The Endocrine System Chapter 17 – Lecture Notes to accompany Anatomy and Physiology: From Science to Life textbook by Gail Jenkins, Christopher Kemnitz, Gerard Tortora 1 Chapter Overview 17.1 Endocrine System Overview 17.2 Hormone Secretion/Pos. & Neg. Feedback 17.3 Hypothalamus and Anterior Pituitary 17.4 Posterior Pituitary 17.5 Thyroid Gland 17.6 Parathyroid Glands 17.7 Adrenal Cortex 17.8 Pancreas 17.9 Gonads 17.10 Pineal Gland 2 Essential Terms hormone chemical mediator that helps maintain homeostasis target cell cell with a receptor that responds to the presence of a hormone 3 Concept 17.1 Endocrine System Overview 4 General Characteristics Glands that secrete chemical signals (hormones) into circulatory system Hormone characteristics Produced in small quantities Secreted into intercellular space Transported some distance in circulatory system Acts on target tissues elsewhere in body Regulate activities of body structures Ligands: more general term for chemical signals 5 Endocrine System Functions Metabolism and tissue maturation Ion regulation Water balance Immune system regulation Heart rate and blood pressure regulation Control of blood glucose and other nutrients Control of reproductive functions Uterine contractions and milk release 6 Endocrine Glands exocrine glands secrete products onto a surface endocrine glands secrete products into the body fluids hormones are carried to target tissues where activity is carried out pituitary, thyroid, parathyroid, adrenal, pineal Other hormone secreting structures hypothalamus, thymus, pancreas, ovaries, testes, kidneys, stomach, liver, small intestine, skin, heart, adipose tissue, placenta 7 Figure 17.1 8 Regulation of Activities: Comparison of Endocrine and Nervous Systems Endocrine: amplitude modulated signals. Amount of hormone determines strength of signal Onset within minutes of secretion of hormone Nervous: frequency-modulated signals. Frequency of action potentials produced by neurons determines strength of signal. Onset within milliseconds Two systems actually inseparable Nervous system secretes neurohormones into circulatory system Nervous system uses neurotransmitters and neuromodulators as ligands Some parts of endocrine system innervated directly by nervous system 9 Intercellular Chemical Signals Hormones: type of intercellular signal. Produced by cells of endocrine glands, enter circulatory system, and affect distant cells; e.g., estrogen Autocrine: released by cells and have a local effect on same cell type from which chemical signals released; e.g., prostaglandin Paracrine: released by cells and affect other cell types locally without being transported in blood; e.g., somatostatin Pheromones: secreted into environment and modify behavior and physiology; e.g., sex pheromones Neurohormone: produced by neurons and function like hormones; e.g., oxytocin Neurotransmitter or neuromodulator: produced by neurons and secreted into extracellular spaces by presynaptic nerve terminals; travels short distances; influences postsynaptic cells; e.g., acetylcholine. 10 Functional Classification of Intercellular Chemical Signals 11 Functional Classification of Intercellular Chemical Signals 12 Control of Secretion Rate Most hormones controlled by negative feedback systems Most hormones are not secreted at constant rate, but their secretion is regulated by three different methods 1. 2. 3. The action of a substance other than a hormone on an endocrine gland. Neural control of endocrine gland. Control of secretory activity of one endocrine gland by hormone or neurohormone secreted by another endocrine gland 13 Positive and Negative Feedback • • • • POSITIVE During the menstrual cycle, before ovulation, small amounts of estrogen are secreted from the ovary. Estrogen stimulates the release of gonadotropin-releasing hormone (GnRH) from the hypothalamus and luteinizing hormone (LH) from the anterior pituitary GnRH also stimulates the release of LH from the anterior pituitary LH causes the release of additional estrogen from the ovary. The GnRH and LH levels in the blood increase because of this positivefeedback effect. 14 Positive and Negative Feedback • • • NEGATIVE During the menstrual cycle, after ovulation, the ovary begins to secrete progesterone in response to LH. Progesterone inhibits the release of GnRH from the hypothalamus and LH from the anterior pituitary. Decreased GnRH release from the hypothalamus reduces LH secretion from the anterior pituitary. GnRH and LH levels in the blood decrease because of this negative-feedback effect. 15 Table 17.1 16 Concept 17.2 Hormone Secretion 17 Receptors hormones only affect target cells water soluble hormone receptors on outside surface and trigger response inside the cell lipid soluble hormone receptors on inside of cell and trigger response inside cell target cells generally have between 2,000 and 100,000 receptors for a given hormone 18 Chemical Classes of Hormones water soluble amino acid based lipid soluble steroids thyroid hormones nitric oxide transported in blood by transport proteins slow rate of loss in kidneys ready reserve of hormone in blood stream 19 Table 17.2 pt 1 20 Table 17.2 pt 2 21 Hormone Action variable depending on hormone and target cell various targets respond differently to same hormone some hormones activate synthetic or stimulatory processes others activate degradation or inhibitory processes 22 Figure 17.2 23 Figure 17.3 24 Hormone Interactions permissive effects one hormone allows the other to function synergistic effects one hormone intensifies the effects of the other antagonistic effects one hormone inhibits or reduces the effects of the other 25 Control of Hormone Secretion hormones secreted in bursts as stimulation increases bursts increase in frequency in absence of stimulation, bursts are minimal or inhibited regulated by neural signals chemical changes in blood other hormones 26 Concept 17.3 Hypothalamus & Pituitary Gland 27 Hypothalamus controls the activity of the pituitary gland major integrating link between the nervous and endocrine systems hormones that stimulate anterior pituitary are all either releasing hormones or inhibiting hormones 28 Releasing and Inhibiting Hormones Tropins or tropic hormones: hormones that regulate the hormone secretions of target endocrine tissues. All anterior pituitary hormones are tropins. Releasing hormones: GHRH. Growth hormone-releasing hormone. Causes the anterior pituitary to release growth hormone. TRH. Thyroid-releasing hormone. Causes the anterior pituitary to release thyroid-stimulating hormone (TSH). CRH. Corticotropin-releasing hormone. Causes anterior pituitary to produce adrenocorticotropic hormone. GnRH. Gonadotropin-releasing hormone. Causes anterior pituitary to produce FSH (follicle stimulating hormone) and LH (luteinizing hormone). PRH. Prolactin-releasing hormone. Causes the anterior pituitary to release prolactin. Inhibiting hormones: GHIH. Growth hormone-inhibiting hormone, somatostatin. Causes the anterior pituitary to decrease release of growth hormone. PIH. Prolactin-inhibiting hormone. Causes the anterior pituitary to decrease release of prolactin. 29 Figure 17.4 30 Pituitary Gland two lobes anterior lobe stimulated by tropic hormones from hypothalamus hypophyseal portal system posterior lobe neural tissue that releases hormones produced in the hypothalamus neurosecretory cells 31 Table 17.3 32 Figure 17.5 33 Figure 17.6 34 Figure 17.11 35 FSH & LH released by the anterior pituitary triggered by GnRH target tissue gonads FSH in females initiates development of ovarian follicles in males stimulates sperm production LH in females triggers ovulation in males triggers testosterone secretion 36 PRL released by the anterior pituitary trigger is PRH and PIH from hypothalamus initiates and maintains milk secretion and production by mammary glands in females in males can cause erectile dysfunction 37 ACTH secreted by anterior pituitary triggered by CRH also triggered by stress controls production and secretion of hormones called glucocorticoids cortisol from adrenal cortex cause negative feedback regulation of CRH and ACTH release 38 Figure 17.16 39 MSH secreted by anterior pituitary function unknown in humans presence of MSH receptors in brain suggests it may influence brain activity excessive CRH stimulates MSH release PIH inhibits MSH release 40 Table 17.4 pt 1 41 Table 17.4 pt 2 42 Concept 17.4 Posterior Pituitary 43 Posterior Pituitary AKA neurohypophysis store and release two hormones produced by hypothalamus ADH OT 44 Figure 17.4 45 OT oxytocin targets uterus and mammary glands during and after delivery uterus contracts milk ejection (“let down”) function in non-reproducing women and in men is unknown animal studies seem to indicate parental caretaking behavior toward offspring sexual pleasure during and after intercourse 46 ADH antidiuretic hormone decreases urine production kidneys return water to blood decreases sweating causes constriction of arterioles increases blood pressure AKA vasopressin 47 Figure 17.8 48 Table 17.5 49 Concept 17.5 Thyroid Gland 50 TSH follicular cells produce thyroxine (T4) triiodothyronine (T3) parafollicular cells produce calcitonin involved in calcium homeostasis brings calcium levels down when too high 51 Figure 17.11 52 Actions of Thyroid Hormones thyroxine and triiodothyronine regulate oxygen use and BMR 53 Abnormal Thyroid Conditions 54 Concept 17.6 Parathyroid 55 PTH parathyroid hormone major regulator of calcium, magnesium, and phosphate ions in blood PTH brings blood levels of calcium up when too low 56 Figure 17.13 57 Table 17.7 58 Causes and Symptoms of Hypersecretion and Hyposecretion of Parathyroid Hormone 59 Concept 17.7 Adrenal Gland 60 Adrenal Cortex divided into three zones each secretes its own hormone mineralocorticoids glucocorticoids androgens 61 Figure 17.15 62 Figure 17.16 63 Figure 17.5 64 Glucocorticoid Functions Protein breakdown Glucose formation Triglyceride breakdown Resistance to stress Anti-inflammatory effects Depression of immune responses 65 Adrenal Medulla modified sympathetic ganglion of ANS epinephrine norepinephrine 66 Figure 17.8 67 Figure 17.17a 68 Figure 17.17b 69 Figure 17.17c 70 Figure 17.17d 71 Symptoms of Hypersecretion and Hyposecretion of Adrenal Cortex Hormones 72 Concept 17.8 Pancreas 73 Blood Glucose Level Regulation glucagon released when blood glucose is low insulin released when blood glucose is high 74 Figure 17.18 75 Table 17.9 76 Concept 17.9 Gonads 77 FSH & LH estrogens progesterone testosterone androgens 78 Table 17.10 79 Concept 17.10 Pineal Gland 80 Pineal Gland hormone secreted is melatonin contributes to setting the body’s biological clock promotes sleepiness in small doses in animals with breeding seasons, melatonin inhibits reproductive functions outside the season 81 End Chapter 17 82