Chapter 10: Photosynthesis Objectives The student is responsible for: 1. Knowing the definition of all the bold faced words in the chapter 2. Knowing the basic structure of chloroplasts 3. Understanding the electromagnetic spectrum and how it relates to PS 4. Flow of electrons in Photosystems I and II 5. Knowing the light reaction and chemiosmosis 6. Understanding the Calvin Cycle 7. Alternative mechanisms of carbon fixation: How have plants adapted to harsh environments? Figure 10.0 Sunbeams Figure 10.1 Photoautotrophs In Land Plants Euglena, a protist Multicellular Algae Cyanobacteria, prokaryotes Other bacteria Figure 10.2 Focusing in on the location of photosynthesis in a plant Figure 10.3 Tracking atoms through photosynthesis Figure 10.4 An overview of photosynthesis: cooperation of the light reactions and the Calvin cycle (Layer 3) Figure 10.5 The electromagnetic spectrum Figure 10.6 Why leaves are green: interaction of light with chloroplasts Figure 10.7 Determining an absorption spectrum Figure 10.8 Evidence that chloroplast pigments participate in photosynthesis: absorption and action spectra for photosynthesis in an alga Figure 10.9 Location and structure of chlorophyll molecules in plants Movie link practice Figure 10.10 Excitation of isolated chlorophyll by light Figure 10.11 How a photosystem harvests light Figure 10.12 How noncyclic electron flow during the light reactions generates ATP and NADPH (Layer 1) Figure 10.12 How noncyclic electron flow during the light reactions generates ATP and NADPH (Layer 2) An enzyme splits the water molecule releasing oxygen and electrons to fill the “electron hole” produced by light. Figure 10.12 How noncyclic electron flow during the light reactions generates ATP and NADPH (Layer 3) Figure 10.12 How noncyclic electron flow during the light reactions generates ATP and NADPH (Layer 4) Figure 10.12 How noncyclic electron flow during the light reactions generates ATP and NADPH (Layer 5) Figure 10.14 Cyclic electron flow Figure 10.15 Comparison of chemiosmosis in mitochondria and chloroplasts Figure 10.16 The light reactions and chemiosmosis: the organization of the thylakoid membrane Figure 10.17 The Calvin cycle (Layer 1) Figure 10.17 The Calvin cycle (Layer 2) Figure 10.17 The Calvin cycle (Layer 3) Figure 10.20 A review of photosynthesis Photorespiration Photorespiration is the uptake of carbon dioxide without the production of a sugar or ATP and consumes oxygen. Why would such a metabolic process that is so counterproductive, exist? Occurs in soybeans, rice, wheat. Photorespiration occurs on hot, dry days when the stomata would close. 1. CO2 levels drop in the leave; oxygen levels begin to increase. 2. Rubisco binds to oxygen and forms a two-carbon compound which leaves the chloroplast for the mitochondria and peroxisomes. 3. It is here the two-carbon compound is broken down 4. There are other ways to deal with stomata closing on hot dry days that are beneficial to plants, C4 plants and CAM plants. Figure 10.18 C4 leaf anatomy and the C4 pathway So here CO2 is kept high enough in the Bundle-sheath cells to keep rubisco binding to CO2. Photorespiration is minimized; sugar production is high. Sugar cane Figure 10.19 C4 and CAM (Crassulacean acid metabolism) photosynthesis compared