Introduction – Landscape Ecology • Landscape Ecology: Study of landscape structure and processes. – Landscape: Heterogeneous area composed of several ecosystems. – Landscape Elements: Visually distinctive patches in an ecosystem. Human Land Use Practices Ecosystem simplification: Intensive Agriculture & Clean Farming Timber Extraction & Fragmentation Roads: Formation of Barriers in Landscapes Formation of Terrestrial “Islands” Habitat Fragmentation • Process of breaking contiguous unit into smaller pieces; area & distance components • Leads to: • Community & Ecosystem processes altered #patches Patch isolation Patch size Edge Habitat Fragmentation • area-sensitive species: species that require minimum patch size for daily life requirements • Edge effects: influence of factors from outside of a patch Increased Edge Habitat Increased Edge Habitat Edge Effects • Habitat surrounding a patch can: - change abiotic conditions; e.g., temp. - change biotic interactions, e.g., predation Example of nest predation = edge effect of approximately 50 m into forest patch Habitat Fragmentation • First-Order Effects: fragmentation leads to change in a species’ abundance and/or distribution Habitat Fragmentation • Higher-Order Effects: fragmentation indirectly leads to change in a species abundance and/or distribution via altered species interactions Habitat Fragmentation: SpeciesSpecific Sensitivity? • Rare species = more vulnerable • Wide ranging species = large-area requirements • Species with reduced mobility = more vulnerable • Species with low fecundity (related to rarity?) • Species with short life cycle (or multistage life cycle?) Habitat Fragmentation: SpeciesSpecific Sensitivity? • Interior-dependent species • Species vulnerable to human exploitation or disturbance • Specialist species? Implications of Changes in Scale Insects sampled at 10-m intervals for 100 m 45 40 35 30 25 Predator Prey 20 15 Pr ey Pr ed at or 10 5 0 Implications of Changes in Scale Insects sampled at 2000-m intervals for 20,000 m 45 40 35 30 25 Predator Prey 20 15 Pr ey Pr ed at or 10 5 0 Landscape Processes • Landscape structure influences processes such as the flow of energy, materials, and species between the ecosystem within a landscape. Landscape Structure and Dispersal of Small Mammals Habitat Patch Size and Isolation and Density of Butterfly Populations Introduction – Geographical Ecology • MacArthur defined geographical ecology as search for patterns of plant and animal life that can be put on a map. – Above level of landscape ecology Island Area and Species Richness Species-Area Relationship Island Biogeography • equilibrium model suggesting number of species occurring on an island represents a balance between immigration (in) and extinction (out) • Robert MacArthur & E.O. Wilson Equilibrium Model of Island Biogeography • Proposed rates of extinction on islands would be determined mainly by island size. – LG near islands will support highest number. – SM far islands will support lowest number. – SM near and LG far will support intermediate number.