Wildlife Habitat Requirements OVERVIEW INSTRUCTOR: UNIT: Wildlife, Fisheries, and Ecology Management LESSON: Wildlife Habitat Requirements IMS REFERENCE: #8986-A TOPIC NOTES INTRODUCTION The management of wildlife is a complicated science. Wildlife management is to establish a healthy, diverse wild population of plants and animals. In nature, plants and animals form a symbiotic relationship. Plants provide food and cover for the animals. Animals control the growth of plants and return nutrients from the plants back to the soil. Rainfall fills rivers and lakes providing water for plants and animals. The vastness of the countryside provides space for plants and animals to live. The same vast expanse provides for the proper arrangement of food, cover, water, and space. Food, water, cover, space, and arrangement are all essential for the survival of wildlife. Human destruction of habitat breaks down these elements. Agriculture, roads, homes, and industry requires land. It is impossible to use land for these purposes without destroying wildlife habitat. Instructional Materials Service 1 www.myimsservices.com Wildlife Habitat Requirements Habitat management is the attention to the needs of wildlife. When properly conducted, it will provide the food, water, cover, space, and arrangement required by wildlife. Where humans need land for their own survival, they can also provide for the needs of all wildlife. FOOD When caring for livestock, a rancher provides food in the form of prepared feeds or pasture. To attract deer to a site, a hunter will usually feed corn. Homeowners will hang feeders on their porches to attract hummingbirds. Birdseed is available for homeowners to attract and feed birds. Where these are ways to feed wildlife, they are impractical or ineffective on a large scale. A game manager understands that there is a diversity of wildlife and each has its own food preferences. Plants (primary producers) take nutrients from the soil, energy from the sun, and sufficient water. Plants are the largest group in the food chain. Plants provide the base of the food chain. Herbivores (songbirds, squirrel, deer) will eat only plants and seeds. Carnivores (coyotes, mountain lions, snakes) prey on other animals. Omnivores (raccoons, skunks, opossums) will feed on vegetation and prey on other animals. All forms of wildlife serve a specific function in the ecosystem. Species have selective tastes even within their classification. Some herbivores prefer forbs while others prefer grasses. Some herbivores feed on succulents such as cactus while others feed on berries and seeds. Each species has their preference for food. Even plants are selective where they grow. Some plants only grow in an aquatic area while others thrive only in drier climates. A successful game manager will know what wildlife is present and the type of food the animals prefer. Plants adapt to certain climates and weather conditions. A game manager will know the plants native to the targeted area as well as those plants that can adapt to the area. Plant cover protects the soil from erosion and helps keep the soil moisture stable in a natural ecosystem. In that system, a balance of plants supports a community of animals. Those animals in turn provide a food supply to the predators. Predators prevent the herbivores from over populating and consuming all of the plants. This prevents soil erosion, maintains soil nutrients, and prevents loss of soil moisture. The soil provides plants and the plants feed the herbivores. This continuous cycle of plants and animals feeding on and being fed on by other members of the ecosystem is the food chain (see Figure 1). Figure 1. Food Chain Instructional Materials Service 2 www.myimsservices.com Wildlife Habitat Requirements Destruction of habitat breaks the cycle. This destruction kills plants, exposes the ground, and displaces animals. There is not enough vegetation for food or cover. The herbivores do not have enough food to produce healthy litters nor do they have the cover for natal activities. In the absence of enough wild prey, carnivores will prey on domestic animals. This causes the rancher to begin predator control activity. Predator control and a decreased food supply will reduce the number of predators. Invader plants replace the preferred plants of the area. Herbivores adapt to the new plant growth and begin to increase in numbers. Without predators to keep them in balance, they soon overpopulate. This again reduces the plant population. The animals look for new food sources generally cultivated crops. The farmer now has crop damage from herbivores. It is easy to understand the need for a balanced ecosystem. Each level of the food chain depends on the health and size of the other levels. A skilled wildlife manager understands this relationship and works to maintain a balanced ecosystem. A diversity of preferred plants promotes a strong population of plant eaters. This in turn provides a plentiful food supply for the predators. Managing food sources for wildlife can be simple or complex. The more elements in an ecosystem, the more complex it is to manage. An ecosystem with a wetland, timber, and grassland is more diverse and complex than a prairie. Within each type of system exists a food web (see Figure 2). In the food web, one stage of an animal’s life cycle may expose it as prey to animals that they may hunt when they are adults. Figure 2. Food Web Instructional Materials Service 3 www.myimsservices.com Wildlife Habitat Requirements A game manager should understand the concept of “good” or “bad” years, competitors, seasons, and available shelter. Plant nutrient value varies; rainfall affects plant growth, length of seasons affect plant growth. A rancher knows how many head of cattle are in a pasture. The rancher also knows how many cattle the grass can support and for how long. These are two major factors in managing the wildlife food supply. A game manager must know the population of all species in the area under management. It is also necessary to know the food source of each species. With these two items of information, a wildlife manager can effectively control populations to meet the food supply available or work to increase the volume of preferred plants. Food Requirements for Selected Game Species Managing food for wildlife requires an understanding and knowledge of the food requirements for various species. The following are selected game species often targeted by landowners and wildlife managers for development. Providing food for these species often have residual effects for other wildlife species. Consult other references for specific food needs of wildlife species not listed in this topic. White-tailed Deer Plants eaten by deer fall into one of three general categories: (1) browse or woody plants, (2) forbs or weeds (herbaceous broadleaf plants), and (3) grasses. The percentage these three types of food comprise in the diet of deer varies between seasons, vegetative areas, soils, individuals, and competition. Deer are very selective if they have a choice of foods and select foods because of their palatability, succulence, availability, and nutritive value. A good deer diet consists of three to seven pounds of dry matter per day and contains 14 to 18 percent protein. The leaves, twigs, and fruit of woody brush species, along with weeds and forbs, make up the bulk of the diet. Increased use of grass may indicate over population, poor conditions, or too much competition. A listing of preferred food items is at the end of this section. In addition to native plants, deer also like many agricultural crops such as peanuts, corn, peas, clover, oats, ryegrass, and wheat. In some areas, deer will eat the tender parts of cotton plants. Deer will eat available salt, but it is not essential. Mule Deer Browse is the staple item in the yearlong diet of the mule deer. Mule deer will eat succulent grasses and forbs when available. Pronghorn Antelope The diet of the pronghorn antelope consists primarily of forbs, browse, and a few grasses, varying with the seasons and the habitat types. Generally, their diet will be 65 percent forbs, 30 percent browse, and less than 10 percent grasses. When available and accessible, small grains such as wheat, barley, and oats may make up as high as 70 percent of the winter diet of pronghorns. Pronghorns are unique in that they can eat several forbs that are poisonous to domestic livestock. On poor condition and overgrazed ranges, and during periodic droughts, pronghorns have suffered losses from browsing woolly locoweed, creosote bush, and tarbush. Instructional Materials Service 4 www.myimsservices.com Wildlife Habitat Requirements Javelina Javelina is chiefly an herbivore and will feed on various cacti, especially prickly pear, mesquite beans, sotol, lechuguilla, and other succulent vegetation. Food studies have shown that prickly pear makes up 80 to 95 percent of the daily diet of javelinas. Javelinas also root for tubers, rhizomes, and bulbs. Most food studies found that javelina also eat certain animals such as rats, snakes, grubs, and carrion. Javelinas eat seasonal mast (acorns), grapes, hackberries, persimmons, and agarito berries when available. Squirrel The daily food requirement of squirrel is 0.2 pounds. Acorns are the main stay food of squirrel and makes up about 60 percent by weight of the total amount of food consumed. Squirrels eat acorns to some extent 12 months out of the year. Most hardwood trees must be at least 25 years old before acorn production begins. Antlers, bones, seeds, and fruit provide calcium when eaten by squirrels. Pecans and other nuts are taken in considerable quantities in spring, summer, and fall by squirrels. Pecans are preferred over all other foods. Squirrels begin to feed on pecans when blooms occur and continues through the nut stage. Quail The quail’s food consists of about 15 percent animal matter and 85 percent plant matter. The animal food is predominantly insects such as beetles, weevils, caterpillars, and crickets. Young birds feed almost exclusively on insects. The plant matter is composed mainly of seeds and fruits, although quail eat some green vegetation. Plants that produce a smooth, hard seed such as ragweed, croton, and partridge pea are important food plants. Quail use agricultural crops including corn, grain sorghum, legumes, soybeans, and other small grains. Pheasant Pheasants are basically seedeaters. Domestic grains (corn, grain sorghum, wheat, barley, soybeans) provide the bulk of a pheasant’s diet. In the spring and summer, insects provide a significant amount of the animal matter in the diet. Pheasant chicks subsist entirely on insects for the first five weeks of their life. Seeds such as ragweed, careless weed, smartweed, and crotons make up a significant amount of the winter diet. Gravel and calcium (in the form of snail shells) is an important part of the pheasants’ diets. This is especially important during the breeding and egg laying period. Turkey The turkey is opportunistic and uses a wide range of food. During the spring and summer, food items are green grasses and forbs, buds, flowers, seed, and insects. Poults’ diet during their first two weeks of life is 90 percent insects found primarily in openings and on the edge of pastures. Fall and winter foods include seed, green grass and forb shoots, acorns, and agriculture crops such as corn, grain sorghum, and wheat. Instructional Materials Service 5 www.myimsservices.com Wildlife Habitat Requirements Mourning Dove A mourning dove’s main diet consists of seeds from native grasses, cultivated grains, and forbs such as croton and sunflower. Insects are a minor item in their diet. Gravel or grit aid in digesting seed. The most popular feeding areas of mourning dove are recently harvested crop fields where waste grain and bare ground are present. WATER Water is basic to all living organisms, both domestic and wild. All plants and animals require water. How much, how often, and by what species is the problem faced by wildlife managers. Plants adapt to the amount of rainfall. Dense ground cover will conserve soil moisture for plant growth. Some animals get water from the dew or the plants they eat. Most animals have to travel to their water source. A wildlife manager sees the importance of plants in an ecosystem. Reduction of plants may be due to overgrazing or drought. It may be necessary to replant to strengthen a plant community. Native plants should be the first choice of the wildlife manager. They already have adapted to the type of weather conditions in the area. Water needs for animals vary, often within species. Turkeys can travel to get water. However, nesting hens need nearby water. The bobwhite quail will use no free water. Amphibians live, travel, find food, locate shelter, and often reproduce in water. Large animals will travel longer distances to water than will smaller animals. Smaller birds (quail and dove) can travel longer distances than larger birds (turkey and pheasant). Nesting or nurturing females need closer water than males or females not caring for young. Some animals (javelina) can meet their water needs from plants. Other animals (frogs, turtles) need a constant water source for survival. Water Sources Water sources for wildlife consumption include surface water, snow; dew, succulent plants, insects and other animal life; and metabolic water. Good wildlife water management involves developing and maintaining surface water. Rainfall and groundwater cares for the plants. In the absence of rain, there is little the wildlife manager can do to remedy the situation. Rivers, streams, creeks, ponds, and lakes provide water for most wild animals. Sometimes these sources are unavailable or drought affects the supply. It is often necessary for the wildlife manager to provide water for the animals. Understanding the needs of the species is the first step in providing water. Design and development of water sources depend on the needs of the target species. Design and Development Pond construction must take place on a suitable site. The site should contain a soil high in clay content. Clay reduces seepage and provides a stable embankment. Some pond construction provides for only seasonal water. Seasonally flooded areas do not require a Instructional Materials Service 6 www.myimsservices.com Wildlife Habitat Requirements soil with high clay content. Soils with low clay content should have a wide top and less slope. This provides a more stable structure. A pond should have an emergency spillway with a flat, grass bottom. A spillway will divert water away from the top of the embankment. It will also reduce erosion on site and downstream. Pond design will vary according to the type of facility. Four factors affect spillway design. They are watershed soil type, slope, vegetative cover, and the number of acres supplying the watershed. Constructed Permanent Impoundments There are two types of ponds that provide drinking water for wildlife. A pit-type pond is usually excavated on flat or level ground. Rainfall or a groundwater pump will fill this structure. Earthen dams are larger structures that collect water from a watershed. The depth of these structures should allow for normal evaporation loss and seepage. Design can include shallow areas and islands. These structures can enhance the food and habitat value of these structures for wildlife. Location is the critical element in construction of either type of pond. Proper placement ensures use by wildlife. Watering facilities should be no more than one-half mile apart. As a rule of thumb, there should be one pond for each 100 acres. This will benefit both large and small animals. Seasonal water structures meet the water needs of many wild species. One design of a shallow pond can provide a small area of permanent water. Another design will allow for larger areas of seasonal water. Both designs benefit the needs of waterfowl and other species that prefer water. Flat, poorly drained soils are excellent for shallow water impoundments. These areas are subject to shallow flooding during the winter. Areas that have deep seasonal flooding are hard to manage. They are also expensive to maintain and construct. Ideal sites are accessible by farm equipment to allow for food planting and maintenance of structures. Waterfowl prefer sites away from actively traveled roads. Design of any impoundment should include flooding and draining structures. Pipes and drains meet these needs. Drainage systems should allow for rapid removal of water during the growing season. Flooding begins in October and ends in mid-March. Drains should be open during the growing season to allow for food production. It also prevents damage to trees in the area. The preferred depth for these areas ranges from 6 to 18 inches of water. Natural Occurring Water Supplies Springs, perennial streams, and potholes are natural water supplies. They are important to all wildlife for drinking water and habitat. A wildlife manager should make resource decisions adjacent to and in these areas. Livestock exclusion, streamside management zones, and pest management are only three practices that can affect wildlife water. Their use can affect sediment load, water quality, improved nesting habitat, food supplies, and wildlife travel corridors. Instructional Materials Service 7 www.myimsservices.com Wildlife Habitat Requirements Water has one requirement for a good wildlife habitat. It must accompany good food and cover. Water is not enough to improve wildlife populations. For example, quail will not leave protective cover to get to water. Improved water quality and quantity can increase wildlife populations with good food and cover. Water Requirements for Selected Game Species A person must understand and know the water needs for many species of wildlife to manage it correctly. This section will identify water needs for selected game species. Water for one species will often benefit other species. Still, each species has its own needs. Consult other references for specific water needs of wildlife species not in this topic. White-tailed Deer White-tailed deer need available water for drinking and their needs are greatest during hot weather. Deer consume one-half to one gallon of water daily. In dry portions of the state, water may limit deer densities and distribution. Reliable available water from ponds, wells, springs, or creeks should supply the managed area. Water would be near cover and a water source should be located within one to three miles from the next available source. Mule Deer Mule deer should have water that is no further than three miles apart with a preferred range of one to three miles. Adequate cover should be adjacent to the water supply. Poorly spaced water supplies may concentrate the deer herds around available water. This can cause an overuse of forage species. Pronghorn Antelope Pronghorn antelope do not need a close water supply in their habitat but it is helpful. The pronghorn is able to conserve body water for a long time. Pronghorns will drink from livestock water troughs or ponds with livestock. Javelina Javelinas live in the Southwest United States. Much of this area contains plants known as succulents. Succulents (prickly pear cactus, lechuguilla) make up a large part of their diet. This food source makes javelinas less dependent on free water. Not all areas contain this type of succulent growth. When this occurs, javelinas will seek out streams, livestock ponds, and water troughs. Normally, absence of succulents either naturally or from brush control will cause javelina to leave the area. Squirrel Good squirrel habitat should have an available water source. Water is not a daily requirement for squirrel. Succulent plants will provide water in the absence of a surface water source. Ponds and creeks are common sources of water for squirrels. They even drink from tree hollows that collect water. Squirrels can survive for several weeks without free water. If water is absent for long periods, squirrels will leave the area. Instructional Materials Service 8 www.myimsservices.com Wildlife Habitat Requirements Quail Water requirements are generally met from the food consumed, dew, or surface water. Pheasant Water does not appear to be a critical habitat element for pheasant. Irrigated cropland and insects make up the pheasants’ habitat. This seems to meet the water needs of this species. Turkey Turkeys need water daily. They can get the water from food or from free water. Turkeys will nest and roost close to a water source. A nesting hen should have drinking water within 400 feet of her nest. Mourning Dove Mourning doves have a high need for water. Water is very important during the nesting season. The mourning dove is very mobile and can fly long distances to water. Because of this, water sources that meet the needs of other species will serve doves. COVER Cover is any material used by animals for protection from predators or shelter for nesting or natal activities. Animals can travel in it, use it for bedding, or seek protection from the weather in it. Animals can use cover to hide from predators or hide when stalking prey. There is a misconception that habitat is cover. While cover is a part of the animals’ habitat, habitat is not just cover. An animal can find cover in plants, rocks, caves, crevices, or ravines. Cover requirements of any wildlife species include several vegetation types. For example, a sage grouse requires open and bare areas for breeding seasons in addition to dense sage areas for feeding. A mule deer finds food in a sagebrush flat during winter but requires juniper groves for shelter during the same season. Factors that influence the amount of food also influence the amount of cover. The annual cycle of plant growth influences cover. For example, dense grassland is suitable for shelter in the spring but often loses its usefulness as the grass dies in the fall. Soil conditions, weather changes, and artificial conditions influence plant growth. Any change in the natural surroundings eventually has an effect upon the wildlife of the area. A single production operation, such as harvesting a grain crop, produces both favorable and unfavorable changes in the habitat. For example, harvesting corn destroys valuable cover for wildlife but at the same time waste kernels of grain are available for food. Agricultural practices used to promote wildlife cover include rotating crops, liming and fertilizing soils, planting winter cover crops, strip cropping, and applying other soil and water conservation practices. There are some innovative practices used to promote cover in urban areas. One is the use of multi-flora roses for living fences rather than ordinary Instructional Materials Service 9 www.myimsservices.com Wildlife Habitat Requirements wire fences. This presents cover for birds and small mammals while serving as a property barrier. Seeding ditches with wildlife plants will provide cover and food. Trails built under a highway provide cover and safe corridors for wildlife in urban areas. Cover Requirements for Selected Game Species A person must understand and know the cover needs for many species of wildlife to manage it correctly. This section will identify cover needs for selected game species. Cover for one species will often benefit for other species. Still, each species has its own needs. Consult other references for specific cover needs of wildlife species not in this topic. White-tailed Deer White-tailed deer require some type of woody cover. Cover may take many forms such as trees and brush in draws and along slopes, rough topography, or large, solid tracts of timber and brush. It may be motts or thickets in open areas, or along streams in prairie areas. Cover must be adequate to provide shelter from weather, predators, and man. A mixed cover type with a diverse plant community is more desirable than one with one species. The ideal percentages of cover would vary from 40 to 50 percent of an area and would depend on the brush species, deer densities, and hunting pressures. Does frequently hide fawns in areas of tall grass. The manager should protect good cover from overgrazing and wildfire. Habitat lacking in cover may need protection or there may be a need for a change in the native woody plants. Tall leafy plants can connect brushy areas and function as a temporary cover. Mule Deer Mule deer require shrubs, trees, and tall grasses for shelter and cover. The amount of woody cover needed by mule deer depends on the type and quality of the cover, topography, soils, and grazing pressure by domestic animals. Land with 25 to 35 percent of brush cover can maintain high populations of deer. It can cause grass production to increase. Pronghorn Antelope The pronghorn antelope is a Plains animal that prefers open, rolling landscapes. This habitat will allow for early detection of intruders. It requires no protective vegetative cover, but it will use draws and headers for protection from the elements during the winter months. Javelina Javelinas prefer the dry conditions in the southwestern United States. The most preferred habitat is dense thicket of prickly pear, scrub oak, guajillo, whitebrush, and mesquite. They take advantage of rough canyons and gullies for travelways and protection during bad weather. Javelinas are most active during the cooler hours of the day and at night. They rest in the denser thickets, under rock ledges, or shaded and cooler areas. Instructional Materials Service 10 www.myimsservices.com Wildlife Habitat Requirements Squirrel Fox squirrels adapt to a variety of forest habitats. Open upland forests with mixed trees support the largest populations. Mature oak and hickory woodlands provide the best habitat. The trees should be irregular shaped tracts of 5 to 20 acres. Strips of woodland connecting the tracts serve as corridors. Pine, elm, beech, pecan, maple, and other foodproducing trees will enhance the habitat. River valleys make up the western part of their range. This area supports pecans, walnuts, oaks, and other required trees. Gray squirrels prefer tree cover along streams. They prefer dense hammocks of live oak, willow oak, water oak, white oak, cypress, blackgum, and magnolia. Bottomland tracts with strong oak and pecan populations produce high populations of gray squirrels. Swamps and uplands are less productive. Squirrels prefer hollow trees for offspring to be born and for nurseries. They use leaf nests to some extent, especially in the spring and summer. Leaf nests also serve as refuge, resting, and feeding stations and occasionally as nurseries. Production is 2½ times more successful in tree cavities than in leaf nests. On an average, hardwood trees must be 40 years old to provide den cavities. An area must contain 5 to 10 percent mature hardwood trees to maintain good habitat. Gray squirrel den trees must have vines and heavy cover around the tree. Quail Quail are edge animals and prefer early successional stage vegetation. They inhabit idle fields, open woodland, crop fields with weedy edges, and pastures. Nesting, loafing, escape, and protective cover near dusting and feeding areas are necessary. Loafing or resting cover is used during the middle of the day between feeding periods. Low growing woody plants will provide loafing cover and should not be over 200 to 300 feet apart. Last year’s growth of herbaceous cover provides ideal nesting sites. Location of the nest is often near the edge of rough grass. This permits the hen to lead her chicks to thinner cover for feeding. Therefore, the edge of roads, fencerows, or firelanes is favorite spots selected for nesting. Shrub thickets provide escape cover and should be available on about every 15 acres. Roosting cover must provide warmth at the ground level and not restrict flight in any direction. Quail roost in a circle, heads looking outward with their tails toward the center. Pheasant Pheasant need several types of cover. Loafing cover, travel lanes, roosting cover, nesting cover, and winter cover meet these needs. Of these, nesting cover and winter cover are the most critical. Winter cover such as weedy fencerows and windbreaks provide protection from the elements. Turkey Mature timber provides good roosting sites and shelter from the weather. Large brush or timbered stands, dense grassy fields, and thickets offer good escape cover. The best habitat is diversity land use types with half in mature brush or timber with random openings. Interspersion of trees or brush and openings is the key. Good habitat will Instructional Materials Service 11 www.myimsservices.com Wildlife Habitat Requirements include a range of plants. It should have a variety of mast-producing hardwood trees. There should be a moderate to open understory cover with a variety of mast-producing plants such as sand shinoak. It should also have an abundant ground cover that provides green forage and seeds. Open areas are grasslands, fallow fields, roadways, power lines, and pipelines. Mourning Dove The mourning dove is one of the most adaptable of all species as indicated by its wide range. However, they do not inhabit forests, plains, vast open fields, marshes, or prairies. The broken topography of trees and openings brought about by agriculture seem to be more desirable. Trees, hedgerows, shrubs, and wires between utility poles furnish the dove nesting and roosting sites. Doves prefer to feed on the ground where the vegetation is sparse rather than on dense perennial grasses. SPACE Space requirements for wildlife allow room to move about, avoid predators, locate a mate, seek food and water, and rest. Home range is the term describing the native habitat of a species. Size of the home range will vary with the species. Still, quality and quantity of food, cover, and water will determine the amount of space required. Space requirements also depend on the size of the animal. In general, larger animals need more space but can live in a variety of habitats. Dietary preferences affect space requirements. For example, carnivores have larger territories and need more space than herbivores. Another factor is the species ability to tolerate crowded conditions. If the quality and quantity of a habitat is sufficient, crowding can occur with limited negative results. Wild animals have different space requirements based on the type of movement. Two types of space apply to wildlife movement. Internal movement will include migration and home range. Migration is the seasonal travel of a species from one habitat to another then back again. For example, geese leave their northern habitat and fly thousands of miles to their southern habitat. The following spring, they leave their winter habitat and return to their northern home range. Home range is the area where a species spends all or most of its time. Home range will vary in size with each species. A mountain lion may defend a 150-mile territory. On the other hand, a nestling wren may have a territory of only 150 square feet. There is also movement outside the population range. This movement includes dispersal of the target species. Dispersal will include emigration and immigration activities. Emigration is movement out of an area. Immigration is movement into an area. Habitat destruction by land development projects is the cause for movements of this type. Space Requirements for Selected Game Species A person must understand and know the space needs for many species of wildlife to manage it correctly. This section will identify space needs for selected game species. Space for one species will often benefit for other species. Still, each species has its own Instructional Materials Service 12 www.myimsservices.com Wildlife Habitat Requirements needs. Consult other references for specific space needs of wildlife species not in this topic. White-tailed Deer The area to maintain a deer herd link directly to the arrangement of food, cover, and water. Several hundred acres of food, cover, and water is needed for good quality habitat. Deer densities of one deer to three acres are very unusual and typically seasonal. Carrying capacity of the land for deer will depend on the potential of the site. Site potential depends on the soil type, vegetation, and current rainfall. Mule Deer Space is seldom a limiting factor in the natural range of mule deer. Quality of the habitat is often the limiting factor for mule deer numbers. There is approximately one mule deer per 100 acres in the Trans-Pecos area. The Panhandle will have a density of one animal per 300 acres. Good range should support about 10 to 15 deer per section. Pronghorn Antelope A free-ranging herd of pronghorns can cover great distances. The distance they travel will depend on the forage available. On fenced rangelands, three or four section pastures are a minimum size for a small herd of 8 to 12 pronghorns. Males with harems defend their territory during the breeding season. During this time, larger pastures provide less conflict between breeding males. The conflict is between breeding males defending their harem against bachelor males. Javelina Javelinas usually travel in herds of 5 to 15 animals. The herd has no particular leadership or hierarchy. The home range of a herd relates directly to the density of brush on that range. In South Texas, herd home ranges vary from 180 to 975 acres. Dense brushy cover and the abundance of prickly pear are the main criteria that determine the home range size. There is a herd boundary overlap of about 200 yards. When herds meet on this common ground a fight ensues with much chasing and loud popping of the tusks. Squirrel Fox squirrel may range up to 40 acres over a period of a year. The average range is about 10 acres with the minimum range being about two acres. The gray squirrel may range up to 20 acres over a period of a year. The average range is about eight acres with the minimum range being about two acres. Quail The home range of a covey is normally ¼-square mile and rarely exceeds a square mile. When cover and food are close together, quail generally have a smaller annual range. Research by F. Guthery shows the carrying capacity for quail can exceed three birds per acre. This is only with intense management and favorable weather conditions. Instructional Materials Service 13 www.myimsservices.com Wildlife Habitat Requirements Pheasant Pheasants will roam widely to fulfill its life needs. If adequate food and cover are available throughout the year, it will spend most of its time on a square mile or less. Turkey Annual range of a turkey is frequently 8 to 10 miles from winter roost sites to summer nesting sites (about 20,000 acres). Birds are very sensitive to human disturbance. People should not hunt or disturb them within a quarter mile of their roost or nest. Morning Dove Space is not a limiting factor for mourning dove except during the nesting season. ARRANGEMENT The welfare factors for wildlife are food, cover, and water. The space relationship between these factors and their abundance will determine what species can survive in the habitat. Arrangement unites the four needs of wildlife. Spatial heterogeneity is the diverse nature of the space available to the wildlife in a given area. That is, how much and what quality of food, cover, and water exist in the area. There is food, cover, and water available inside each area that may go unused. This is because they are outside the travel lanes of the animals. Arrangement brings food, cover, and water into the space used by the animals. An ecotone is the area where two edges or boundaries of adjoining habitats meet. The ecotone contains components of both habitats. Animals from each habitat move in and out of the ecotone. This creates an area with a greater diversity of wildlife species that each habitat individually. Species such as the cottontail rabbit, white-tailed deer, and the bobwhite quail are common inhabitants of ecotones, living on the edge and inside the adjoining habitat. Management to maximize the edge will increase the horizontal diversity. Horizontal diversity is the variety of plants along the ground. This will increase the number of species present. Establishing ecotones will not benefit all species. Some species such as migratory birds depend on large, unbroken tracts of habitat. Predation and parasitic infection of songbirds increases in ecotones. Creating an edge effect by bisecting the habitat into smaller units can reduce populations. Some individuals refer to edges as ecological traps for this reason. Two types of edge or ecotone systems are available as management practices. Inherent edge is a result of natural features. Soil, topography, or water creates the edge effect naturally. The result is ecotones that last longer and are relatively stable. It is possible to create induced edges artificially or naturally. They are the result of fire, wind, logging, planting, seeding, or other disturbances to the area. This edge is very unstable and may last as little as 2 years or no more than 10 years. Instructional Materials Service 14 www.myimsservices.com Wildlife Habitat Requirements Where induced edges are present, corridors between habitats will allow for movement between them without excessive predation. Corridors can be effective if they meet three criteria. They must protect known travel routes. They should follow areas of least resistance to movement. Finally, they should connect seeps, springs, and riparian areas. Proper arrangement of dissimilar habitats can support a greater diversity of wildlife. This mean food should be adjacent to cover. Nesting cover should be near brood feeding areas. Under these conditions, two or more different habitats preferred by a species can maximize energy loss and reduce exposure to predatory animals. The greater the diversity in habitat types, the better the opportunity for more abundant wildlife. It is also necessary to consider vertical diversity. Vertical diversity is the arrangement of plants from the ground upwards. At the lowest level are grasses and forbs. The understory is the next level and is composed of shrubs and young trees. The upper part of vertical diversity is the canopy. This is the cover produced by mature trees. Arrangement of these three units is related directly to the successional stage of development by the plant community. A diverse wild animal population is often associated with the number and density of vertical layers in the habitat. SUMMARY Food, cover, water, space, and arrangement are the five needs of wildlife. Each is critical to a healthy, diverse population of wild animals in a habitat. The successful wildlife manager is aware of the individual needs of the species and works to stimulate population growth by meeting their essential needs. Although all are important, close attention to arrangement can greatly enhance the use of the first four needs. This in turn will maximize the diversity and numbers of both game and non-game animals. ACKNOWLEDGEMENTS Dr. Joe Dettling, Associate Professor, Instructional Materials Service, Texas A&M University, College Station, Texas researched and developed this topic. Tamara Trail, Director of Education and Special Events, Texas Wildlife Association, and Dr. Terry Blankenship, Wildlife Biologist, Welder Wildlife Foundation, Sinton, Texas, reviewed this topic. Vickie Marriott, Office Software Associate, Instructional Materials Service, Texas A&M University, prepared the layout and design for this topic. Christine Stetter, Artist, Instructional Materials Service, Texas A&M University, prepared the illustrations for this topic. REFERENCES Instructional Materials Service 15 www.myimsservices.com Wildlife Habitat Requirements Demers, Chris, Leslie Hawkins, Alan Long, and Chuck McKelvy. 2001. “Providing Wildlife Cover.” SS-FOR-15, School of Forest Resources and Conservation, Florida Cooperative Extension Service, Institute of Food and Agricultural Sciences, University of Florida. [On-line]. Available: http://edis.ifas.ufl.edu/BODY_FR124 “Habitat Improvement for Wildlife.” 1998. Virginia Department of Forestry. [On-line]. Available: http://www.dof.state.va.us/mgt/wildlife/habimp.htm Miller, G. Tyler. 1996. Living in the Environment: Principles, Concepts, and Solutions. Wadsworth Publishing Company: Washington. “USDA Farm Bill Conservation Provisions – Wildlife Habitat Incentive Programs.” United States Department of Agriculture, Natural Resource Conservation Service. 2001. [On-line]. Available: http://www.nhq.nrcs.usda.gov/CCS/FB96OPA/WhipQ%26A.html “Wildlife and Recreation Management – State Study Guide.” 2001. Instructional Materials Service, Texas A&M University, College Station, Texas, Catalog Number 4400. GLOSSARY OF TERMS Competitors – Plants or animals that compete for the same food, space, territory. Food web – The interlocking food chains within an ecological community. Mast – Nuts (i.e., acorns, hickory, pecan). Metabolic – Related to the ongoing interrelated chemical actions that take place in living organisms that provide the energy and nutrients needed to sustain life. Natal – Relating to birth or to the time and place of birth. Poults – A young turkey. Riparian – Situated or taking place along or near the bank of a river. Section – Area of one square mile. Seep – Small pool or spring where liquid escapes from the ground. Succulent – With thick fleshy leaves and stems that can store water. Symbiotic – A close association of animals or plants of different species that is often, but not always, of mutual benefit. Watershed – The land area that drains into a particular area. ALL RIGHTS RESERVED Reproduction prohibited without written permission. Instructional Materials Service Texas A&M University 2588 TAMUS College Station, Texas 77843-2588 http://www-ims.tamu.edu 2002 Instructional Materials Service 16 www.myimsservices.com