Origin of living Things and Natural Selection DBQ • Around 30 000 species of vertebrate animals with four limbs (tetrapods) are found today. They are thought to have evolved from a single amphibian species that crawled on to land around 400 million years ago during the Devonian geological period. The graph shows how the number of families of tetrapod species varied in each geological period since their first appearance. (a)State in which geological period reptiles first appeared. [1] (b) Calculate how many families of mammal were present five million years ago. [1] (c) Analyze the data in the graph to conclude whether mammals are more likely to have evolved from birds or from reptiles. [1] (d) Suggest how scientists collected the data about the numbers of families. [1] (e) The junction between the Cretaceous and Tertiary periods is called the K–T boundary. Using the data in the graph, discuss whether evolution at the K–T boundary fits the hypothesis of punctuated equilibrium or gradualism more closely. [8] (mammal/bird; at a constant or gradual rate; amphibian/reptile; Tertiary; rapid bursts; species; stability; Cretaceous) In punctuated equilibrium evolution occurs in _______________ interspersed by long periods of _____________ . The evidence shows very little change in number of __________________ families occurred during ____________________ period but large increase during the _________________ period. In gradualism evolution occurs _______________. The evidence shows the number of _____________________ families did not change much but a change in the number of families do not reflect what happens at the ____________________ level. The Origin of Organic Molecules and Life (A Scientific Perspective) Origin of Life and Organic Molecules (The Scientific Perspective) 1500 2000 2500 3000 3500 ARCHEAN Millions of years ago 1000 Paleozoic Bacteria Archae- Protista Plantae bacteria Fungi Animalia Colonization of land by animals Appearance of animals and land plants First multicellular organisms Oldest definite fossils of eukaryotes PRECAMBRIAN 500 Cenozoic Mesozoic PROTEROZOIC 0 Appearance of oxygen in atmosphere Oldest definite fossils of prokaryotes 4000 Molten-hot surface of earth becomes cooler 4500 Formation of earth The evolutionary tree of life can be documented with evidence. The Origin of Life on Earth is another story… The Origin of Life is a • Extra-terrestrial Origin – Was the original source of organic (carbon) materials comets & meteorites striking early Earth? – Testable • Spontaneous Abiotic Origin – Did life evolve spontaneously from inorganic molecules? – Testable Hypothesis Abiotic Origin of Life • Water vapor condensed (did not crystallize) - Formed oceans -Contained chemicals of life (primordial soup) • Most researchers believe life first appeared in shallow water or moist sediments. • However some now feel the first organisms may have developed on the sea floor due to the harsh conditions on the surface during that time. • This position was strengthened in the 1970's by the discovery of the deep-sea vent communities. • Hot water and minerals emitted from such deep-sea volcanic vents may have provided the energy and chemicals necessary for development of the early protobionts. Conditions on early Earth • Reducing atmosphere: this enhanced the joining of simple molecules to form more complex ones – water vapor (H2O), CO2, N2, NOx, H2, NH3, CH4, H2S – lots of available Hydrogen & its electron – no free oxygen • Energy sources - lightning, UV radiation, volcanic What’s missing from that atmosphere? A: Atmosphere was almost devoid of oxygen (which steals electrons away from other atoms). Origin of Organic Molecules • Abiotic synthesis – 1920 Oparin & Haldane propose reducing atmosphere hypothesis – 1953 Miller & Urey test hypothesis • formed organic compounds Stanley Miller University of Chicago Produced: -amino acids -hydrocarbons -nitrogen bases -other organics • The findings were significant for a number of reasons: Why was The early Earth could not have had oxygen present this (supported by other geological evidence) experiment • The early Earth had conditions that could have important??! allowed for the generation of molecules that would sustain life • Other variations of the experiment showed that molecules that could be used in DNA and RNA could be produced. • So this experiment supports the chemosynthetic theory, but because we weren't there to observe it, it is NOT proven. Spontaneous Origin of Life on Earth • Life is defined partly by two properties: accurate replication and metabolism. Neither property can exist without the other. Self–replicating molecules and a metabolism–like source of the building blocks must have appeared together. How did that happen? • Four processes needed for the spontaneous origin of life on Earth – The non-living synthesis of simple organic molecules – The assembly of these molecules into polymers – The origin of self replicating molecules that made inheritance possible – The packaging of these molecules into membranes with an internal chemistry different from their surroundings • The necessary conditions for life may have been met by protobionts, aggregates of abiotically produced molecules surrounded by a membrane or membrane–like structure. • Protobionts exhibit some of the properties associated with life, including simple reproduction and metabolism, as well as the maintenance of an internal chemical environment different from that of their surroundings. Key Events in Origin of Life • Origin of Cells (Protobionts) – lipid bubbles separate inside from outside w/metabolism & reproduction Laboratory experiments demonstrate that protobionts could have formed spontaneously from abiotically produced organic compounds. For example, small membrane–bounded droplets called liposomes can form when lipids or other organic molecules are added to water. • Origin of Genetics – RNA is likely first genetic material – multiple functions: encodes information (self-replicating), enzyme, regulatory molecule, transport molecule (tRNA, mRNA) • makes inheritance possible • makes natural selection & evolution possible • Origin of Eukaryotes – endosymbiosis First Eukaryotes ~2 bya • Development of internal membranes – create internal micro-environments – advantage: specialization = increase efficiency • natural selection! infolding of the plasma membrane plasma membrane endoplasmic reticulum (ER) nuclear envelope nucleus DNA cell wall Prokaryotic cell Prokaryotic ancestor of eukaryotic cells plasma membrane Eukaryotic cell 1 st Endosymbiosis • Evolution of eukaryotes – origin of mitochondria – engulfed aerobic bacteria, but did not digest them – mutually beneficial relationship • natural selection! internal membrane system aerobic bacterium mitochondrion Endosymbiosis Ancestral eukaryotic cell Eukaryotic cell with mitochondrion 2 nd Endosymbiosis • Evolution of eukaryotes Eukaryotic cell with mitochondrion – origin of chloroplasts – engulfed photosynthetic bacteria, but did not digest them – mutually beneficial relationship • natural selection! photosynthetic bacterium chloroplast Endosymbiosis Eukaryotic cell with chloroplast & mitochondrion mitochondrion Theory of Endosymbiosis • Evidence – structural • mitochondria & chloroplasts resemble bacterial structure – genetic • mitochondria & chloroplasts have their own circular DNA, like bacteria – functional • mitochondria & chloroplasts move freely within the cell • mitochondria & chloroplasts reproduce independently from the cell Lynn Margulis Cambrian explosion • Diversification of Animals The Day the Mesozoic Died – within 10–20 million years most of the major phyla of animals appear in fossil record 543 mya Extraterrestrial Life? “Hmmmm….. Life on other planets is there?” Seeding the Universe – ClEvR Graphic Organizer (on wiki) Scientific Explanation to Origin of Life • Panspermia -- some organic compounds may have reached Earth by way of meteorites and comets. – Organic compounds (e.g., amino acids) have been recovered from modern meteorites. • These extraterrestrial organic compounds may have contributed to the pool of molecules that formed early life (seeds of life are prevalent throughout the universe and life on Earth began by such seeds landing on Earth and propagating). • There is some evidence to suggest that bacteria may be able to survive for very long periods of time even in deep space (and may therefore be the underlying mechanism behind Panspermia). • Streptococcus mitus bacteria that had accidentally been taken to the moon on the Surveyor 3 spacecraft in 1967, could easily be revived after being taken back to earth three years later. Origin of Life Assignment The Origin of Life Challenge