Classification (Taxonomy) -naming and classification of species Topics: • History of classification • Current Scheme • Five/six kingdom system • Kingdoms overview Introduction • To trace phylogeny (the evolutionary history of life) biologists use evidence from paleontology, molecular data, comparative anatomy, etc. • Tracing phylogeny is one of the main goals of systematics, (the study of biological diversity in an evolutionary context.) • Systematics includes taxonomy, which is the naming and classsification of species and groups of species. As Darwin correctly predicted, “our classifications will come to be, as far as they can be so made, genealogies.” Aristotle (384 BC to 322 BC) His classification system was based on structural differences that were seen. Carolus Linnaeus (1707-1778) Swedish Botantist based his classification on characteristics of organisms that were similar. Developed the two word system used to identify species: binomial nomenclature. Taxonomy employs a hierarchical system of classification • First formally proposed system. • Systema naturae in the 18th century, has two main characteristics. • Each species has a two-part name. • Species are organized hierarchically into broader and Carolus Linnaeus (1707-1778) Binomial Nomenclature. • Each species is assigned a two-part latinized name, a binomial. • The first part, the genus, is the closest group to which a species belongs. • The second part, the specific epithet, refers to one species within each genus. • The first letter of the genus is capitalized and both names are Hierarchical Classification Taxons • Groups species into broader taxonomic categories. • Species that appear to be closely related are grouped into the same genus. • For example, the leopard, Panthera pardus, belongs to a genus that includes the African lion (Panthera leo) and the tiger (Panthera tigris). • Biology’s taxonomic scheme formalizes our tendency to group related objects. From Kingdom to Species Timeline of Classification 1. 384 – 322 B.C. Aristotle 2 Kingdom Broad Classification 2. 1735 - Carl Linnaeus 2 Kingdom Multi-divisional Classification Kingdom, Phylum, Class, Order, Family Genus, Species 3. Evolutionary Classification – (After Darwin) Group By lines of Evolutionary Descent 4. 5 Kingdom System – 1950s 5. 6 Kingdom System – 1990s 6. 3 Domain System – 1990s Phylogenetic Trees reflect the hierarchical classification of many taxonomic groups. Phylogeny is determined by a variety of evidence including fossils, molecular data, anatomy, and other features. Modern phylogenetic systematics is based on cladistic analysis • Most systematists use cladistic analysis, developed by a German entomologist Willi Hennig to analyze the data • A phylogenetic diagram or cladogram is constructed from a series of dichotomies. Systematics-the study of biological diversity in an evolutionary context. • These dichotomous branching diagrams can include more taxa. • The sequence of branching symbolizes historical chronology. • The last ancestor common to both the cat and dog families lived longer ago than the last common ancestor shared by leopards and domestic cats. branch or clade Can not always base clade on appearance. • It is especially important to distinguish similarities that are based on shared ancestry or homology from those that are based on convergent evolution or analogy. • These two desert plants are not closely related but owe their resemblance to analogous adaptations (convergent evolution). • The presence good distinguish the mammals tetrapods. of hair is a character to clade of from other Analyzing the taxonomic distribution of homologies enables us to identify the sequence in which derived characters evolved during vertebrate phylogeny. A cladogram presents the chronological sequence of branching during the evolutionary history of a set of organisms. Systematics is the study of the diversity of organisms using information from cellular to population levels. 1. Classification reflects phylogeny; one goal of systematics is to create phylogenetic trees. 2. Phylogeny is the evolutionary history of a group of organisms. 3. A phylogenetic tree indicates common ancestors and lines of descent. 4. A primitive character is a trait that is present in a common ancestor and all members of a group. 5. A derived character is present only in a specific line of descent. 6. Different lineages diverging from a common ancestor may have different derived characters. Molecular clocks may keep track of evolutionary time • The timing of evolutionary events has rested primarily on the fossil record. • Recently, molecular clocks have been applied to place the origin of taxonomic groups in time. • Molecular clocks are based on the observation that some regions of genomes evolve at constant rates. • For these regions, the number of nucleotide and amino acid substitutions between two lineages is proportional to the time that has elapsed since they branched. • For example, the homologous proteins of bats and dolphins are much more alike than are those of sharks and tuna. • This is consistent with the fossil evidence that sharks and tuna have been on separate evolutionary paths much longer than bats and dolphins. • In this case, molecular divergence has kept better track of time than have changes in morphology. Six Kingdoms of Life Eukarya Archaea Archaebacteria Bacteria EuBacteria Plantae Protista Fungi Animalia Six Kingdoms of Life •Archaebacteria •Eubacteria •All Single celled prokaryotes •Anaerobes and aerobes •One circular chromosome •Heterotrophic and autotrophic Six Kingdoms of Life Protista •Widest variety of organisms •All eukaryotes •Single and multicellular •Heterotrophic and autotrophic Six Kingdoms of Life Fungi •All Heterotrophic •Eukaryotic •Single and multicellular •Important as decomposers •Cell walls made of chitin •4 divisions Six Kingdoms of Life Plantae •All Autotrophic •Eukaryotic •All multicellular •Cell walls made of cellulose •10 divisions Six Kingdoms of Life Animalia •All heterotrophic •All Eukaryotic •All multicellular •35 phyla Mrs. Payne