Accessory Sex Glands The accessory glands are located along the pelvic portion of the urethra, with ducts which empty their secretions into the urethra. They include the vesicular glands, the prostate gland and the bulbourethral glands .They contribute greatly to the fluid volume of semen. In addition, their secretions are solution of buffers, nutrients ,and other substances needed to assure optimum motility and fertility of semen. Vesicular Glands The vesicular glands (sometimes called seminal vesicles) are a pair of lobular glands that are easily that identified because of their knobby appearance. They have been described as having the appearance of a "cluster of grapes." They are of similar length in the bull, boar ,and stallion (35 to 15 cm),but the width and thickness of the vesicular glands of the bull is approximately half that of the boar and stallion. They vesicular glands of the ram and buck are much smaller, being about 4 cm in length .The excretory ducts of the vesicular glands open near the bifurcation where the ampulla merge with the urethra. In bulls, they contribute well over half of the total fluid volume of semen ,and appear to make a substantial contribution in other species. Several organic compounds found in secretion of the vesicular glands are unique in that they are not found in substantial quantities elsewhere in the body. Two of these compounds, fructose and sorbitol ,are major sources of energy for bull and ram spermatozoa but are found in lower concentration in boar and stallion .Both phosphate and carbonate buffers are found in these secretions and are important in that they protect against shifts in the pH of semen .Such shifts in pH would be detrimental to spermatozoa. Prostate Gland The prostate is a single gland located around and along the urethra just posterior to the excretory ducts of the vesicular glands. A prostate body is visible in excised tracts and can be palpated in bulls and stallions.In rams, all of the prostate is embedded in urethra muscles as is part of this glandular tissue in bulls and boars. It makes a small contribution to the fluid volume of semen in most species studied .However ,some report that the contribution of the prostate gland is at least as substantial as that of the vesicular glands in boars. The prostate of the boar is larger than that of the bull. The secretions of the prostate are high in inorganic ions with sodium, chlorine, calcium, and magnesium all in solution. Bulbourethral Glands The bulbourethral (Cowpers)glands are a pair of glands located along the urethra near the point where it exits from the pelvis. They are about the size and shape of walnuts in bulls, but are much larger in boars. In bulls, they are embedded in the bulbospongiosum muscle .They contribute very little to the fluid volume of semen. In bulls, their secretions flush urine residue from the urethra before ejaculation. These secretions are seen as dribbling from the prepuce just before copulation. In boars, their secretions account for that portion of boar semen which coagulates. This strained from boar semen before it is used for artificial insemination. During natural service ,the white lumps formed by coagulation may prevent semen from flowing back through the cervix into the vagina of sows.