. Ecology and conservation of wild Bactrian Camels. (co-Advisor).
Leo, Viyanna. Ph.D. Candidate. University of New South Wales, Sydney, Australia. Planned completion: June 2016.
The role of dingos as top down regulations. (Field Supervisor)
Suuri, Buyandelger. Ph.D. Student. National University of Mongolia. Planned Completion: June 2016.
Marmots as Keystone Species in Mongolia. (co-Supervisor).
Sodnompil, Batdorj. Ph.D. Student. National University of Mongolia. Planned Completion: June 2016.
Hedgehog Resource Partitioning in Mongolia. (co-Supervisor).
Committee Member
Molitoris, Sandra. Honors B.S. in Animal Sciences, Colorado State University. May 1998.
Palatability of a Gel Fish Substitute for Marine Mammals: Harbor Seals (Phoca vitulina) and California Sea
Lions (Zalophus californianus).
Bestelmeyer, Stephanie. Ph.D. in Biology, Colorado State University. June 2000.
Solitary, Reproductive, and Parental Behavior in Captive and Free-Ranging Maned Wolves.
Zapico, Tom. M.S. in Biology, University of Denver. December 2000.
Effects of environment on the behavior of captive common hippos (Hippopotamus amphibius) and observations
of sociosexual investigatory behaviors.
Rosmarino, Nicole. Ph.D. in Political Science, University of Colorado. May 2002.
Endangered Species Act Under Fire: ESA Controversies, Science, Values, and the Law.
Dolan, Bridgette. M.S. in Geography, University of Denver. June 2002.
Prairie Conservation and GIS: Mapping Change and Predicting Prairie Dog Habitat Preference for the Southern
Plains Land Trust in Baca County, CO.
Renee, Tatja. M.S. in Biology, University of Denver. June 2002.
Allometric Analysis of Voluntary Dive Times in Fresh Water Turtles.
Bernal Stoopen, Jose F. Ph.D. in Wildlife & Fisheries Science, Texas A&M University. May 2004.
Binational Collaboration in recovery of endangered species: The Mexican wolf as a case study.
Wingard, Ganchimeg. M.S. in Wildlife Biology, University of Montana. December 2005.
Seasonal food habits of argali and dietary overlap with domestic livestock in Mongolia
Magle, Seth. Ph.D. in Wildlife Ecology, Colorado State University. June 2007.
Ecology and dynamics of an urban complex of prairie dog colonies in the Denver metro area.
Insalaco, Emily. M.A. George Mason University. May 2010.
Evaluating enrichment devices with Denver Zoo orangutans with the goal of identifying appropriate options for
orangutans in Borneo.
Bragin, Nanette. M.A.S. University of Denver. May 2010.
Creating a habitat suitability index model for argali sheep (Ovis ammon), and Siberian ibex (Capra sibirica) in
Ikh Nart Nature Reserve, Mongolia.
Zapletal, Mirka. M.S. Antioch University. May 2012.
R. P. Reading, Ph.D. — Page 4
Habitat utilization by Daurian hedgehogs (Mesechinus dauuricus) in the Gobi Steppe Environment.
SCHOLARSHIPS / HONORS
1980
1980-1984
1980-1984
1983
1983
1982-1984
1984
1984
1988-1993
1997
1997
2001
2003
2003
2004
2004
2007
2007
2009
2012
2014
2014
2014
2015
Regent’s Scholarship. New York State Board of Regents.
Alfred E. Ives Scholarship. Alfred E. Ives Foundation.
Trinity Scholarship. Trinity College.
Phi Beta Kappa. Trinity College.
John Small Award. (Male scholar-athlete award) Trinity College.
Faculty Honors List. Trinity College.
Graduated 10th in class with Honors. Trinity College.
Bookhout Scholarship. Duke University.
Yale University Fellowship. Yale University Graduate School.
Biographee. Who’s Who in Science and Engineering.
Certificate of Appreciation. Ministry of Natural Resources and Environment, Victoria, Australia.
Certificate of Appreciation. Colorado Division of Wildlife.
E-chievement Award. E-Town Radio and Whole Food.
Certificate of Appreciation. Southern Plains Land Trust, Pritchett, Colorado.
Biographee. Who’s Who in Sciences Higher Education
Honorary Membership. Mongolian Association of Colorado.
Significant Achievement in International Conservation. Association of Zoos and Aquariums.
Distinguished Environmentalist. Highest honor of the Mongolian Ministry for Nature and Environment
Recognition Award for Conservation Research. Dornogobi Province, Mongolia.
Best Monitoring & Evaluation Award - Lake Titicaca Frog (Telmatobius culeus) Conservation Project,
Peru. Eco-Index, Rainforest Alliance.
Recognition Award for Contributions to Conservation. Dalanjargal Soum (County), Mongolia.
Kublai Khan Gold Medal. Highest honor of the Mongolian Academy of Sciences.
Distinguished Scientist. Highest honor of the Mongolian Ministry of Education and Science.
Honorary Doctorate, Mongolian Education University.
BOARDS OF DIRECTORS
2000-2004
2001-2004
2001-2003,
2005-2006
2003-2008
1999-Present
2001-Present
2007-Present
2011-Present
2013-Present
2015-Present
Board of Trustees. The Colorado-Mongolian Project: Cultural and Education Exchanges, Denver, CO.
Board of Directors. The Predator Conservation Alliance, Bozeman, MT. (Treasurer 2002-2003, President
2004)
Board of Directors. Southern Plains Land Trust, Pritchett, CO. (Newsletter Editor 2004- Present)
Board of Directors. Denver-Ulaanbaatar Sister Cities Committee, Denver, CO.
Board of Directors. Argali Wildlife Research Center, Ulaanbaatar, Mongolia.
Board of Directors. The Mongolian Conservation Coalition, Ulaanbaatar, Mongolia (President 2004Present)
Board of Directors. Idea Wild, Fort Collins, CO.
Board of Directors. Sand Creek Regional Greenway Partnership, Denver, CO.
Steering Committee, Ivory Free Colorado, Denver, CO.
Board of Directors. African Network for Animal Welfare-US, Denver, CO.
ADVISORY BOARDS & JOURNAL EDITOR
1998-2001
1999-2000
2000-2003
2004-2012
2012-2014
2000-Present
2010-Present
2013-Present
Member (Chair 1998-2000). Lynx-Wolverine Advisory Team to the Colorado Division of Wildlife.
Scientific Advisor. Southern Plains Lands Trust, Pritchett, CO.
Member. AZA Special Task Force on Science and Technology
Board of Advisors. Grasslands Foundation, Lincoln, NE.
Board of Advisors. Lowry Range Stewardship Advisory Committee, Denver, CO
Scientific Advisor. Rocky Mountain Wild (formerly Center for Native Ecosystems), Denver, CO.
Scientific Advisor. Prairie Dog Coalition of the Human Society US, Boulder, CO.
Assistant Editor. Journal of Asia-Pacific Biodiversity.
R. P. Reading, Ph.D. — Page 5
2014-Present
2014-Present
Board of Scientific Advisors. Butterfly Pavilion, Westminister, CO.
Review Editor. Frontiers in Ecology and Evolution & Frontiers in Environmental Science
MEMBERSHIPS
1988 - 1989
1991-1992
Present
President. Yale Chapter of The Wildlife Society, Yale University, New Haven, CT.
Senator. Yale University Graduate School Senate, New Haven, CT.
Professional Membership. Bison, Reintroduction, Captive Breeding, & Caprinae Groups of the IUCN's
Species Survival Commission, Society for Conservation Biology, Flora and Fauna International, the
Oriental Bird Club, Asian Raptor Research and Conservation Network, The Wildlife Society, American
Association for the Advancement of Science, American Institute of Biological Sciences, Natural Areas
Association, and Large Herbivore Initiative.
SKILLS
Grants. Received support from the American Association of Zoo Keepers – Rocky Mountain Chapter, Anza-Borrego
Foundation, Aquarium and Zoo Facility Association, Chicago Zoological Society, Cleveland Metroparks Zoo,
Conservation Endowment Fund of the AZA, David Traylor Zoo, Disney Worldwide Conservation Fund, Earthwatch
Institute, Edna Sussman Fund, ESRI, Folsom Children’ Zoo and Botanical Gardens, Gerald R. Dodge Foundation,
the German Technical Advisory Group (GTZ), Global Environment Facility, Montana Department of Fish, Wildlife,
& Parks, International Iguana Foundation, Minnesota Zoo’s Ulysses S. Seal Conservation Grant Program, Mohamed
bin Zayed Species Conservation Fund, Mongolian Open Forum, National Fish & Wildlife Foundation, Nature
Conservation International, Northern Rockies Conservation Cooperative, Partners Without Borders, Thaw Charitable
Trust, Turner Endangered Species Fund, Trust for Mutual Understanding, United Nations Development Programme,
U.S. Bureau of Land Management, U.S. Embassy to Mongolia’s Ambassador’s Fund, U.S. Fish & Wildlife Service,
Victorian Department of Natural Resources, Wildlife Reserves Singapore, World Bank-Netherlands Trust Fund for
Environmental Reform, World Society for the Protection of Animals, WWF - International, World Wildlife Fund U.S., and a variety of private sources.
Art. Published biological illustrations and photographs.
Equipment. Experience operating four wheel drive vehicles (including double-clutch trucks), motorcycles, all-terrain
vehicles, small outboard boats, snow mobiles, radio telemetry equipment, winches, electronic planimeters, global
positioning systems, several computer operating systems and programs, and more.
First Aid & CPR Certification. American Red Cross.
Certified Open Water Diver. (P.A.D.I.)
Scientific Referee. Review manuscripts for their scientific rigor and merit for a variety of scientific publications, including
books and journal articles. Journals include Ambio, American Midland Naturalist, Animal Conservation, Biological
Conservation, Conservation Biology, Ecology, Ecological Applications, Great Plains Research, Human Dimensions
of Wildlife, Integrated Zoology, Journal of Applied Ecology, Journal of Arid Ecology, Journal of Mammalogy
Journal of Asia-Pacific Biodiversity, Journal of Wildlife Management, Mammalian Species, Mongolian Journal of
Biological Sciences, Southwestern Naturalist, Oryx, Ursus, Wildlife Biology, Wildlife Society Bulletin, and Zoo
Biology.
TECHNICAL PUBLICATIONS
1986
1. Reading, R. P. and C. W. Schneider. 1986. On the male conceptacles of two terete species of Gracilaria (Rhodophyta,
Gigartinales) from North Carolina. Journal of Phycology 22:395-398.
1987
2. Schneider, C. W. and R. P. Reading. 1987. A revision of the genus Peyssonnelia (Rhodophyta, Crytonemiales) from
North Carolina, including P. atlantica sp. nov. Bulletin of Marine Science 40:175-192.
1989
3. Reading, R. P., J. J. Grensten, S. R. Beissinger, and T. W. Clark. 1989. Attributes of black-tailed prairie dog colonies
in Phillips County, MT, with management recommendations for the conservation of biodiversity. Montana Bureau
of Land Management Wildlife Technical Bulletin 2:13-27.
R. P. Reading, Ph.D. — Page 6
1990
4. Miller, B., C. Wemmer, D. Biggins and R. P. Reading. 1990. A proposal to conserve black-footed ferrets and the
prairie dog ecosystem. Environmental Management 14:763-769.
5. Reading, R. P. and T. W. Clark. 1990. Black-footed ferret annotated bibliography, 1986-1990. Montana Bureau of
Land Management Wildlife Technical Bulletin 3:1-22.
1991
6. Reading, R. P., T. W. Clark , S. R. Kellert. 1991. Toward an endangered species reintroduction paradigm. Endangered
Species Update 8(11):1-4.
7. Reading, R. P., T. W. Clark, P. Goldstraw, A. J. Watson, and J. H. Seebeck. 1991. An overview of eastern barred
bandicoot reintroduction programs in Victoria, Australia: With recommendations for future reintroductions. Victoria
Department of Conservation and the Environment. Melbourne, Victoria, Australia. 21 pp.
1992
8. Clark, T. W., P. Schuyler, T. Donnay, P. Curlee, T. Sullivan, M. Cymerys, L. Sheeline, R. Reading, R. Wallace, T.
Kennedy, Jr., A. Marcer-Battle, and Y. DeFretes. 1992. Conserving biodiversity in the real world: Professional
practice using a policy orientation. Endangered Species Update 9(5/6):5-8.
9. Reading, R. P., T. W. Clark, A. Arnold, J. Fisher, P. Myroniuk, and J. H. Seebeck. 1992. Analysis of a threatened
species restoration program: The eastern barred bandicoot (Perameles gunnii) in Victoria, Australia. Victoria
Department of Conservation and the Environment. Melbourne, Victoria, Australia. 26 pp.
1993
10. Reading, R. P. and S. R. Kellert. 1993. Attitudes toward a proposed black-footed ferret (Mustela nigripes)
reintroduction. Conservation Biology 7:569-580.
1994
11. Backhouse, G. N., T. W. Clark, and R. P. Reading. 1994. The Australian eastern barred bandicoot recovery program:
Evaluation and reorganization. Pp. 251-271, in: Endangered Species Recovery: Finding the Lessons and Improving
the Process. T. W. Clark, R. P. Reading, and A. L. Clarke (eds.). Island Press.
12. Backhouse, G. N., T. W. Clark, and R. P. Reading. 1994. Reintroductions for recovery of the eastern barred
bandicoot Perameles gunii in Victoria, Australia. Pp. 209-218, in: Reintroduction Biology of Australian and New
Zealand Fauna, M. Serena (ed.). Surrey Beatty and Sons, Chipping Norton, Victoria.
13. Clark, T. W., and R. P. Reading. 1994. A professional performance perspective: Professional roles, problem solving,
sharing knowledge across expertise, and learning. Pp. 351-369, in: Endangered Species Recovery: Finding the
Lessons and Improving the Process. T. W. Clark, R. P. Reading, and A. L. Clarke (eds.). Island Press.
14. Curlee, A. P., T. W. Clark, D. Casey, and R. P. Reading. 1994. Large carnivore conservation: Back to the future.
Endangered Species Update. 11(3-4):1-4.
15. Miller, B. J., G. Ceballos, and R. P. Reading. 1994. The prairie dog and biotic diversity. Conservation Biology 8:677681.
16. Miller, B. J., R. P. Reading, C. Conway, J. Jackson, M. Hutchins, S. Forrest, N. Snyder, and S. Derrickson. 1994. A
model for improving endangered species recovery programs. Environmental Management. 18:637-645.
17. Reading, R. P., T. W. Clark, S. R. Kellert. 1994. Attitudes and knowledge of people living in the Greater
Yellowstone Ecosystem. Society and Natural Resources 7:349-365.
18. Reading, R. P., T. W. Clark, and A. Arnold. 1994. Values and attitudes of Hamilton, Victoria residents toward the
threatened eastern barred bandicoot (Perameles gunnii). Anthrozoös 7:255-269.
19. Reading, R. P. and B. J. Miller. 1994. The black-footed ferret recovery program: Unmasking professional and
organizational weaknesses. Pp. 73-100, in: Endangered Species Recovery: Finding the Lessons and Improving the
Process. T. W. Clark, R. P. Reading, and A. L. Clarke (eds.). Island Press.
1995
20. Clark, T. W., G. N. Backhouse, and R. P. Reading. 1995. Prototyping in endangered species recovery programmes:
The eastern barred bandicoot experience. Pp. 50-62, in: People and Nature Conservation: Perspectives on Private
Land Use and Endangered Species Recovery. A. Bennet, G. N. Backhouse, and T. W. Clark (eds.). The Royal
Zoological Society of New South Wales, Mosman, New South Wales, Australia.
21. Clark, T. W., R. P. Reading, and G. N. Backhouse. 1995. Prototyping for successful conservation: The eastern barred
bandicoot program. Endangered Species Update 12(10/11):5-7, 10.
R. P. Reading, Ph.D. — Page 7
22. Mix, H., R. P. Reading, and B. Lhagvasuren. 1995. A systematic census of various large mammals in Eastern and
Southern Mongolia. Proceedings from the Conference on Asian Ecosystems and Their Protection, August 1995.
Ulaanbaatar, Mongolia. (In Russian).
1996
23. Clark, T. W., A. P. Curlee, and R. P. Reading. 1996. Large carnivore conservation: Identifying problems and
developing strategies. Conservation Biology 10:940-948.
24. Reading, R. P. and T. W. Clark. 1996. An interdisciplinary examination of carnivore reintroductions. Pp. 296-336, in:
Carnivore Behavior, Ecology, and Evolution, Vol. II. J. L. Gittleman (ed.). Cornell University Press, Ithaca, NY.
25. Reading, R. P., T. W. Clark, J. H. Seebeck, and J. Pearce. 1996. Habitat suitability index model for the eastern barred
bandicoot (Perameles gunnii). Wildlife Research 23:221-235.
26. Reading, R. P., T. W. Clark, A. Vargas, L. R. Hanebury, B. J. Miller, and D. Biggins. 1996. Recent directions in
black-footed ferret (Mustela nigripes) recovery. Endangered Species Update 13(10-11):1-6.
1997
27. Biggins, D. E., B. J. Miller, T. W. Clark, and R. P. Reading. 1997. Management of an endangered species: The blackfooted ferret. Pp. 420-426, in: Principles of Conservation Biology, 2nd Ed. G. K. Meffe, C. R. Carroll, and
Contributors. Sinauer Assoc., Inc., Sunderland, Mass.
28. Mallon, D. P., S. Dulamtseren, A. Bold, R. P. Reading, and S. Amgalanbaatar. 1997. Mongolia. Pp. 193-201, in:
Wild Sheep and Goats and Their Relatives: Status Survey and Conservation Action Plan for Caprinae, D. M.
Shackleton (ed.) and the IUCN/SSC Caprinae Specialist Group. IUCN, Gland, Switzerland.
29. Reading, R. P. 1997. Wild camels in Mongolia. International Zoo News 44(7):422-423.
30. Reading, R. P., S. Amgalanbaatar, H. Mix, and B. Lhagvasuren. 1997. Argali Ovis ammon surveys in Mongolia’s
South Gobi. Oryx 31:285-294.
31. Reading, R. P., T. W. Clark, and B. Griffith. 1997. The influence of valuational and organizational considerations to
translocation success. Biological Conservation 79:217-225.
32. Reading, R. P., T. W. Clark, A. Vargas, L. R. Hanebury, B. J. Miller, D. Biggins, and P. Marinari. 1997. Blackfooted ferret (Mustela nigripes) conservation update. Small Carnivore Conservation 17:1-6.
33. Reading, R. P. and R. Matchett. 1997. Attributes of black-tailed prairie dog colony in northcentral Montana. Journal
of Wildlife Management 61:664-673.
34. Reading, R. P. and B. J. Miller. 1997. Zoos and a land ethic vision. Pp. 545-460, in: American Zoo and Aquarium
Association Annual Conference Proceedings 1997. Albuquerque Biological Park, Albuquerque, NM.
1998
35. Miller, B., R. Reading, J. Strittholt, C. Carroll, R. Noss, M. Soulé, O. Sánchez, J. Terborgh, D. Brightsmith, T.
Cheeseman, and D. Foreman. 1998/99. Focal species in the design of reserve networks. Wild Earth 8(4): 81-92.
36. Reading, R. P., H. Mix, B. Lhagvasuren, and D. Tseveenmyadag. 1998. The commercial harvest of wildlife in
Dornod Aimag, Mongolia. Journal of Wildlife Management 62:59-71.
1999
37. Clark, T.W., R. P. Reading, and R. L. Wallace. 1999. Research in endangered species conservation: An introduction
to multiple methods. Endangered Species Update 16(5):90-97.
38. Lhagvasuren, B., S. Dulamtseren, L. Amgalan, D. Mallon, G. Schaller, R. Reading, and H. Mix. 1999. Status and
conservation of antelopes in Mongolia. Scientific Research of the Institute of Biological Sciences 1:96-108.
39. Miller, B., K. Ralls, R. P. Reading, J. M. Scott, and J. Estes. 1999. Biological and technical considerations of
carnivore translocation: A review. Animal Conservation 2(1):59-68.
40. Reading, R. P., S. Amgalanbaatar, and L. Lhagvasuren. 1999. Biological assessment of Three Beauties of the Gobi
National Conservation Park, Mongolia. Biodiversity & Conservation 8:1115-1137.
41. Reading, R. P., M. Johnstad, S. Amgalanbaatar, Z. Batjargal, and H. Mix. 1999. Expanding Mongolia’s system of
protected areas. Natural Areas Journal 19(3):211-222.
42. Reading, R. P., B. Miller and S. R. Kellert. 1999. Values and attitudes toward prairie dogs. Anthrozoös 12(1):43-52.
43. Reading, R. P., B. J. Miller, and M. Stanley Price. 1999. Reintroducing animals into the wild: Lessons for giant
pandas. Pp. 18-29, 146-155, in: International Workshop on the Feasibility of Giant Panda Re-introduction,
Woolong Nature Reserve, Sichuan, China, September, 25-29, 1997. S. Mainka (ed.). China Forestry Publishing
House, Beijing, China. (In Chinese and English).
44. Reading, R. P., H. Mix, B. Lhagvasuren, and E. Blumer. 1999. The status wild Bactrian camel and other large
ungulates in south-western Mongolia. Oryx 33(3):247-255.
R. P. Reading, Ph.D. — Page 8
2000
45. Amgalanbaatar, S. and R. P. Reading. 2000. Altai argali. Pp. 5-9, in: Endangered Animals: Conflicting Issues,
Reading, R. P. and B. Miller (eds.). Greenwood Press, Westport, CT.
46. Amgalanbaatar, S., R. P. Reading, and J. Ganchimeg. 2000. Concerns about the effective management and
conservation of Argali sheep in Mongolia. Pp. 16 – 20, in Strategic Planning for Conservation of Mongolian Argali
Sheep (Ovis ammon). Mongolian Ministry for Nature and Environment and World Wide Fund for Nature –
Mongolia. Ulaanbaatar, Mongolia. (in Mongolian).
47. Kleiman, D. G., R. P. Reading, B. J. Miller, T. W. Clark, J. M. Scott, J. Robinson, R. Wallace, R. Cabin, and F.
Fellman. 2000. The importance of improving evaluation in conservation. Conservation Biology 14 (2):1-11.
48. Miller, B., R. P. Reading, and T. W. Clark. 2000. Black-footed ferret. Pp. 54-59, in: Endangered Animals:
Conflicting Issues, Reading, R. P. and B. Miller (eds.). Greenwood Press, Westport, CT.
49. Miller, B., R. Reading, J. Hoogland, T. Clark, G. Ceballos, R. List, S. Forrest, L. Hanebury, P. Manzano, J. Pacheco,
and D. Uresk. 2000. The role of prairie dogs as keystone species: Response to Stapp. Conservation Biology
14:318-321.
50. Reading, R. P. 2000. Giant panda. Pp. 112-118, in: Endangered Animals: Conflicting Issues, Reading, R. P. and B.
Miller (eds.). Greenwood Press, Westport, CT.
2001
51. Berger, J.,S. Dulamtseren, S. Cain, D. Enkhbileg, P. Lichtman, Z. Namshir, G. Wingard, and R. P. Reading. 2001.
Back-casting sociality in extinct species: New perspectives using mass death assemblages and sex ratios.
Proceedings of the Royal Society: Biological Sciences (London) 268:131-139.
52. Clark, T. W., D. J. Mattson, R. P. Reading, and B. J. Miller. 2001. Interdisciplinary problem solving in carnivore
conservation: An introduction. Pp. 223-240, in: Carnivore Conservation. J. L. Gittleman, S. M. Funk, D. W.
Macdonald, and R. K. Wayne (eds.). Cambridge University Press, Cambridge, UK.
53. Kenny, D. and R. P. Reading. 2001. Field research into argali. International Zoo News 48(4): 257-258.
54. Miller, B., B. Dugelby, D. Foreman, C. Martinez del Río, R. Noss, M. Phillips, R. Reading, M. Soulé, J. Terborgh,
and L. Wilcox. 2001. The importance of large carnivores to healthy ecosystems. Endangered Species UPDATE 18:
202-210.
55. Reading, R. P., S. Amgalanbaatar, and G. Wingard. 2001. Argali Sheep Conservation and Research Activities in
Mongolia. Open Country 3: 25-32.
56. Reading, R. P. and B. J. Miller. 2001. Release and Reintroduction of species. Pp. 1053-1057, in: Encyclopedia of the
World’s Zoos. C. Bell (ed.). Fitzroy Dearborn Publishers, Chicago, IL.
57. Reading, R. P. and B. J. Miller. 2001. Translocation. Pp. 1262-1263, in: Encyclopedia of the World’s Zoos. C. Bell
(ed.). Fitzroy Dearborn Publishers, Chicago, IL.
58. Reading, R. P., H. Mix, B. Lhagvasuren, C. Feh, D. Kane, S. Dulamtseren, and S. Enkhbold. 2001. Status and
Distribution of khulan (Equus hemionus) in Mongolia. Journal of Zoology (London) 254: 381-389.
2002
59. Amgalanbaatar, S., R. P. Reading, S. Dulamtseren, Yo. Onon, Sh. Tumentsetseg, and N. Batsukh. 2002. Assessment
of argali sheep (Ovis ammon) distribution in Mongolia using GIS. Proceedings of the Institute of Biology of the
Mongolian Academy of Sciences (Mammalogical Studies in Mongolia and Its Adjacent Territories) 24: 26-34. (In
Mongolian)
60. Amgalanbaatar, S., R. P. Reading, B. Lhagvasuren, and N. Batsukh. 2002. Argali sheep (Ovis ammon) trophy hunting
in Mongolia. Pirineos 157: 129-150.
61. Blumer, E. S, Z. Namshir, T. Tuya, B. Mijiddorj, R. P. Reading, and H. Mix. 2002. Veterinarian aspects of wild
Bactrian camel conservation in Mongolia. Pp. 115-122, in: Ecology and Conservation of Wild Bactrian Camels
(Camelus bactrianus ferus). R. P. Reading, D. Enkhbileg, and T. Galbaatar (eds.). Mongolian Conservation
Coalition & Admon Printing, Ulaanbaatar, Mongolia.
62. Clark, T. W., R. P. Reading, R. L. Wallace, and B. Wilson. 2002. If the Tasmanian tiger was found, what should we
do? An interdisciplinary guide to endangered species recovery. Endangered Species Update 19:194-200.
63. Enkhbileg, D., S. Dulamtseren, S. Namshir, L. Sandui, R. Reading, and J. Berger. 2002. Distribution and population
of the Asiatic wild ass khulan, Equus hemionus, in the Small Gobi Strictly Protected Area, adjacent territory.
Proceedings of the Biological Institute of the Mongolian Academy of Sciences 23: 105-116. (In Mongolian).
64. Feh, C., N. Shah, M. Rowan, R. Reading, and S. P. Goyal. 2002. Status and action plan for the Asiatic wild ass
(Equus hemionus). Pp. 62-71, in: Equids: Zebras, asses and horses. Status survey and conservation action plan. P.
D. Moehlman (ed.). IUCN, Gland, Switzerland and Cambridge, UK.
R. P. Reading, Ph.D. — Page 9
65. Lyda, R., J. Kirkpatrick, R. P. Reading, and E. S Blumer. 2002. Bactrian camel fecal steroid analysis. Pp. 79-84, in:
Ecology and Conservation of Wild Bactrian Camels (Camelus bactrianus ferus). R. P. Reading, D. Enkhbileg, and
T. Galbaatar (eds.). Mongolian Conservation Coalition & Admon Printing, Ulaanbaatar, Mongolia.
66. McCain, L. A., R. P. Reading, and B. J. Miller. 2002. Prairie dog gone: Myth, persecution, and preservation of a
keystone species. Pp. 230-235, in:Welfare Ranching: The Subsidized Destruction of the American West. M.
Wuerthner and G. Wuerthner (eds.). Island Press, Washington, D.C.
67. Miller, B. and R. P. Reading. 2002. The black-tailed prairie dog: Threats to survival and a plan for conservation.
Wild Earth 12(1): 46-55.
68. Mix, H., R. P. Reading, E. S Blumer, and B. Lhagvasuren. 2002. Status and distribution of wild Bactrian camels in
Mongolia. Pp. 39-48, in: Ecology and Conservation of Wild Bactrian Camels (Camelus bactrianus ferus). R. P.
Reading, D. Enkhbileg, and T. Galbaatar (eds.). Mongolian Conservation Coalition & Admon Printing,
Ulaanbaatar, Mongolia.
69. Onon. Yo., S. Amgalanbaatar, and R. P. Reading. 2002. Influence of winter on the argali population of Ikh Nart.
Proceedings of the Institute of Biology of the Mongolian Academy of Sciences (Mammalogical Studies in
Mongolia and Its Adjacent Territories) 24:111-119. (In Mongolian).
70. Onon, Yo., S. Amgalanbaatar, R. P. Reading, and J. Ganchimeg. 2002. Behavior and home range of argali sheep
(Ovis ammon L., 1758). Proceedings of the Biological Institute of the Mongolian Academy of Sciences 23: 81-88.
(In Mongolian).
71. Reading, R. P., T. W. Clark, L. A. McCain, and B. J. Miller. 2002. Black-tailed prairie dog conservation: A new
approach for a 21st century challenge. Endangered Species Update 19: 162-170.
72. Reading, R. P., D. Enkhbileg, and T. Galbaatar. 2002. An introduction to the wild Bactrian camel. Pp. 5-10, in:
Ecology and Conservation of Wild Bactrian Camels (Camelus bactrianus ferus). R. P. Reading, D. Enkhbileg, and
T. Galbaatar (eds.). Mongolian Conservation Coalition & Admon Printing, Ulaanbaatar, Mongolia.
73. Reading, R. P., A. Magash, T. Galbaatar, and R. Indra. 2002. Conclusions: Prospects for the wild Bactrian camel’s
future. Pp. 149-152, in: Ecology and Conservation of Wild Bactrian Camels (Camelus bactrianus ferus). R. P.
Reading, D. Enkhbileg, and T. Galbaatar (eds.). Mongolian Conservation Coalition & Admon Printing,
Ulaanbaatar, Mongolia.
74. Reading, R. P., H. Mix, E. S Blumer, S. Amgalanbaatar, T. Galbaatar, G. Wingard, Ts. Tuya, M. D. Johnstad, Z.
Namshir, and B. Lhagvasuren. 2002. Wild ungulate conservation in Mongolia. Proceedings of the Institute of
Biology of the Mongolian Academy of Sciences (Mammalogical Studies in Mongolia and Its Adjacent Territories)
24:120-132.
75. Reading, R. P., H. Mix, E. Blumer, T. Galbaatar, and B. Lhagvasuren. 2002. Wild Bactrian camel conservation
project: Recent activities and future plans. Pp. 129-138, in: Ecology and Conservation of Wild Bactrian Camels
(Camelus bactrianus ferus). R. P. Reading, D. Enkhbileg, and T. Galbaatar (eds.). Mongolian Conservation
Coalition & Admon Printing, Ulaanbaatar, Mongolia.
76. Wallace, R. L., T. W. Clark, and R. P. Reading. 2002. Introduction: Interdisciplinary endangered species
conservation: A new approach for a new century. Endangered Species Update 19: 70-73.
77. Wallace, R. L., T. W. Clark, and R. P. Reading. 2002. Conclusion: Becoming a more effective professional: The next
steps in learning and applying an interdisciplinary approach to the conservation of biological diversity. Endangered
Species Update 19: 201-202.
2003
78. Reading, R. P., S. Amgalanbaatar, D. Kenny, Yo. Onon, Z. Namshir, and A. DeNicola. 2003. Argali ecology in Ikh
Nartiin Chuluu Nature Reserve: Preliminary Findings. Mongolian Journal of Biological Sciences 1(2): 3-14.
79. Smith, J., J. Clanahan, M. DeMarco, R. Howard, D. Foreman, R. Reading, and B. Miller. 2003. Implementation and
conservation action. Pp. 115-133, in: Southern Rockies Wildlands Network Vision: A science-based approach to
rewilding the Southern Rockies. B. Miller, D. Foreman, M. Fink, D. Shinneman, J. Smith, M. DeMarco, M. Soule,
and R. Howard (eds.). Colorado Mountain Club Press, Golden, CO.
80. Tuya, Ts., R. P. Reading, R. R. Ramey II, O. A. Ryder, Ts. Zhanchiv, and Kh. Tumennasan. 2003. New perspectives
on the subspecies, population structure, and conservation of Mongolian argali sheep (Ovis ammon). Proceedings of
the Institute of Biology of the Mongolian Academy of Sciences (Mammalogical Studies in Mongolia and Its
Adjacent Territories) 24:141-158.
2004
81. Adiya, Ya., R. P. Reading, E. Blumer, H. Mix, B. Mijiddorj, and S. Choijin. 2004. Summary of recent research on
wild Bactrian camels (Camelus bactrianus ferus Przewalski, 1883). Pp. 38-45, in: Some Studies in the Trans-Altai
R. P. Reading, Ph.D. — Page 10
82.
83.
84.
85.
Gob Ecosystem. Ya. Adiya and K. Ulikpan (eds.). United Nations Development Project-Global Environment
Facility, Conservation of the great Gobi and its Umbrella Species, Ulaanbaatar, Mongolia. (In Mongolian).
Miller, B., W. Conway, R. P. Reading, C. Wemmer, D. Wildt, D. Kleiman, S. Monfort, A. Rabinowitz, B.
Armstrong, and M. Hutchins. 2004. Evaluating the conservation mission of zoo, aquariums, botanical gardens, or
natural history museums. Conservation Biology 18: 1-8.
Reading, R. P., D. G. Kleiman, and B. J. Miller. 2004. Conservation and behavior: Species reintroductions. Pp. 426 435, in: Encyclopedia of animal behavior. Volume 1: A-C. M. Bekoff (ed.). Greenwood Press, Westport, CT.
Tserenbataa, T., R. R. Ramey II, O. A. Ryder, T. W. Quinn, and R. P. Reading. 2004. A population genetic
comparison of argali sheep (Ovis ammon) in Mongolia using the ND5 gene of mtDNA; Implications for
conservation. Molecular Ecology 13: 1333-1339.
Zahler, P., B. Lhagasuren, R.P. Reading, J.R. Wingard, S. Amgalanbaatar, S. Gombobaatar, N.W.H. Barton, and Y.
Onon. 2004. Illegal and unsustainable wildlife hunting and trade in Mongolia. Mongolian Journal of Biological
Sciences 2:23-32.
2005
86. Mandakh, B., G. Wingard, and R. Reading. 2005. Winter pasture conditions and forage use by argali (Ovis ammon)
in Gobi Gurvan Saykhan National Park. Erforschung Biologischer Ressourcen der Mongolei (Halle/Saale) 9: 7176.
87. Meadow, R., R. P. Reading, M. Phillips, M. Mehringer, and B. J. Miller. 2005. The influence of persuasive
arguments on public attitudes toward a proposed wolf restoration in the southern Rockies. Wildlife Society Bulletin
33(1): 154-163.
88. Miller, S. D., R. P. Reading, B. Haskins, and D. Stern. 2005. Overestimation bias in estimate of black-tailed prairie
dog abundance in Colorado. Wildlife Society Bulletin 33: 1444-1451.
89. Reading, R. P., S. Amgalanbaatar, D. Kenny, and B. Dashdemberel. 2005. Cinereous vulture nesting ecology in Ikh
Nartyn Chuluu Nature Reserve, Mongolia. Mongolian Journal of Biological Sciences 3(1): 13-19.
90. Reading, R. P., S. Amgalanbaatar, G. J. Wingard, D. Kenny, and A. DeNicola. 2005. Ecology of argali in Ikh Nartiin
Chuluu, Dornogobi Aimag. Erforschung Biologischer Ressourcen der Mongolei (Halle/Saale) 9: 77-89.
91. Reading, R. P., E. Blumer, H. Mix, and Ya. Adiya. 2005. Wild Bactrian camel conservation. Erforschung
Biologischer Ressourcen der Mongolei (Halle/Saale) 9: 91-100.
92. Reading, R. P., L. McCain, T. W. Clark, and B. J. Miller. 2005. Understanding and Resolving the Black-tailed Prairie
Dog Conservation Challenge. Pp. 209-223, in: People and Wildlife: Conflict or Coexistance? R. Woodroffe, S.
Thirgood, and A. Rabinowitz (Eds.). Cambridge University Press, Cambridge, UK.
93. Reading, R. P., L. McCain, T. W. Clark, and B. J. Miller. 2005. Understanding the Controversy Surrounding Blacktailed Prairie Dog Restoration Efforts. Pp. 166-189, in: Facets of Grassland Restoration. A. Struchov and J.
Kuleshova (Eds.). Biodiversity Conservation Center, Moscow, Russia.
2006
94. Amgalanbaatar, S., O. Shagdarsuren, R. Reading, and Yo. Onon. 2006. Pasture overlap between argali sheep and
livestock in state border area of Uvs Province. Pp. 88-92, in: Natural conditions, reserves, and biodiversity of the
Mongolian Altai-Sayan Ecoregion. D. Dash (Ed.). Altai-Sayan UNDP-GEF Project, Ulaanbaatar, Mongolia. (in
Mongolian).
95. Amgalanbaatar, S., O. Shagdarsuren, R. Reading, and Yo. Onon. 2006. Collared argali movement and home ranges
in Bokhmoron Soum of Uvs Aimag. Pp. 93-97, in: Natural conditions, reserves, and biodiversity of the Mongolian
Altai-Sayan Ecoregion. D. Dash (Ed.). Altai-Sayan UNDP-GEF Project, Ulaanbaatar, Mongolia. (in Mongolian).
96. Biggins, D. E., B. J. Miller, T. W. Clark, and R. P. Reading. 2006. Restoration of an endangered species: The blackfooted ferret. Pp. 581-585, in: Principles of Conservation Biology, 3rd Edition. M. A. Groom, G. K. Meffe, and C.
R. Carroll (Eds.). Sinauer Associates, Sunderland, MA.
97. Kotliar, N. B., B. Miller, R. P. Reading, and T. W. Clark. 2006. The prairie dog as a keystone species. Pp. 248-260,
in: Conservation of the black-tailed prairie dog: Saving North America’s western grasslands. J. L. Hoogland (Ed.).
Island Press, Washington, D.C.
98. Lamb, B. L., R. P. Reading, and W. F. Andelt. 2006. Attitudes and perceptions about prairie dogs. Pp. 108-114, in:
Conservation of the black-tailed prairie dog: Saving North America’s western grasslands. J. L. Hoogland (Ed.).
Island Press, Washington, D.C.
99. Miller, B., and R. P. Reading. 2006. A proposal for effective conservation of prairie dogs. Pp. 53-64, in:
Conservation of the black-tailed prairie dog: Saving North America’s western grasslands. J. L. Hoogland (Ed.).
Island Press, Washington, D.C.
R. P. Reading, Ph.D. — Page 11
100. Murdoch, J. D., S. Batdorj, S. Buyandelger, D. Kenny, and R. P. Reading. 2006. Ecology of the Daurian hedgehog
(Hemiechinus dauuricus) in Ikh Nart Nature Reserve, Mongolia: Preliminary findings. Mongolian Journal of
Biological Sciences 4(2): 25-32.
101. Murdoch, J. D., Ts. Munkhzul, S. Amgalanbaatar, and R. P. Reading. 2006. Checklist of mammals in Ikh Nart
Nature Reserve. Mongolian Journal of Biological Sciences 4(2): 69-74.
102. Reading, R. P., D. J. Bedunah, and S. Amgalanbaatar. 2006. Conserving biodiversity on Mongolian rangelands:
Implications for protected area development and pastoral uses. Pp 1-17, in: D. J. Bedunah,E. D. McArthur, M.
Fernandez-Gimenez (Comps.) Rangelands of Central Asia: Transformations, Issues and Future Challenges. 2004
January 27; Salt Lake City, UT. RMRS-P-39. Fort Collins, CO: U.S. Department of Agriculture, Forest Service,
Rocky Mountain Research Station.
103. Reading, R. P., D. Stern, and L. A. McCain. 2006. Attitudes and knowledge of natural resources agency personnel
towards black-tailed prairie dogs (Cynomys ludovicianus). Conservation and Society 4(4): 592-627.
104. Shipley, B. and R. P. Reading. 2006. A comparison of herptofauna and small mammal diversity on black-tailed
prairie dog (Cynomys ludovicianus) colonies and non-colonized grasslands in Colorado. Journal of Arid Environ.
66(1): 27-41.
2007
105. Miller, B. J., R. P. Reading, D. E. Biggins, J. K. Detling, S. C. Forrest, J. L. Hoogland, J. Javersak, S. D. Miller, J.
Proctor, J. Truett, and D. W. Uresk. 2007. Prairie dogs: An ecological review and current biopolitics. Journal of
Wildlife Management 71(8): 2801-2810.
106. Murdoch, J. D., T. Munkhzul, S. Buyandelger, and R. P. Reading. 2007. Biotope of corsac fox and red fox in
Central Asia. Mongolian Journal of Biological Sciences 5(1-2): 7-12.
107. Reading, R. P. and B. J. Miller. 2007. Attitudes and attitude change among zoo visitors. Pp. 63-91, in: Zoos in the
21st Century: Catalysts for Conservation? A. Zimmermann, M. Hatchwell, L. Dickie and C. West (Eds.).
Cambridge University Press, Cambridge UK.
108. Reading, R. P., S. Amgalanbaatar, D. Kenny, A. DeNicola, and E. Tuguldur. 2007. Siberian ibex (Capra sibirica)
home ranges in Ikh Nart Nature Reserve, Mongolia: preliminary findings. Mongolia Journal of Biological
Sciences 5(1-2): 29-37.
2008
109. Batbayar, N., R. Reading, D. Kenny, T. Natsagdorgj, and P.W. Kee. 2008. Migration and movement patterns of
cinereous vultures in Mongolia. Falco 32: 5-7.
110. Dulamtseren S., R. Reading, Shar S., J. D. Murdoch, Gantulga J., and Usukhjargal D. 2008. Contribution to the
previous distribution range of marbled polecat (Vormela peregusna) in Mongolia. Proceedings of the Institute of
Biology of the Mongolian Academy of Sciences 27:12-14. (In Mongolian)
111. Harris, R .B. and R. Reading. 2008. Ovis ammon. In: IUCN 2012. IUCN Red List of Threatened Species. Version
2012.2.
112. Kenny, D. E., A. J. DeNicola, S. Amgalanbaatar, Z. Namshir, G. Wingard, T. Tuya, and R. P. Reading. 2008. Three
field capture techniques for free-ranging argali sheep (Ovis ammon) in Mongolia. Zoo Biology 27: 137-144.
113. Murdoch, J. D., K. Ralls, B. L. Cypher, and R. P. Reading. 2008. Barking vocalizations in San Joaquin kit foxes
(Vulpes macrotis mutica). Southwestern Naturalist 53(1): 118-124.
114. Murdoch, J. D., K. Ralls, B. Cypher, and R. P. Reading. 2008. Social interactions among San Joaquin kit foxes
before, during, and after the mating season. Journal of Mammalogy 89(5): 1087-1093.
115. Shipley, B., R. P. Reading, and B. Miller. 2008. Comparing capture rates of reptiles and amphibians in black-tailed
(Cynomys ludovicianus) prairie dog colonies and uncolonized prairie in Colorado, USA. Western North American
Naturalist 62(2): 245-248.
2009
116. Bernal Stoopen, J., J. M. Packard, and R. P. Reading. 2009. Mexican wolf recovery: Insights from transboundary
stakeholders. Pp. 115-132, in L. López-Hoffman, E.D. McGovern, R.G. Varady and K.W. Flessa (eds.).
Conservation of Shared Environments: Learning from the United States and Mexico. Tucson: University of
Arizona Press, Tucson, AZ.
117. Dulamtseren, S., S. Shar, J. D. Murdoch, R. P. Reading, J. Gantulga, D. Usukhjargal, and S. Buyandelger. 2009.
Contributions to the distribution of the marbled polecat (Vormela peregusna Guldenstaedt, 1770) in Mongolia.
Small Carnivore Conservation 40: 29-32.
118. Kenny, D., N. Batbayar, P. Tsolmonjav, M. J. Willis, J. Azua, and R. P. Reading. 2009. Dispersal of Eurasian Black
Vulture (Aegypius monachus) Fledglings from the Ikh Nart Nature Reserve, Mongolia. Vulture News 59: 13-19.
R. P. Reading, Ph.D. — Page 12
119. Murdoch, J., T. Munkhzul, S. Buyandelger, and R. Reading. 2009. Body size and sexual dimorphism among a
population of corsac and red foxes in central Mongolia. Mammalia 73: 72-75.
120. Murdoch, J.D., T. Munkhzul, S. Buyandelger, R.P. Reading, and C. Sillero-Zubiri. 2009. The Siberian marmot as a
keystone species: Observations of burrow use by corsac foxes in Mongolia and implications for conservation.
Oryx 43(3): 431-434.
121. Reading, R. P. 2009. Vertebrate species associated with prairie dogs. Pp. 139-146, in Prairie dogs: Communication
and community in an animal society. C. Slobodchikoff, B. S. Perla, and J. L. Verdolin. Harvard University Press,
Cambridge, MA.
122. Reading, R.P., D. Kenny, S. Amgalanbaatar, A. DeNicola, and G. Wingard. 2009. Argali Lamb (Ovis ammon)
morphometric measurements and survivorship in Mongolia. Mammalia 73: 98-104.
2010
123. Clark, S.G., B.J. Miller, and R.P. Reading. 2010. Policy: Integrated problem-solving as an approach to wolf
management. Pp. 147-157, in: Awakening Spirits: Wolves in the Southern Rockies. R. P. Reading, B. J. Miller, A.
Masching, R. Edward, and M. Phillips (Eds.). Fulcrum Publishing, Golden, CO.
124. Edwards, R. and R. P. Reading. 2010. Saving the world’s grasslands: An introduction. Great Plains Research 20(1):
5-8.
125. Maude, G. and R. P. Reading. 2010. The role of ecotourism in biodiversity and grassland conservation in Botswana.
Great Plains Research 20(1): 109-120.
126. Miller, B.J., R.P. Reading, and H. Gosnell 2010. History of public attitudes toward wolves and wolf recovery. Pp.
103-117, in: Awakening Spirits: Wolves in the Southern Rockies. R. P. Reading, B. J. Miller, A. Masching, R.
Edward, and M. Phillips (Eds.). Fulcrum Publishing, Golden, CO.
127. Miller, B.J., R.P. Reading, K. Ralls, S.G. Clark, and J. Estes. 2010. Variables influencing carnivore translocation.
Pp. 61-75, in: Awakening Spirits: Wolves in the Southern Rockies. R. P. Reading, B. J. Miller, A. Masching, R.
Edward, and M. Phillips (Eds.). Fulcrum Publishing, Golden, CO.
128. Murdoch, J.D., T. Munkhzul, S. Buyandelger, R. P. Reading, and C. Sillero-Zubiri. 2010. Seasonal food habits of
corsac and red foxes in Mongolia and the potential for competition. Mammalian Biology 75: 36-44.
129. Murdoch, J. D., B. Suuri, and R. P. Reading. 2010. Estimates of toad-headed agama density in three steppe habitats
of Mongolia. Erforschung Biologischer Ressourcen der Mongolei (Halle/Saale) 11:383-389.
130. Paquet, P., B.J. Miller, K. Kunkel, R.P. Reading, and M. Phillips. 2010. The importance of large carnivores. Pp. 4959, in: Awakening Spirits: Wolves in the Southern Rockies. R. P. Reading, B. J. Miller, A. Masching, R. Edward,
and M. Phillips (Eds.). Fulcrum Publishing, Golden, CO.
131. Reading, R. P., D. J. Bedunah, and S. Amgalanbaatar. 2010. Conserving Mongolia's grasslands with challenges,
opportunities, and lessons for America's Great Plains. Great Plains Research 20(1): 85-108.
132. Reading, R. P., D. Kenny, J. Azua, T. Garrett, M. J. Willis, and P. Tsolmonjav. 2010. Ecology of Eurasian black
vultures (Aegypius monachus) in Ikh Nart Nature Reserve, Mongolia. Erforschung Biologischer Ressourcen der
Mongolei (Halle/Saale) 11: 177-188.
133. Reading, R. P., D. Kenny, S. Batdorj, and J. Murdoch. 2010. Comparative morphology of two sympatric species of
hedgehog in Ikh Nart Nature Reserve, Mongolia. Erforschung Biologischer Ressourcen der Mongolei
(Halle/Saale) 11:323-328.
134. Reading, R. P. and B. Miller. 2010. Captive breeding ethics. Pp. 101-105, in: Encyclopedia of Animal Rights and
Animal Welfare, 2nd Edition, Volume 1: A-H. M. Bekoff (Ed.). Greenwood Press, Santa Barbara, CA.
135. Reading, R.P., B. Miller, A. Masching, R. Edward, and M. Phillips. 2010. Introduction: A comprehensive approach
to evaluating the potential for wolf restoration in the Southern Rockies. Pp. 1-9, in: Awakening Spirits: Wolves in
the Southern Rockies. R. P. Reading, B. J. Miller, A. Masching, R. Edward, and M. Phillips (Eds.). Fulcrum
Publishing, Golden, CO.
136. Singh, N. J., S. Amgalanbaatar, and R. P. Reading. 2010. Temporal dynamics of group size and sexual segregation
in Ibex. Erforschung Biologischer Ressourcen der Mongolei (Halle/Saale) 11: 315-322.
137. Singh, N. J., S. Amgalanbaatar, and R. P. Reading. 2010. Grouping patterns of argali in Ikh Nart Nature Reserve,
Mongolia. Mongolian Journal of Biological Sciences 8(2): 7-13.
2011
138. Reading, R. P., A. Bräunlich and H. Mix. 2011. Red-fronted Serin Serinus pusillus: First record for Mongolia.
Birding Asia 15: 102-103.
139. Reading, R. P., S. Harris, and A. Bräunlich. 2011. Rufous-bellied woodpecker Dendrocopos hyperythrus: First
records for Mongolia. Birding Asia 15: 104-105.
R. P. Reading, Ph.D. — Page 13
140. Reading, R. P., R. Haeffner, V. Veer, and N. Mitchell. 2011. Reintroduction of Turks and Caicos rock iguana
(Cyclura carinata) to Long Cay. Pp. 120-124, in: Global Re-introduction Perspectives: 2011. More case studies
from around the globe, P.S. Soorae (Ed.). IUCN/SSC Re-introduction Specialist Group. Abu Dhabi, UAE.
141. Reading, R.P., T.J. Weaver, J.R. Garcia, R. Elias Piperis, M.T. Herbert, C. Cortez, A. Muñoz, J.E. Rodríguez & Y.
Matamoros (Eds.). 2011. Taller para establecer la estrategia de Conservación de la Rana del Titicaca (Telmatobius
culeus). 13-15 de diciembre, 2010. Facultad de Ciencias Biológicas, Universidad Nacional del Altiplano, Puno,
Perú. Conservation Breeding Specialist Group (SSC/IUCN)/CBSG Mesoamerica
142. Wingard, G. J., R. B. Harris, S. Amgalanbaatar, and R. P. Reading. 2011. Estimating abundance of mountain
ungulates incorporating imperfect detection: Argali in the Gobi Desert, Mongolia. Wildlife Biology 17(1): 93 101.
143. Wingard, G. J.., R. B. Harris, D. H. Pletscher, D. J. Bedunah, M. Bayart, A. Sukh, and R. P. Reading. 2011. Argali
food habits and dietary overlap with domestic livestock in Ikh Nart Nature Reserve, Mongolia. Journal of Arid
Environments 75(2): 138-145.
144. Young, J. K., K. A. Olson, R. P. Reading, S. Amgalanbaatar, and J. Berger. 2011. Is wildlife going to the dogs?
Impacts of feral and free-ranging dogs on wildlife populations. BioScience 61(2): 125-132.
2012
145. Amgalanbaatar, S., R. P. Reading, and Ts. Z. Dorzhiev. 2012. Argali sheep (Ovis ammon) current population level
and the issues of trophy hunting in Mongolia. Vyestnik Spyevipusk V 2012: 290-293. (In Russian)
146. Amgalanbaatar, S., R. P. Reading, and Ts. Z. Dorzhiev. 2012. The ecology of argali in “Ikh Nart” Nature Reserve,
Mongolia. Vyestnik Biologia, Gegrafia 2012 (4): 105-108. (In Russian).
147. Amgalanbaatar, S., R. P. Reading, and Ts. Z. Dorzhiev. 2012. Argali sheep (Ovis ammon) current population level
and the issues of trophy hunting in Mongolia. Vyestnik Biologia, Gegrafia 2012 (4): 109-112. (In Russian)
148. Biggins, D. E., B. J. Miller, T. W. Clark, and R. P. Reading. 2012. Restoration of an endangered species: The blackfooted ferret. In: Principles of Conservation Biology, 4th Ed. G. K. Meffe, C. R. Carroll, and Contributors.
Sinauer Assoc., Inc., Sunderland, Mass.
149. Davie, H. S., J. D. Murdoch, N. Nandin-Erdene, and R. P. Reading. 2012. A survey of bat diversity at Ikh Nart
Nature Reserve, Mongolia. Mongolian Journal of Biological Sciences 10(1-2): 33-40.
150. Magle, S. B., K. A. Salamack, K. R. Crooks, and R. P. Reading. 2012. Effects of habitat fragmentation and blacktailed prairie dogs on urban avian diversity. Biodiversity and Conservation 21: 2803-2821.
151. Miller, B. J., and R. P. Reading. 2012. Challenges to black-footed ferret recovery: Protecting prairie dogs. Western
North American Naturalist 72 (2): 228-240.
152. Munkhzul, Ts., B. Buuveibaatar, J. D. Murdoch, R. P. Reading, and R. Samiya. 2012. Factors affecting home ranges
of red foxes in Ikh Nart Nature Reserve, Mongolia. Mongolia. Mongolian Journal of Biological Sciences 10(1-2):
51-58.
153. Munkhzul, T., J. D. Murdoch, and R. P. Reading. 2012. Home range characteristics of corsac and red foxes in
Mongolia. Erforschung Biologischer Ressourcen der Mongolei (Halle/Saale) 12: 93-104.
154. Sidle, J. G., D. J. Augustine, D. H. Johnson, S. D. Miller, J. F. Cully, Jr., and R. P. Reading. 2012. Aerial surveys
adjusted by ground surveys to estimate area occupied by black-tailed prairie dogs. Wildlife Society Bulletin 36(2):
248-256.
155. Yadamsuren, A., E. Dulamtseren, and R. P. Reading. 2012. Conservation status and management of wild Bactrian
camels in Mongolia. Pp. 45-54, in: Camels in Asia and North Africa: Interdisciplinary perspectives on their
significance in past and present, E Knoll and P. Burger (eds.). Austrian Academy of Science, Vienna, Austria.
156. Zapletal, M., Batdorj S., J. Atwood, J. D. Murdoch, and R. P. Reading. 2012. Home range characteristics and habitat
selection by Daurian hedgehogs (Mesechinus dauuricus) in Ikh Nart Nature Reserve, Mongolia. Mongolian
Journal of Biological Sciences 10(1-2): 41-50.
2013
157. Aber, B., R. Callas, G. Chapron, J. Clark, J. P. Copeland, B. Giddings, R. Inman, J. Ivan, R. Kahn, C. Long, A.
Magoun, J. Mattisson, D. McCaouley, K. McKelvey, M. Miller, R. Monello, B. Oaklead, E. Odell, J. Persson, R.
Reading, S. Sarorius, M. Schwartz, T. Shenk, M. Sirochman, J. Squires, S. Wait, M. Wild, and L. Wolfe. 2013.
Restoration of wolverines: Considerations for translocation and post-release monitoring. 51 pp. Wildlife
Conservation Society, New Work. http://www.wcsnorthamerica.org/Wildlife/Wolverin.aspx.
158. Bragin, N., N. J. Singh, and R. P. Reading. 2013. Creating a ruggedness layer for use in habitat suitability modeling
for Ikh Nart Nature Reserve, Mongolia. Mongolian Journal of Biological Sciences 11(1-2): 19-23.
R. P. Reading, Ph.D. — Page 14
159. Chimeddorj, B., B. Buuveibaatar, Yo. Onon, O. Munkhtogtokh, and R. P. Reading. 2013. Identifying potential
conservation corridors along the Mongolia-Russian border using resource selection functions: A case study on
Argali Sheep. Mongolian Journal of Biological Sciences 11(1-2): 45-53.
160. Hiener, J. B. Yunden, J. Kiesecker, G. Davaa, M. Ganbaatar, O. Ichinkhorloo, H. von Wehrden, R. Reading, K.
Olsen, R. Jackson, J. Evans, B. McKenny, J. Oakleaf, K. Sochi, and E. Oidov. 2013. Identifying conservation
priorities in the face of future development: Applying development by design in the Mongolian Gobi. The Nature
Conservancy, Ulaanbaatar, Mongolia. 62 pp. + append.
161. Kenny, D. E., C. L. Bickel, and R. P. Reading. 2013. Veterinary assessment for free-ranging Eurasian Black Vulture
(Aegypius monachus) chicks in southeastern Mongolia. Topics in Companion Animal Medicine 28(4): 143-150.
162. Murdoch, J., H. Davie, M. Galbadrakh, T. Donovan, and R. P. Reading. 2013. Do Siberian marmots influence toadheaded agama occupancy? Examining the influence of marmot colonies and three steppe habitats in Mongolia.
Journal of Arid Environments 92: 76-80.
163. Otgonbayar B., Buyandelger S. and R. Reading. 2013. Chestnut-flanked White-eye Zosterops erythropleurus: First
record for Mongolia. Birding Asia 20: 111-112.
164. Reading, R. P., D. E. Kenny, and K. Fitzgerald. 2013. The critical contribution of veterinarians to conservation
biology. Topics in Companion Animal Medicine 28(4): 131-134.
165. Reading, R. P., B. Miller, and D. Shepherdson. 2013. The value of enrichment to reintroduction success. Zoo
Biology 32: 332-341.
2014
166. Amgalanbaatar, S., Ts. Z. Dorzhiev, and R. P. Reading. 2014. Structure of argali sheep Ovis ammon habitat in
Mongolia and its dynamics in the initial period of the 21 st century. Vyestnik Biologia, Gegrafia 2014 4(1): 58-66.
(In Russian)
167. Batsaikhan , N., B. Buuveibaatar, B.Chimed-Ochir, O. Enkhtuya, D. Galbrakh, O. Ganbaatar, B. Lkhagvasuren,
D.Nandintsetseg, J. Berger, J.M. Calabrese, A. E. Edwards, W. F. Fagan, T. K. Fuller, M. Heiner, T. Y. Ito, P.
Kaczensky, P. Leimgruber, A. Lushchekina, E. J. Milner-Gulland, T. Mueller, M. G. Murray, K. A. Olson, R.
Reading, G. B. Schaller, A. Stubbe, M. Stubbe, C. Walzer, H. von Wehrden, and T. Whitten. 2014. Crossing a
new line: conserving the world’s finest grassland amidst ambitious national development. Conservation Biology
28(6): 1736-1739.
168. Davie, H.S., J. D. Murdoch, A. Lhagvasuren, and R. P. Reading. 2014. Measuring and mapping the influence of
landscape factors on livestock predation by wolves in Mongolia. Journal of Arid Environments 103:85-91.
169. Kaczensky, P. Y. Adiya, H. von Wehrden, B. Mijiddorj, C. Walzer, D. Güthlin, D. Enkhbileg, and R. P. Reading.
2014. Space and habitat use by wild Bactrian camels in the Transaltai Gobi of southern Mongolia. Biological
Conservation 169: 311-318.
170. Reading, R. P., G. Maude, P. Hancock, D. Kenny, and R. Garbett. 2014. Comparing different types of patagial tags
for use on vultures. Vulture News 67: 33-42.
2015
171. Reading, R. P., G. Wingard, T. Selenge, and S. Amgalanbaatar. 2015. The crucial importance of protected areas to
conserving Mongolia’s natural heritage. Pp. 257-265, in: Protecting the wild: Parks and wilderness, the foundation
for conservation, Wuerthner, G., E. Crist, and T. Butler (eds.). Island Press, Washington, D.C.
172. Kenny, D., Y. J. Kim, H. Lee, and R. P. Reading. 2015. Blood lead levels for Eurasian Black Vultures (Aegypius
monachus) migrating between Mongolia and the Republic of Korea. Journal of Asian-Pacific Biodiversity 8(3):
199-202.
173. Kenny, D., R. P. Reading, and H. Lee. 2015. Blood-gas analysis from free-ranging Eurasian Black Vultures
(Aegypius monachus) in Mongolia and the Republic of Korea. Research & Reviews: Journal of Veterinary
Sciences 1(1): 1-9.
174. Kenny, D., R. P. Reading, G. Maude, P. Hancock, and R. Garbett. 2015. Blood lead levels in White-Backed
Vultures (Gyps africanus) from Botswana, Africa. Vulture News 68:25-31.
175. Zapletal, M., Batdorj S., J. Atwood, J. D. Murdoch, and R. P. Reading. 2015. Fine-scale habitat use by Daurian
hedgehogs in the Gobi-steppe of Mongolia. Journal of Arid Environments 114:100-103.
R. P. Reading, Ph.D. — Page 15
In press
176. Kenney, J. J., J. Detling, and R. P. Reading. In press. Influence of black-tailed prairie dogs (Cynomys ludovicianus)
on short-horned grasshoppers (Orthoptera: Acrididae) on the shortgrass steppe of Colorado. Journal of Arid
Environments.
177. Leo, V., M. Letnic, and R. P. Reading. In press. Responses of free-ranging red foxes (Vulpes vulpes) to the odour an
apex predator, the dingo (Canis Dingo). Oecologia.
178. Mudoch, J. D., H. S. Davie, M. Galbadrakh, and R. P. Reading. In press. Factors influencing red fox occupancy
probability in central Mongolia. Mammalian Biology.
BOOKS / JOURNAL SPECIAL ISSUES / DISSERTATION
1993
1. Reading, R. P. 1993. Toward an endangered species reintroduction paradigm: A case study of the black-footed ferret.
Ph.D. Dissertation. Yale University, New Haven, CT.
1994
2. Clark, T. W., R. P. Reading, and A. L. Clarke (eds.). 1994. Endangered Species Recovery: Finding the Lessons and
Improving the Process. Island Press, Washington, D.C.
3. Reading, R. P., D. Sumya, R. Samya, and N. Batsaikhan (eds.). 1994. Latin-Mongolian-Russian-English-German
Dictionary of the Vertebrate Species of Mongolia. Sukhbaatar Publishing House, Ulaanbaatar, Mongolia.
1995
4. Johnstad, M. and R. P. Reading. 1995. Protected Areas Rangers Training Manual. United Nations Development
Programme/Global Environment Facility & Ministry for Nature and the Environment’s Mongolia Biodiversity
Project. Mongolian Ministry for Nature and the Environment. Ulaanbaatar, Mongolia. (In Mongolian).
1996
5. Miller B. J., R. P. Reading, and S. Forrest. 1996. Prairie Night: Black-footed Ferrets and the Recovery of Endangered
Species. Smithsonian Press, Washington, D.C.
6. Johnstad, M., R. P. Reading and J. R. Wingard. 1996. Protected Areas Rangers Training Manual, 2nd Ed. United
Nations Development Programme/Global Environment Facility & Ministry for Nature and the Environment’s
Mongolia Biodiversity Project and German Technical Advisory Group (GTZ). Mongolian Ministry for Nature and
the Environment. Ulaanbaatar, Mongolia. (In Mongolian).
7. Contributing writer and photographer to: Finch, C. (ed.). 1996. Mongolia’s Wild Heritage: Biological Diversity,
Protected Areas, and Conservation in the Land of Chingis Khan. Mongolia Ministry for Nature and the
Environment, United Nations Development Programme-Global Environmental Facility, and World Wide Fund for
Nature, Avery Press, Boulder, CO.
2000
8. Reading, R. P. and B. Miller (eds.). 2000. Endangered Animals: A Reference Guide to Conflicting Issues. Greenwood
Press, Westport, CT.
2002
9. Wallace, R. L, T. W. Clark, and R. P. Reading. (eds.). 2002. An Interdisciplinary Approach to Endangered Species
Recovery: Concepts, Applications, Cases. Endangered Species Update Special Edition 19: 70-202.
10. Reading, R. P., D. Enkhbileg, and T. Galbaatar. (eds.). 2002. Ecology and Conservation of Wild Bactrian Camels,
Camelus bactrianus ferus. Mongolian Conservation Coalition & Admon Printing, Ulaanbaatar, Mongolia.
2006
11. Reading, R. P., D. Kenny, G. Wingard, B. Mandakh, and B. Steinhauer-Burkart. 2006. Ikh Nart Nature Reserve.
Nature-Guide No. 4, Mongolia. ECO Nature Edition Steinhauer-Burkart OHG, Oberaula, Germany. (English &
Mongolian editions).
2010
12. Miller, R., and R. P. Reading (eds). Saving the World’s Grasslands. Special Issue of Great Plains Research 20(1): 1160.
13. Reading, R. P., B. Miller, A. L. Masching, R. Edward, and M. K. Phillips. (eds.). 2010. Awakening spirits: Wolves in
the Southern Rockies. Fulcrum Publishing, Golden, Colorado, USA.
R. P. Reading, Ph.D. — Page 16
2011
14. Reading, R. P., D. Kenny, and B. Steinhauer-Burkart. 2011. Ikh Nart Nature Reserve, 2nd Edition. Nature-Guide No.
4, Mongolia. ECO Nature Edition Steinhauer-Burkart OHG, Oberaula, Germany.
NON-TECHNICAL PUBLICATIONS
1988
1. Reading, R. P. 1988. Pudu. Animal Keeper's Forum 15:43-45.
1990
2. Reading, R. P. 1990. Prairie dogs - A link in the chain. BLM Quarterly Steward. Winter:4.
3. Reading, R. P. 1990. A framework for reintroducing endangered species. Northern Rockies Conservation Cooperative
News 3:9-10.
1991
4. Reading, R. P. 1991. Saving the black-footed ferret. Montana Wildlife 15(2):4, 12.
5. Reading, R. P. 1991. Sociology of ferret recovery in Montana. Northern Rockies Conservation Cooperative News 4:13, 13.
1992
6. Reading, R. P. 1992. 'The greatest threat to the natural world.' Northern Rockies Conservation Cooperative News
5:18-19.
7. Reading, R. P., T. W. Clark, and S. R. Kellert. 1992. Knowledge and attitudes toward the Greater Yellowstone
Ecosystem. Northern Rockies Conservation Cooperative News 5:8-9.
8. Reading, R. P., P. Myroniuk, G. Backhouse, and T. W. Clark. 1992. Eastern barred bandicoot Reintroductions in
Victoria, Australia. Species. 19:29-31.
1994
9. Reading, R. P. 1994. Good news “Down Under”: Bandicoot numbers are up and NRCC’s international work
continues. Northern Rockies Conservation Cooperative News 7:16-17.
1995
10. Reading, R. P. 1995. A model for reintroducing endangered animals. Uyakhan Zambuutiv 2(1):22-27. (In
Mongolian).
11. Reading, R. P. 1995. Conserving Biodiversity in Mongolia. Northern Rockies Conservation Cooperative News 8: 47.
1997
12. Mix, H., R. P. Reading, and B. Lhagvasuren. 1997. Zum Status des Wilden Trameltieres (Camelus bactrianus ferus)
in der Mongolei. Zool. Gesellschaft fur Arten- und Populationsschutz 13(1):1-3. (In German)
13. Reading, R. P. 1997. Conservation of Mongolia’s wild Bactrian camels. The Zoo Review Summer 1997:10-13.
14. Reading, R. P. 1997. Mongolia’s biodiversity. Zoogoer 26(5):23-27.
1998
15. Radcliffe, C., V. Veer, and R. P. Reading. 1998. Twilight of the iguana. Zoo Review Summer:14-15.
16. Reading, R. P. 1998. China’s giant pandas. The Zoo Review Spring:3-6.
17. Reading, R. P., S. Amgalanbaatar, and H. Mix. 1998. Recent conservation activities for argali (Ovis ammon) in
Mongolia - Part 1. Caprinae August:1-3.
18. Reading, R. P., S. Mainka, L. Zhi, and G. Mills. 1998. Giant panda re-introduction feasibility. Re-introduction News
15:4-6.
1999
19. Miller, B., R. Nuñez, J. Terborgh, C. Lopéz, A. Miranda, and R. Reading. 1999. La importancia de los grandes
carnivoros en la conservación. Especies 8(3):4-8.
20. Reading, R. P., S. Amgalanbaatar, and H. Mix. 1999. Recent Conservation Activities for Argali (Ovis ammon) in
Mongolia, Part 2. Caprinae January: 1-4.
2000
21. Reading, R. P., H. Mix, and E. S Blumer. 2000. Bactrian camel survey. Endangered Species 1(3):34-37.
R. P. Reading, Ph.D. — Page 17
22. Reading, R. P., E. S Blumer, and H. Mix. 2000. Los camellos silvestres: En ayuda de otras especies in peligro.
Especies 9(4):25-27.
23. Reading, R. P. 2000. Excerpts from a field biologist’s journal: Argali sheep research in Mongolia. Zoo Review
Summer:10-13.
2001
24. Kenny, D. and R. P. Reading. 2001. The first argali. Zoo Review Spring: 6-7.
25. Kenny, D., R. P. Reading, S. Amgalanbaatar, and G. Wingard. 2001. The feasibility of immobilizing and radiocollaring free-ranging argali sheep (Ovis ammon) in Mongolia. Caprinae November: 1-3.
26. Reading, R. P. 2001. Erhaltung der Saiga in Kalmyien. ZGAP Mitteilungen 17 (2): 5 – 7. (in German).
27. Reading, R. P. and B. Miller. 2001. Denver Zoo’s Department of Conservation Biology celebrates its five-year
anniversary. Zoo Review Fall: 7-9.
2002
28. Reading, R. P. and A. Lushchekina. 2002. Saiga – Heading to extinction? Zoo Review Spring: 16-17.
2003
29. Amgalanbaatar, S. and R. P. Reading. 2003. Argali sheep conservation and research activities in Mongolia. In:
Mongolia: Biodiversity at a Crossroads, Biodiversity at a Crossroads. Biodiversity Briefings from Northern Asia
2(1): 14-16.
30. Johnstad, M. D. and R. P. Reading. 2003. Mongolia’s protected areas system. In: Mongolia: Biodiversity at a
Crossroads, Biodiversity at a Crossroads. Biodiversity Briefings from Northern Asia 2(1): 8-11.
31. McCain, L. and R. Reading. 2003. Happy birthday SPLT! Grasslands Gazette 5(2):3-4.
32. Reading, R. P., E. S Blumer, H. H. Mix, and B. Mijiddorj. 2003. Using satellite telemetry to track wild Bactrian
camels. CMS Bulletin 17: 12-13.
2004
33. McCain, L. and R. Reading. 2004. Prairie dogs as focal species. Grasslands Gazette 6(2):4-5.
34. Reading, R. 2004. How many prairie dogs are there? Grasslands Gazette 6(2):5-6.
2005
35. Murdoch, J. and R. Reading. 2005. Conserving the elusive Pallas’ cat. Zoo Review Spring: 16-17.
36. Reading, R. 2005. Is knowledge-provision enough? The relationship between values, attitudes and knowledge with
respect to wildlife conservation. Living Forests 10: 19-22. (In Chinese and English)
37. Reading, R., J. R. Wingard, and S. Amgalanbaatar. 2005. Trophy hunting of argali: An example of unsustainable
legal and illegal hunting and trade. P. 12, in: Going, going, gone…The illegal trade in wildlife in East and
Southeast Asia. The World Bank, Environment and Social Development East Asia and Pacific Region Discussion
Paper, The World Bank, Washington, D.C.
2006
38. Augeri, D. M., R. P. Reading, M. Fisher, D. J. Chivers, and J. A. Royle. 2006. Good estimates of density: A response
to Garshelis and McLellan. International Bear News 15 (2): 8-11.
39. Mix, H. M. and R. P. Reading. 2006. Usbekistans Wildschafe. Optimismus wird gebraucht. Und Geld. ZGAP
Mitteilungen 22 (1): 3 – 6. (in German).
40. Murdoch, J., T. Munkhzul, and R. Reading. 2006. Pallas’ cat (Otocolobus manul) ecology and conservation in the
semi-desert steppes of Mongolia. Cat News 45: 18-19.
41. Murdoch J. D., R. P. Reading, and T. Munkhzul. 2006. Conserving Pallas’ cats in Dornogobi, Mongolia. Pp. 15-18,
in: North American Felid Taxon Advisory Group (TAG) 2006 Annual Report. Edited by W. F. Swanson and N.
Flecthall, Felid TAG official website (www.felidtag.org).
42. Reading, R. P. and D. Kenny. 2006. Conservation in Mongolia: The Denver Zoo’s Mongolian Wildlife Conservation
Program. Communiqué May: 10-12.
2007
43. Reading, R. P. and D. Kenny. 2007. Denver Zoo’s Mongolian Wildlife Conservation. Connect Special Conservation
Issue: 24-27.
44. Reading, R. P., S. Amgalanbaatar, J. D. Murdoch, G. Wingard, and K. Kenny. 2007. Working to conserve
Mongolia’s Naural Heritage. Proceedings of the AZA National Conference, Sept 16 – 21, 2007, Philadelphia, PA.
AZA, Bethesda, MD or http://www.aza.org/AZAPublications/2007Proceedings/#07ac.
R. P. Reading, Ph.D. — Page 18
2008
45. McCain, L., R. P. Reading, and D. Stern. 2008. How natural resources personnel feel about black-tailed prairie dogs.
Current Conservation 2.2: 17-18.
46. Reading, R. P. 2008. Preface. Pp. v, in: Global Re-introduction Perspectives: Re-introduction case-studies from
around the globe, P.S. Soorae (Ed.). IUCN/SSC Re-introduction Specialist Group. Abu Dhabi, UAE.
2013
47. Reading, R. 2013. A few facts about the argali. P. 197, in: I heard a ram call my name. D. Duca. CreateSpace,
Amazon.com.
2013
48. Reading, R. 2014. Conserving vultures. Connect January: 14-15.
PRESENTATIONS, ABSTRACTS, POSTERS, & BOOK REVIEWS
1986
1. Schneider, C. W. and R. P. Reading. 1986. On the male conceptacles of two terete species of Gracilaria (Rhodophyta,
Gigartinales) from North Carolina. ASLO/PSA Meeting, Univ. of Rhode Island. (Abstract).
1991
2. Reading, R. P. 1991. Conservation of the endangered black-footed ferret. Chicago Zoological Society, Brookfield Zoo,
IL.
3. Reading, R. P. 1991. A holistic approach to black-footed ferret conservation. Royal Melbourne Zoological Park,
Victoria, Australia.
4. Reading, R. P., C. Schriver, S. Kellert, and J. Trent. 1991. Perceptions of local landowners toward prairie dogs and the
possible reintroduction of black-footed ferrets. Western Social Science Association Abstracts of Papers. 33rd
Annual Conference. Reno, NV. April 24-27, 1991. (Abstract).
1992
5. Reading, R. P. 1992. Reintroducing endangered species into the wild: A case study of the black-footed ferret. Univ. of
Hawaii, Honolulu.
1993
6. Reading, R. P. 1993. Lessons from recent (1988-1992) black-footed ferret (Mustela nigripes) recovery efforts.
Workshop on implementing endangered species policy: Reviewing the experience and learning the lessons.
University of Michigan, Ann Arbor.
7. Reading, R. P. 1993. Organizational aspects of black-footed ferret reintroduction and recovery efforts. Yale
University, New Haven, CT.
1994
8. Reading, R. P. 1994. Identifying and protecting critical habitat for threatened and endangered species: The American
experience. Arthur Rylah Institute, Department of Conservation and Natural Resources, Heidelberg, Victoria,
Australia.
9. Reading, R. P. 1994. Sport hunting and the conservation of biodiversity in Mongolia. Mongolian Sports Hunting
Conference, June 3-8, 1994. Mongol An Hunting Organization, Bulgan Hotel, Bulgan, Mongolia. (Abstract).
10. Reading, R. P. 1994. Biological aspects of Mongolian gazelle (Procapra gutturosa) conservation. Mongolian Gazelle
Seminar, September 9-10, 1994. Mongolia Biodiversity Project and the Society for the Conservation of the
Mongolian Gazelle, Ulaanbaatar, Mongolia. (Abstract).
11. Reading, R. P. 1994. Expanding Mongolia’s system of protected areas. Round Table Conference on Protected Areas
Guidance, October 6-7, 1994. German Technical Advisory Group’s (GTZ) Protected Areas and Development of
Buffer Zone Project and Mongolian National Park and Ecotourism Service, Ulaanbaatar, Mongolia. (Abstract).
12. Reading, R. P. 1994. Conservation of the Black-footed Ferret (Mustela nigripes) in North America. Institute for Zoo
Biology and Wildlife Research. Berlin, Germany.
1995
13. Reading, R. P. 1995. Conservation and management of the endangered species of Mongolia. Yale University School
of Forestry & Environmental Studies. New Haven, CT.
14. Reading, R. P. 1995. Applying a policy orientation to the conservation of endangered species and biodiversity. Yale
University School of Forestry & Environmental Studies. New Haven, CT.
R. P. Reading, Ph.D. — Page 19
15. Reading, R. P. 1995. The valuational aspects of prairie dog conservation. Society for Conservation Biology Annual
Meeting, 9 June 1995. Fort Collins, CO. (Abstract).
1996
16. Reading, R. P. 1996. Mongolia’s biological diversity. ZooMontana. Billings, MT.
17. Reading, R. P. 1996. Field conservation in developing countries: Lessons from Mongolia. Environmental,
Population, and Organismic Biology Department’s Fall Colloquium Series, University of Colorado, Boulder, CO.
18. Reading, R. P. 1996. Black-footed ferret (Mustela nigripes) conservation: An interdisciplinary assessment.
Department of Zoological Research’s Seminar Series, National Zoological Park, Smithsonian Institution,
Washington, D.C.
19. Reading, R. P. 1996. Conserving Mongolia’s biological heritage. Keynote Address, Jackson Hole Alliance for
responsible Planning Annual Meeting, Jackson, WY.
20. Reading, R. P. 1996. Development and management of Mongolia’s protected areas network. Department of Biology,
University of Denver, CO.
1997
21. Miller, B., R. List, R. P. Reading, T. Clark, A. Miranda, R. Medellin, O. Sanchez, and J. Bernal. 1997. International
Cooperation and the black-footed ferret. Proceedings of the International Theriological Congress, 10-12
September, 1997, Acapulco, Mexico. (Abstract).
22. Reading, R. P. 1997. Moving beyond biology in conservation: Lessons from the black-footed ferret recovery
program. Department of Wildlife and Fisheries Seminar Series, Texas A&M University, College Station, TX.
23. Reading, R. P. 1997. Interdisciplinary approaches to conservation: Lessons from Australia, Mongolia, and the United
States. Keynote Address, Thirteenth Annual Doctoral Research Conference, Yale School of Forestry &
Environmental Studies, New Haven, CT.
24. Reading, R. P. and B. J. Miller. 1997. Accurate evaluation: Why it’s important - Why it’s elusive. Society for
Conservation Biology Annual Conference, 6-9 June 1997, University of Victoria, British Columbia, Canada.
(Abstract).
25. Reading, R. P. and B. J. Miller. 1997. Zoos and a land ethic vision. Proceedings of the AZA National Conference, 1418 September, 1997, Albuquerque, NM. (Abstract).
26. Reading, R. P., B. J. Miller, and M. Stanley Price. 1997. Reintroducing animals into the wild: Lessons for giant
pandas. Giant Panda Reintroduction Workshop, 24-29 September, 1997, Woolong, China. (Abstract).
1998
27. Reading, R. P. 1998. Black-footed ferrets and conservation biology at the Denver Zoo. Animal Sciences and
Conservation Biology, Colorado State University.
28. Reading, R. P. 1998. Reintroduction theory and practice: Lessons for Bali mynahs. Bali Mynah Reintroduction
Workshop, 26-27 June, 1998, Lincoln Park Zoo, Chicago, IL.
29. Reading, R. P. 1998. Wild Bactrian Conservation. Large Herbivore Coordination Group, Second Meeting, 18-21
November, 1998, World Wide Fund for Nature, Lelystad, the Netherlands.
30. Reading, R. P., T. W. Clark, D. Mattson, and B. Miller. 1998. An interdisciplinary framework for effective carnivore
conservation. Carnivore Conservation Symposium, 20-21 November, 1998, Zoological Society of London,
London, UK. (Abstract).
1999
31. Blumer, E. S, Z. Namshir, T. Tuya, B. Mijiddorj, R. P. Reading, and H. Mix. 1999. Field studies of wild Bactrian
camels (Camelus bactrianus ferus) in Mongolia. Proceedings of the Annual Conference of the American
Association of Zoo Veterinarians, Columbus Ohio, Oct. 1999. C. Kirk (ed.). American Association of Zoo
Veterinarians, Media, PA. (Abstract).
32. Blumer, E. S, R. P. Reading, H. Mix, B. Lhagvasuren, and B. Mijiddorj. 1999. Development of a conservation
program for wild Bactrian camels in Mongolia. Proceedings of the 7th World Conference on Breeding Endangered
Species: Linking Zoo and Field Research to Advance Conservation, May 22-26, 1999. T. Roth (ed.). Cincinnati
Zoo, Cincinnati, OH. (Abstract).
33. Reading, R. P. 1999. Conservation of black-tailed prairie dogs: A keystone species of the Great Plains. Current
Events Club of Boulder, CO.
34. Reading, R. P. 1999. Przewalski's horse: The history and biology of an endangered species. Mongolian Studies 12:
105-106 (Book Review).
R. P. Reading, Ph.D. — Page 20
2000
35. Blumer, E. S, Z. Namshir, T. Tuya, B. Mijiddorj, R. P. Reading, and H. Mix. 2000. Veterinary aspects of wild
Bactrian camel (Camelus bactrianus ferus) conservation in Mongolia. Conservation of Wild Bactrian Camels and
the Gobi Desert Ecosystem Workshop, 3-5 April 2000. Mongolian Academy of Sciences & Denver Zoological
Foundation, Ulaanbaatar, Mongolia.
36. Lydia, R., J. Kirkpatrick, R. P. Reading, and E. S Blumer. 2000. Bactrian camel steroid fecal analysis. Conservation
of Wild Bactrian Camels and the Gobi Desert Ecosystem Workshop, 3-5 April 2000. Mongolian Academy of
Sciences & Denver Zoological Foundation, Ulaanbaatar, Mongolia.
37. Mix, H., R. P. Reading, E. S Blumer, and B. Lhagvasuren. 2000. Wild Bactrian camel status and distribution in
Mongolia. Conservation of Wild Bactrian Camels and the Gobi Desert Ecosystem Workshop, 3-5 April 2000.
Mongolian Academy of Sciences & Denver Zoological Foundation, Ulaanbaatar, Mongolia.
38. Reading, R. P. 2000. Toward a more holistic approach to reintroductions. Colorado Wildlife Society Annual Meeting,
January 19-21. Fort Collins, Colorado.
39. Reading, R. P. 2000. Interdisciplinary approaches to the reintroduction of threatened and endangered species.
Department of Zoology Seminar Series, February 4. University of Manitoba, Winnipeg, Manitoba.
40. Reading, R. P. 2000. A broader view of conservation: A review of Conservation in a Changing World. Zoo Biology
19:87-89. (Book Review).
41. Reading, R. P., H. M. Mix, E. S Blumer, S. Amgalanbaatar, T. Galbaatar, G. Wingard, T. Tserenbataa, M. D.
Johnstad, Z. Namshir, B. Lhagvasuren, and D. Kane. 2000. Pp. 118-123, in Wild ungulate conservation in
Mongolia. Central Asian Ecosystems – 2000, 5-7 Sept. 2000. Mongolian Academy of Sciences, Ulaanbaatar,
Mongolia. (Abstract).
42. Reading, R. P., H. M. Mix, E. S Blumer, S. Amgalanbaatar, T. Galbaatar, G. Wingard, T. Tserenbataa, M. D.
Johnstad, Z. Namshir, B. Lhagvasuren, and D. Kane. 2000. Wild ungulate conservation in Mongolia. MiniConference on Mongolian Paleoclimatology and Environmental Research, November 3-4, 2000, Lamont-Doherty
Earth Observatory and Columbia University, Palisades, NY.
43. Reading, R. P., H. Mix, E. Blumer, T. Galbaatar, and B. Lhagvasuren. 2000. Wild Bactrian camel conservation
program: Recent activities and future plans. Conservation of Wild Bactrian Camels and the Gobi Desert Ecosystem
Workshop, 3-5 April 2000. Mongolian Academy of Sciences & Denver Zoological Foundation, Ulaanbaatar,
Mongolia.
44. Tserenbataa, T., R. Ramey III, R. P. Reading, and O. Ryder. 2000. Genetic structure of argali sheep populations in
Mongolia. Workshop on Caprinae Taxonomy, Ankara, Turkey, 8-10 May 2000. Middle East Technical University,
Ankara. IUCN Caprinae Specialist Group, Gland, Switzerland. (Poster).
2001
45. Kenny, D., R. P. Reading, S. Amgalanbaatar, and G. Wingard. 2001. The feasibility of immobilizing and
radiocollaring free-ranging argali sheep (Ovis ammon) in Mongolia. Proceedings of the AZAWestern Regional
Conference, 22-25 March 2001, Cheyenne Mountain Zoo, Colorado Springs, CO. (Abstract).
46. Miller, B., R. P. Reading, and D. E. Wildt. 2001. What does a “Mission of Conservation” mean? Proceedings of the
AZAWestern Regional Conference, 22-25 March 2001, Cheyenne Mountain Zoo, Colorado Springs, CO.
(Abstract).
47. Reading, R. P. and B. Miller. 2001. Values, Learning, and Zoo Conservation. Proceedings of the AZAWestern
Regional Conference, 22-25 March 2001, Cheyenne Mountain Zoo, Colorado Springs, CO. (Abstract).
48. Mix, H. M., R. P. Reading, B. Lhagvasuren, and N. Batbayar. 2001. Abundance and distribution of antelope species
in Mongolia. Scientific Programme and Abstracts, 8th International Theriological Congress, 12-17 August 2001.
Sun City, South Africa (Abstract).
2002
49. Reading, R. P. 2002. The relationship between values, attitudes, and knowledge with respect to wildlife conservation.
44th Annual Western Social Science Association Conference, April 10-13, 2002, Albuquerque, New Mexico.
50. Amgalanbaatar, S., B. Lhagvasuren, and R. P. Reading. 2002. Caprinae hunting system and conservation in
Mongolia. III World Congress on Mountain Ungulates, Saragossa (Aragon), Spain, 10-15 June 2002. Saragossa
University, Aragon, Spain. (Abstract).
51. Amgalanbaatar, S., B. Lhagvasuren, and R. P. Reading. 2002. Current status of Caprinae in Mongolia. III World
Congress on Mountain Ungulates, Saragossa (Aragon), Spain, 10-15 June 2002. Saragossa University, Aragon,
Spain. (Abstract).
52. Reading, R. P. 2002. Uncommon ungulates of Mongolia: Ecology and conservation. June 15, 2002, Denver Museum
of Nature and Science, Denver, CO.
R. P. Reading, Ph.D. — Page 21
53. Reading, R. P. 2002. Conservation of Mongolia’s large ungulates. December 2, 2002, Loma Linda University, Loma
Linda, CA.
54. Reading, R. P., T. W. Clark, L. A. McCain, and B. Miller. 2002. Understanding and resolving the black-tailed prairie
dog conservation challenge. Symposium on People and Wildlife: Conflict or Co-existence, December 5-6, 2002,
Zoological Society of London and Wildlife Conservation Society, London, UK.
2003
55. Reading, R. P. 2003. Wildlife Restoration: Techniques for Habitat Analysis and Animal Monitoring by Michael L.
Morrison. Natural Areas Journal 23: 288-289 (Book Review).
56. Reading, R. P., L. A. McCain, T. W. Clark, and B. J. Miller. 2003. Understanding the controversy surrounding blacktailed prairie dog restoration efforts. International Field Seminar on Grassland Restoration, June 16-22, 2003,
Galichya Gora Nature Reserve, Russia.
57. Reading, R. P., L. A. McCain, T. W. Clark, and B. J. Miller. 2003. Understanding the controversy surrounding prairie
dog restoration efforts: Black-tailed prairie dog case study. Keynote address. 2003 Prairie Dog Summit, July 26-27,
2003, Denver, Colorado.
58. Reading, R. P., G. Wingard. 2003. Argali sheep ecology and conservation in Mongolia. Earthwatch Institute Annual
Conference, November 6-8, Cambridge, Massachusetts. (Poster).
2004
59. Murdoch, J., K. Ralls, B. Cypher, and R. Reading. 2004. Scent marking behavior by San Joaquin kit foxes (Vulpes
macrotis mutica). The Wildlife Society – Western Sectional Annual Conference, February 27-28, Rohnert Park,
CA.
60. Reading, R.P., S. Amgalanbaatar, G. Wingard, D. E. Kenny, and A. DeNicola. 2004. Ecology of Argali in Ikh Nartiin
Chuluu, Dornogobi Aimag. International workshop on Ecosystem research in the arid environments of Central
Asia: Results, challenges and perspectives, June 23-25. National University of Mongolia, Ulaanbaatar, Mongolia.
61. Reading, R. P., D. J. Bedunah, S. Amgalanbaatar. 2004. Conserving biodiversity on Mongolian rangelands:
Implications for protected area development and pastoral uses. Rangelands in Transition, 57th Annual Meeting of
the Society for Range Management, January 24 – 30, Salt Lake City, Utah.
62. Reading, R.P., E. S Blumer, H. Mix, and J. Adiya. 2004. Wild Bactrian Camel Conservation. International workshop
on Ecosystem research in the arid environments of Central Asia: Results, challenges and perspectives, June 23-25.
National University of Mongolia, Ulaanbaatar, Mongolia.
63. Reading, R.P., E. S Blumer, H. Mix, and J. Adiya. 2004. Wild Bactrian Camel Conservation in Mongolia. Second
international workshop on the re-introduction of the Przewalski’s horse, June 14-16. Takhin Tal, Great Gobi B
Strictly Protected Area, Mongolia.
64. Reading, R. P. and B. J. Miller. 2004. Attitudes and attitude change among zoo visitors: Findings and
recommendations. Catalysts for Conservation: A Direction for Zoos in the 21st Century. Zoological Society of
London, Wildlife Conservation Society, and Chester Zoological Gardens, February 19-20, London, U.K.
65. Reading, R. P., and G. Wingard. 2004. Argali sheep ecology and conservation in Mongolia. Earthwatch Institute
Annual Conference, November 4-6, Cambridge, Massachusetts. (Poster).
66. Wingard, G., R. P. Reading, S. Amgalanbaatar, and B. Mandakh. 2004. Seasonal food habits of argali (Ovis ammon)
and dietary overlap with domestic livestock in Mongolia. Rangelands in Transition, 57th Annual Meeting of the
Society for Range Management, January 24 – 30, Salt Lake City, Utah. (Poster).
2005
67. Reading, R. P., and G. Wingard. 2005. Argali sheep ecology and conservation in Mongolia. Earthwatch Institute
Annual Conference, November 3-5, Cambridge, Massachusetts. (Poster).
2006
68. Murdoch, J., T. Munkhzul, and R. Reading. 2006. Ecology and conservation of steppe carnivores in Dornogobi,
Mongolia. National Avian Research Centre, March 1, 2006, Abu Dhabi, United Arab Emirates.
69. Murdoch, J., Ts. Munkhzul, and R. P. Reading. 2006. Ecology and conservation of Pallas’ cats and other sympatric
small carnivores in the Northern Gobi. Felid TAG 2006 Mid-Year Meeting, April 7-9, 2006, Denver Zoo. Denver,
CO.
70. Murdoch, J., Ts. Munkhzul, and R. P. Reading. 2006. Ecology and conservation of steppe carnivores in Dornogobi,
Mongolia. Felid SSP Meetings, April 5-6, 2006, Denver Zoo, Denver, CO
71. Reading, R. P. 2006. The Role of Zoos in Conservation. April 8, 2006. University of Colorado, Boulder, CO.
72. Reading, R. P. 2006. Prairie dog empire: A saga of the shortgrass prairie. Great Plains Research 16(1): 103. (Book
Review).
R. P. Reading, Ph.D. — Page 22
2007
73. Kenny, D. and Reading, R. P. 2007. Veterinary Involvement at the Ikh Nart Nature Reserve, Mongolian. Working
Together, Answering the Call of the Wild. American Zoo and Aquarium Association Western Regional
Conference, March 26-31, Denver, Colorado.
74. Reading, R. P., and D. Kenny. 2007. Mongolian wildlife conservation program. Working Together, Answering the
Call of the Wild. American Zoo and Aquarium Association Western Regional Conference, March 26-31, Denver,
Colorado.
75. Reading, R. P , B. Nyambayar, T. Purevsuren, M. J. Willis, and D. Kenny. 2007. Conservation and Breeding Ecology
of Raptors in Ikh Nart Nature Reserve, Mongolia. Annual Conference of the Association of Zoos and Aquariums,
Sept. 17-21, Philadelphia, PA.
76. Reading, R. P., S. Amgalanbaatar, J. Murdoch, G. Wingard, and D. Kenny. 2007. Conserving Mongolia's Natural
Heritage. Annual Conference of the Association of Zoos and Aquariums, Sept. 17-21, Philadelphia, PA.
2008
77. Reading, R. P. 2008. Conserving Genghis Khan’s natural heritage: Wild Bactrian camels and other beasts from the
East. Alaska Zoo, July 1, Anchorage, AK.
78. Reading, R. P. 2008. Beasts from the East: Conserving wildlife in Mongolia’s Gobi Desert. Keynote address.
Earthwatch Europe Annual Conference, Oct. 16, Royal Geographic Society, London, UK.
79. Reading, R. P. 2008. Moving from no management to sustainable management: Developing a model nature reserve in
Mongolia. Keynote address. Sustainability Day, Oct. 22, University of Colorado-Denver, CO.
80. Reading, R. P. 2008. Conserving Genghis Khan’s natural heritage: Cinereous vultures, wild Bactrian camels and
other beasts from the East. Dover Land Trust, Nov. 9, Dover, MA.
2009
81. Bernal Stoopen, J., J. M. Packard, and R. P. Reading. 2009. Binational collaboration in endangered species recovery:
A framework for consensus analysis. Collaborative Conservation in Rapidly Changing Landscapes, 10th Biennial
Conference of Research on the Colorado Plateau, Society for Conservation Biology, US Geological Survey,
Northern Arizona University and others, Oct.5-8, Flagstaff, AZ. (Abstract, pp. 39-40).
82. Murdoch, J., T. Munkhzul, S. Buyandelger, R. P. Reading, and C. Sillero-Zubiri. 2009. Competition and ecological
separation between corsac and red foxes in Mongolia. Carnivores 2009: Carnivore conservation in a changing
world, Defenders of Wildlife, Denver, CO, Nov. 15-18. (Abstract, p. 184).
83. Reading, R. P. 2009. Beasts from the East: Conserving wildlife in Mongolia’s Gobi Desert. Anza Borrego
Foundation and Institute, Borrego Springs, CA, Jan. 31.
84. Reading, R. P. 2009. Beasts from the East: Conserving wildlife in Mongolia’s Gobi Desert. Denver Museum of
Nature and Sciences, Denver, CO, Nov. 17.
85. Reading, R. P., D. Bedunah, S. Amgalanbaatar, and J. Wigsten. 2009. Conserving Mongolia’s grasslands:
Challenges, opportunities, and lessons for America’s Great Plains. Grasslands of the World Conference, Grasslands
Foundation, Denver Zoological Foundation, and WWF, Lincoln, NE, April 1-2.
86. Wingard, G. J., R. B. Harris, D. H. Pletscher, D. J. Bedunah, B. Mandakh, S. Amgalanbaatar, and R. P. Reading.
2009. Argali food habits and dietary overlap with domestic livestock in Ikh Nart Nature Reserve, Mongolia. V
World Conference on Mountain Ungulates, Junta de Andalucia Consejería de Medio Ambiente & IUCN Caprinae
Specialist Group, Granada, Spain, 10-14 November 10-14.
2010
87. Amgalanbaatar, S. and R. Reading. 2010. Ikh Nart Nature Reserve conservation management and local participation.
International Conference on Mongolian wildlife: Conservation challenges and opportunities in 2010 and beyond.
Ulaanbaatar Hotel, Sept. 12 – 16, Ulaanbaatar, Mongolia. (Poster: In Mongolian).
88. Kaczensky, P., R. Reading, H. von Wehrden, and C. Walzer. 2010. Habitat use and movement patterns of wild
Bactrian camels in Mongolia. Camels in Asia and North Africa: Interdisciplinary workshop on their significance in
past and present, Austrian Academy of Sciences, 5 – 7 October 2010, Vienna, Austria.
89. Maran, T. and R. P. Reading. 2010. The role of zoos in conservation: Shift of paradigm. Nature Conservation Beyond
2010, Tallinn, Estonia, May 27-29 (Abstract).
90. Miller, S. and R. P. Reading. 2010. How best to measure abundance of truly active black-tailed prairie dog
populations using aerial survey techniques? WAFWA Grassland Initiative: Workshop on Methodologies for
Estimating Prairie Dog Abundance, Fort Collins, CO, January 25-28.
91. Reading, R. P. 2010. Conserving Wildlife from the Steppes of Colorado to the Steppes of Mongolia. Evergreen
Audubon Society Annual Meeting, Golden, CO, January 15.
R. P. Reading, Ph.D. — Page 23
92. Reading, R. P. 2010. Integrating social science considerations into conservation programs. Keynote Lecture. Nature
Conservation Beyond 2010, Tallinn, Estonia, May 27-29 (Abstract).
93. Reading, R. P. 2010. Beasts from the East: Conserving wildlife in Mongolia’s Gobi Desert. Curator’s Lecture Series.
Denver Museum of Nature and Science, Denver, CO. Aug. 4.
94. Reading, R. P. and J. Hoogland. 2010. Prairie dog ecology and conservation. Prairie Dog Coalition. REI Denver
Flagship Store, Denver, CO, February 27.
95. Willis, M. J., R. P. Reading, D. Kenny, J. Azua, T. Garrett, and P. Tsolmonjav. 2010. Movement patterns of breeding
cinereous vultures (Aegypius monachus) in Ikh Nart Nature Reserve, Mongolia. Asian raptors: Science and
conservation for present and future. 6th International Conference of Asian Raptors, Ulaanbaatar, Mongolia, June
23-27 (Abstract).
2011
96. Murdoch, J., T. Munkhzul, S. Buyandelger, and R. Reading. Ecology of corsac and red foxes and prospects for their
conservation in Mongolia. Biology Department Seminar Series, University of Vermont, April 2011.
97. Murdoch, J., T. Munkhzul, S. Buyandelger, and R. Reading. Ecology of the corsac fox and prospects for its
conservation in Mongolia. Wildlife Ecology Seminar Series, University of Maine – Orono, February 2011.
98. Reading, R.P. 2011. Establishing non-profit, non-governmental organizations in developing countries to support
conservation initiatives. Zoos and Aquariums Committing to Conservation 2011, Woodland Park Zoo, March 7 –
11, Seattle, WA.
99. Reading, R. P. 2011. Managing the genetic integrity of bison in the field: Lessons from Mongolia. Bison Ecological
Restoration, American Bison Society and Wildlife Conservation Society, March 23-25. Tulsa, OK.
100. Reading, R.P. 2011. The value of enrichment to reintroduction. International Conference on Environmental
Enrichment, August 14-19, Portland, OR (Abstract).
2012
101. Reading, R. P., J. Murdoch, D. E. Kenny, T. Purevsuren, J. Azua, Batdorj, T. Garrett, and M. J. Willis. 2012.
Factors influencing nesting success of Eurasian Black Vultures (Aegypius monachus) in Ikh Nart Nature Reserve,
Mongolia. Pp. 49, in: Proceedings of the 7th Symposium on Asian Raptors: Raptor Migration and Conservation in
Asia, 13-16 January 2012, A. Y. Choi, H. Y. Nam, G. C. Bing, and H. Y. Chae (eds.). Asian Raptor Research and
Conservation Network & National Park Research Institute & Ornithological Society of Korea, Ganghwa and
Cheorwon, Korea. (Abstract).
102. Reading, R.P. 2012. Basic Concept and Principles of Sustainable Wildlife Management. Workshop on Sustainable
Management of Central Asian Game Animals, 22-26 March 2012. German Federal Agency for Nature
Conservation, Insel Vilm International Academy for Nature Conservation, and GIZ (German Agency of
International Development). Isle Vilm, Germany.
103. Reading, R.P. 2012. Management Planning for Species and Wildlife Management Areas. Workshop on Sustainable
Management of Central Asian Game Animals, 22-26 March 2012. German Federal Agency for Nature
Conservation, Insel Vilm International Academy for Nature Conservation, and GIZ (German Agency of
International Development). Isle Vilm, Germany.
104. Reading, R.P. 2012. Monitoring Methods for Mountain Ungulates. Workshop on Sustainable Management of
Central Asian Game Animals, 22-26 March 2012. German Federal Agency for Nature Conservation, Insel Vilm
International Academy for Nature Conservation, and GIZ (German Agency of International Development). Isle
Vilm, Germany.
105. Reading, R. P. 2012. Interdisciplinary approaches to protected area management: A case study from Mongolia.
Keynote Address. Yale School of Forestry & Environmental Studies Doctoral Student Conference, 19 October
2012. Yale University, New Haven, CT.
106. Reading, R. P. 2012. Developing and running a conservation biology program at Denver Zoo. 1 August 2012,
University of Colorado Biology Club. University of Colorado Medical School, Aurora, CO.
107. Suuri, B., O. Ganbold, J. Azua, and R. P. Reading. 2012. Diet of Lesser Kestrels (Falco naumanni) in Ikh Nart
Nature Reserve, Mongolia. Pp. 127, in: Proceedings of the 7th Symposium on Asian Raptors: Raptor Migration
and Conservation in Asia, 13-16 January 2012, A. Y. Choi, H. Y. Nam, G. C. Bing, and H. Y. Chae (eds.). Asian
Raptor Research and Conservation Network & National Park Research Institute & Ornithological Society of
Korea, Ganghwa and Cheorwon, Korea. (Poster & Abstract).
R. P. Reading, Ph.D. — Page 24
2013
108. Elias, R., R. Reading, L. Bermudez, T. J. Weaver, J. R. Garcia, and M. T. Herbert. 2013. Ensuring a future for the
Lake Titicaca Frog. Zoos & Aquariums Committing to Conservation, Blank Park Zoo, Des Moines, IA, July 8 –
12. (Poster).
109. Reading, R. P. 2013. In the field with Denver Zoo: Conserving vultures and more. Boulder Audubon Society,
Boulder, CO, Oct. 22, 2013.
110. Reading, R. P., S. Amgalanbaatar, J. Azua, S. Buyandelger, D. Kenny, M. J. Willis, and G. Wingard. 2013.
Conserving Wildlife in Mongolia’s Gobi Desert. Anza-Borrego Foundation & Institute, Borrego Springs, CA,
Feb. 8, 2013.
111. Reading, R. P., A. Levine, and M. Herbert. 2013. Building a culture of conservation at Denver Zoo: Involving staff
in field conservation. Zoos & Aquariums Committing to Conservation, Blank Park Zoo, Des Moines, IA, July 8 –
12.
112. Reading, R. P., E. Stotz, and G. Wingard. 2013. Working with local stakeholders to develop a model protected area
in Mongolia. Zoos & Aquariums Committing to Conservation, Blank Park Zoo, Des Moines, IA, July 8 – 12.
113. Sidle, J., D. Augustine, D. Johnson, S. Miller, J. Cully, and R. Reading. 2013. A correction for overestimation bias
in estimates of black-tailed prairie dog (Cynomys ludovicianus) abundance based on aerial surveys of colony sites
in Colorado and Montana. 51st Annual Conference of the Montana Chapter of The Wildlife Society, Wildlife and
Change: The Influence of Climate, Politics, and Social Dynamics on Wildlife Conservation and Management,
Whitefish, MT, February 25 – March 1. Pp. 31 (Abstract).
2014
114. Amgalanbaatar, S. and R. P. Reading. 2014. Biology and ecology of argali sheep. Conference on Biological
Diversity, Conservation and Conditions of Ikh Nart Nature Reserve: 15 years of monitoring and research,
Mongolian State University of Education, Ulaanbaatar, Mongolia, 25 November 2015.
115. Batdorj, S. and R. P. Reading. 2014. Resource partitioning of 2 species of hedgehogs in Ikh Nart Nature Reserve.
Conference on Biological Diversity, Conservation and Conditions of Ikh Nart Nature Reserve: 15 years of
monitoring and research, Mongolian State University of Education, Ulaanbaatar, Mongolia, 25 November 2015.
116. Dorzhiev, Ts. Z.. S. Amgalanbaatar, S., and R. P. Reading. 2014. Structure of argali sheep Ovis ammon habitat in
Mongolia and its dynamics in the initial period of the 21st century. Conference on Biological Diversity,
Conservation and Conditions of Ikh Nart Nature Reserve: 15 years of monitoring and research, Mongolian State
University of Education, Ulaanbaatar, Mongolia, 25 November 2015.
117. Kenny, D. E., Y. J. Kim, H. Lee and R. P. Reading. 2014. Blood lead survey for migrating Eurasian Black Vultures
(Aegypius monachus) in the Republic of Korea. Book of Abstracts, International Raptor Conference, 8 th ARRCN
Symposium, Pune, India, 6-9 February. P. 58 (Abstract).
118. Kenny, D. E., R. P. Reading, G. Maude, P. Hancock, and R. Garbett. 2014. Elevated blood levels in avian
scavengers from Botswana, Africa. Botswana Wildlife Research Symposium, Maun, Botswana, February.
119. Reading, R. P., S. Amgalanbaatar, D. Kenny, and G. Wingard. 2014. Fifteen years of research and conservation in
Ikh Nart Nature Reserve. Biobeers, Wildlife Conservation Society and Steppe Forward Program, Ulaanbaatar,
Mongolia. 4 December.
120. Reading, R. P., S. Amgalanbaatar, D. Kenny, and G. Wingard. 2014. Developing a model conservation program: A
case study of Ikh Nart Nature Reserve. Conference on Biological Diversity, Conservation and Conditions of Ikh
Nart Nature Reserve: 15 years of monitoring and research, Mongolian State University of Education, Ulaanbaatar,
Mongolia, 25 November 2015.
121. Reagan, K., M. Byamba, and R. Reading. 2014. Bats and wind energy in Mongolia. Wind Wildlife Research
Meeting X, Broomfield, CO, USA, December (Poster).
122. Zapletal, M., B. Sodnompil, J. L. Atwood, J.D. Murdoch and R. P. Reading. 2014. A question of scale: Using
tracked paths to assess Daurian hedgehog habitat selection at multiple levels. Symposium on Animal Movement
and the Environment, North Carolina Museum of Natural Science, Raleigh, NC, 5-7 May 2014 (Poster).
2015
123. Altangerel T. Dursahinhan, A. T., R. P. Reading, B. Nyamsuren, K. J. Thomas, and S. L. Gardner. 2015 Occidian
(Apicomplexa: Eimeriidae) parasites from ibex (Capra sibirica) and domestic goat (Capra hircus) from Ikh Nart
Nature Rerseve, Dornogobi Province, Mongolia. 46th Annual Meeting of the Rocky Mountain Conference of
Parasitologists, Cedar Point Biological Station, University of Nebraska, Nebraska, 10-12 Sept. 2015.
124. Berenguel, R., R. Elias, M. Arteta, V. Ramos, T. Weaver, and R. Reading. 2015. Presencia de Bratrachochytrium
dendrobatidis en la rana gigante de Lago Titicaca (Telmatobius culeus). P. 122, in: Soler-Tovar, D. and P. E.
R. P. Reading, Ph.D. — Page 25
125.
126.
127.
128.
129.
130.
131.
132.
133.
134.
135.
Navas-Suarez (comp.), Salud y enfermedades de la via silvestre en Latinoamerica. Wildlife Disease Asosciation
– Latin American Section (WDA-LA). Asociación de Veterinarios de Vida Silvestre. Bogotá, Columbia. (Poster,
Abstract).
Bragin, N., D. Kenny, O. Ganbold, G. Wingard, H. Lee, M. J. Willis, J. Azua, T. Garrett, and R. P. Reading. 2015.
Long distance migration by Cinereous Vultures (Aegypius monachus). Zoos and Aquariums Committing to
Conservation Conference, 12-16 October 2015, Denver Zoo, Denver, CO. (Poster).
Maude, G., R. P. Reading, D. Kenny, B. Garbett, and P. Hancock. 2015. Addressing the rapid decline of vultures in
southern Africa. Zoos and Aquariums Committing to Conservation Conference, 12-16 October 2015, Denver
Zoo, Denver, CO.
Ramirez, L., B. Miller, S. Ramakrishan, G. Montoya, R. Larranaga, J. Zebrowski, and R. P. Reading. 2015.
Protecting ecological and cultural values through bison restoration in New Mexico. Zoos and Aquariums
Committing to Conservation Conference, 12-16 October 2015, Denver Zoo, Denver, CO.
Reading, R. P. 2015. Developing and running a conservation program at Denver Zoo. Center for Great Plains
Studies. Keynote, Greater Arkansas River Nature Association Annual Meeting, Salida, CO, 5 February 2015.
Reading, R. P. 2015. An innovative conservation partnership in New Mexico: The Rio Mora National Wildlife
Refuge. Keynote, Great Plains Fellows Luncheon. University of Nebraska, Lincoln, NE, 6 May 2015.
Reading, R. P. 2015. From “paper park” to model protected area: Transforming Ikh Nart Nature Reserve. Center for
Great Plains Studies. University of Nebraska, Lincoln, NE, 6 May 2015.
Reading, R. P. 2015. From “paper park” to model protected area: Conserving the Desert Grasslands of Mongolia.
South Dakota State University, Brookings, SD, 16 Sept. 2015.
Reading, R. P. 2015. From “paper park” to model protected area: Ikh Nart Nature Reserve, Mongolia. Department
of Zoology and Physiology, University of Wyoming, Laramie, WY, 2 October 2015
Reading, R. P. 2015. A One Health Approach to Protected Areas Management: Examples from Mongolia & the
USA. Third Global Risk Forum One Health Summit: Fostering interdisciplinary collaboration for global public
and animal health, 4 – 6 October 2015, Davos, Switzerland.
Wingard, G., R. P. Reading, D. Kenny, and S. Amgalanbaatar. 2015. Expansion of Ikh Nart Nature Reserve in
Dornogobi Province, Mongolia. Zoos and Aquariums Committing to Conservation Conference, 12-16 October
2015, Denver Zoo, Denver, CO. (Poster).
Zevallos, S., R. Elias, R. Berenguel, T. Weaver, and R. Reading. 2015. Presencia de Bratrachochytrium
dendrobatidis en Telmatobius spp. decomidades enla cuidad de Lima. P. 182, in: Soler-Tovar, D. and P. E.
Navas-Suarez (comp.), Salud y enfermedades de la via silvestre en Latinoamerica. Wildlife Disease Asosciation
– Latin American Section (WDA-LA). Asociación de Veterinarios de Vida Silvestre. Bogotá, Columbia. (Poster,
Abstract). (Abstract).