Announcements Exams will be ready on Thursday. I have a sudden deadline I need to meet by the end of this week and have not been able to finish going through them. Reading is available from web site. Read it closely by next Thursday and be prepared to discuss in class!!! Urban Forestry project, we'll start on campus. Meet in Rm. 202, as usual, next week. We'll try some techniques for taking quick digital specimens. Bring your cameras. POPULATIONS ‘metapopulations’ genetic diversity is organized on the landscape by different selection pressures SPECIES Atomic particle – evolutionary unit Extinction and birth Binomial taxonomic name Barriers to gene flow External or internal Neutral or selective Gradual or immediate SPECIES Allopatric speciation Mutations accumulation, limited gene flow Sympatric speciation Thought to be very difficult Assortative mating SPECIES Reproductive isolation mechanisms Pollinator fidelity and service Prepollination or post (prezygotic or postzygotic) SPECIES Prepollination – behavior, morphology, selfpollination across strong environmental gradients. Postpollination (prezygotic) – pollen vigor, stigmas as selective agents, MHC complex. Postpollination (postzygotic) – inviability, abortion, infertility SPECIES “Hybridization” is a dirty word amongst most taxonomists despite the fact that it has been shown to be advantageous in some circumstances. “Good” or “bad” species Many plants remain interfertile after ‘speciation’ SPECIES The abominable mystery of flowering plant diversity Latest work on fossil flowers has been quite amazing. SUPER-SPECIES, SPECIES FLOCK or SYNGAMEON “Ring” species Network of closely related and inter-fertile species Fuzzy species concept. COMMUNITY Shifting ‘selective’ landscapes “Mast fruiting” in trees EVOLUTION and ADAPTATION It is an integration of numerous processes over long periods of time, during which sudden and dramatic events occur and change the selective environment.