Chapter 9: Plant Organization Copyright © The McGraw-Hill Companies, Inc. Permission required for reproduction or display. 9-1 Plant Organs The flowering plants, or angiosperms, have characteristic organs and tissues. An organ is a structure that contains different types of tissues and performs one or more specific functions. 9-2 Organization of the plant body Shoot System Fig. 9.1 Vegetative organs are the leaf, stem and root The body of a plant has a root system and a shoot system. Root System 9-3 The Root System Consists of a main root (taproot) and many lateral roots, which absorb water and minerals from the soil for the plant. 9-4 Root systems Fig. 9.2 -Produce hormones -Perennial plants often store the products of photosynthesis in their roots. 9-5 Shoot System - Stems Fig. 9.1 A stem is the main axis of the plant along with its lateral branches. Transports water and minerals from roots to leaves, and products of photosynthesis in the other direction. A cylindrical stem can expand in girth as well as in length. 9-6 Shoot System - Leaves A leaf is a broad, thin organ (maximizes surface area) that carries on photosynthesis (some have other functions). Fig. 9.1 A leaf attaches to a stem at a node; an internode is the region beween nodes. 9-7 Blade Petiole Attaches to the node here 9-8 Monocot Versus Dicot Plants Flowering plants are divided into two groups depending on their number of cotyledons (seed leaves). Monocots (monocotyledons) have one cotyledon; dicots (dicotyledons) have two. Cotyledons provide nutrients for seedlings before true leaves begin photosynthesizing. 9-9 Monocot and dicot traits Fig. 9.3 9-10 Fig. 9.3 Monocots: Parallel veins (sugarcane, corn) Flowers have 3 or multiples of 3 (6,9,12, etc.) parts Dicots: Veins form a net pattern (oak tree) Flowers have 4 or 5 or multiples of 4 or 5 (8,10, etc.) parts 9-11 Dicot leaves From Page 150 9-12 Plant Tissues 1) Epidermal tissue – outer covering 2) Ground tissue – majority of plant tissue 3) Vascular tissue – transport 9-13 Epidermal Tissue Epidermal tissue forms the outer protective covering of a herbaceous (non-woody) plant. Exposed epidermal cells are covered with waxy cuticle to minimize water loss. 9-14 Root Hairs are Epidermal Tissue Fig. 9.4 Root hairs greatly increase the absorptive capacity of the root. 9-15 Stoma of leaves are part of the Fig. 9.4 epidermal tissue 9-16 Cork (an epidermal tissue) of an older stem Fig. 9.4 Cork is a component of bark. New cork cells are made by a meristem called cork cambium. As cork cells mature, they fill with suberin, a lipid that makes them waterproof and chemically inert. 9-17 Ground Tissue -forms the bulk of the plant. Fig. 9.5 -thin-walled and capable of photosynthesis when they contain chloroplasts. -have thicker walls for flexible support (celery strands). -are hollow, nonliving support cells with secondary walls. 9-18 Vascular Tissue Two types of vascular (transport) tissue: Xylem transports water and minerals from roots to leaves and contains two types of conducting cells: tracheids and vessel elements. Phloem transports organic nutrients from leaves to roots and has sieve-tube elements with companion cells; plasmodesmata extend between cells at sieve plates. 9-19 Xylem structure Xylem transports water and minerals from roots to leaves Contains two types of conducting cells: tracheids and vessel elements. Water Leaves Roots 9-20 Phloem structure Transports organic nutrients from leaves to roots Has sieve-tube elements with companion cells Plasmodesmata extend between cells at sieve plates. Organic nutrients Leaves Roots 9-21 Leaves (produce organic nutrients by photosynthesis) Phloem Carries Organic Nutrients Xylem Carries Water and Nutrients Roots (absorb water and minerals from the soil) 9-22 Organization of Roots Within a root are zones where cells are in various stages of differentiation. 9-23 Dicot root tip In the zone of maturation, mature cells are differentiated and epidermal cells have root hairs. In the zone of elongation, cells become longer as they specialize. The root apical meristem is in the zone of cell division; the root cap is a protective 9-24 covering for the root tip. Dicot root tip Fig. 9.8 Epidermis – single layer of thin-walled, rectangular cells; root hairs present in zone of maturation 9-25 Movement of materials into vascular cylinder of the root Endodermis – between cortex and vascular cylinder, single layer of endodermal cells bordered by the Casparian strip Fig. 9.8 Layer of impermeable lignin and suberin Cortex – thinwalled, loosely-packed parenchyma; starch granules store food regulates entrance of minerals into the vascular cylinder 9-26 In Dicot Roots Vascular Tissue is star-shaped; phloem in separate regions between arms of xylem Fig. 9.8 9-27 Branching and Taproot of dicots Fig. 9.9 Pericycle can start the development of branch roots. In some dicot plants, a primary root, or taproot, grows straight down and is the dominant root; it can be fleshy and stores food. 9-28 Organization of Monocot Roots Monocot roots have the same growth zones as a dicot root but they DO NOT undergo secondary growth (become woody). In a monocot root’s centrally located pith, ground tissue is surrounded by a vascular ring. See Fig. 9.10 9-29 Monocot root Fig. 9.11 Monocots have a large number of slender roots, which make up a fibrous root system, and are known as adventitious roots. Adventitious roots that emerge from the surface to help anchor the plant are called prop roots. -used for support 9-30 Fig. 9.11 Some plants are parasitic on other plants. Their stems have rootlike projections called haustoria that grow into the host plant and extract water and nutrients from the host. 9-31 Other Root Diversity Mycorrhizae - mutualistic association between roots (better water absorption) and fungi (receive sugars, etc.) Peas and other legumes have root nodules in which nitrogen-fixing bacteria live. 9-32 Organization of Stems Shoot tip Fig. 9.12 produces new cells that elongate and add length to the stem. 9-33 Primary tissues are new tissues formed each year from primary meristems right behind apical meristem. Meristem – Embryonic Tissue (undifferentiated) that develops into specialized tissue. 9-34 Fate of Primary Meristems Fig. 9.12 Protoderm gives rise to epidermis. Ground meristem produces parenchyma in the pith and cortex. Procambium produces primary xylem and primary phloem; later, vascular cambium occurs between xylem and phoem. 9-35 Herbaceous (nonwoody) Stems Mature herbaceous stems exhibit only primary growth (not secondary). The outermost tissue is the epidermis (not bark), which is covered by a waxy cuticle. 9-36 Herbaceous Dicot Stem -vascular bundles are in a distinct ring Fig. 9.13 9-37 Monocot stem -vascular bundles are scattered throughout Fig. 9.14 9-38 Woody Stems A woody plant has both primary and secondary tissues. Secondary tissues develop during the second and subsequent years of growth from lateral meristems (vascular cambium and cork cambium). Secondary growth, (annual growth) increases the girth of a plant. 9-39 Dicot stems Fig. 9.15 The secondary tissues produced by the vascular cambium, are called secondary xylem and secondary phloem, 9-40 Secondary growth in a dicot Fig. 9.15 stem Pith rays are composed of living parenchyma cells that allow materials to move laterally. Cork cambium replaces epidermis with cork cells impregnated with suberin. 9-41 Section of woody stem Fig. 9.15 Spring wood followed by summer wood makes up one year’s growth or annual ring. The bark of a tree contains cork, cork cambium, and phloem. A woody stem has three distinct areas: the pith, the wood, and the bark. 9-42 Annual Rings This tree had a pith date of 256 BC and an outer ring of about AD 1320, making this tree nearly 1,600 years old when it died (it's about 3 feet across)! (photo © H.D. Grissino-Mayer and R.K. Adams). 9-43 Tree trunk Fig. 9.16 9-44 Modified stems Fig. 9.17 9-45 Organization of Leaves Fig. 9.18 Helps prevent water loss Photosynthesis Photosynthesis and Increased area for gas exchange gas exchange 9-46 The body of the leaf is composed of mesophyll. Parenchyma cells of these mesophyll layers house chloroplasts. 9-47 Classification of leaves Fig. 9.19 9-48 Leaf diversity The leaves of a cactus are spines attached to a succulent stem. Climbing leaves are modified into tendrils. Fig. 9.20 The leaves of a few plants are specialized for catching insects.9-49