Chapter 26c Reproduction and Development Pregnancy Prevention • Contraceptives (birth control methods) • Abstinence • Total abstinence • Abstinence during periods of female fertility • Interventional methods • • • • Barriers Surgery Blocking implantation Hormone treatments to reduce gamete production Pregnancy Prevention Table 26-5 Fertilization • Sperm capacitation occurs in vagina • Sperm swim “upstream” • Sperm reach oocyte in Fallopian tube • Acrosomal reaction digestive enzymes • Zona pellucida and cell junctions dissolve • Membranes fuse and sperm nucleu enters • Cortical reaction blocks polyspermy • Nuclear fusion creates a diploid cell • 1 sperm + 1 oocyte 1 zygote Fertilization Figure 26-16a Acrosome Reaction Allows Sperm to Reach Egg Figure 26-16b Sperm and Egg Fuse to Form a Zygote First polar body Egg Sperm nucleus Sperm and egg plasma membranes fuse. Sperm nucleus moves into cytoplasm of egg. Figure 26-17 (1 of 4, 2 of 4) Oocyte Completes Meiosis and Nuclei Fuse Sperm nucleus Egg Oocyte nucleus completes meiotic division. Second polar body Sperm and egg nuclei fuse to form zygote nucleus. Figure 26-17 (3 of 4, 4 of 4) Early Development • Rapid cell divisions follow fertilzation • Blastocyst • Hollow ball of ~ 100 cells • Arrives in uterus on day 4 or 5 • Implantation • Blastocyst invades the endometrium Early Development: Ovulation to Implantation 3 Days 2-4: Cell division takes place. 2 Day 1: Fertilization Zygote 4 Day 4-5: Blastocyst reaches uterus. Fallopian tube Egg 1 Ovulation Inner cell mass Blastocyst Ovary Uterus 5 Days 5-9: Blastocyst implants. Figure 26-18 Early Development: Ovulation Fallopian tube Egg 1 Ovulation Ovary Uterus Figure 26-18, step 1 Early Development: Fertilization 2 Day 1: Fertilization Zygote Fallopian tube Egg 1 Ovulation Ovary Uterus Figure 26-18, steps 1–2 Early Development: Rapid Cell Divisions 3 Days 2-4: Cell division takes place. 2 Day 1: Fertilization Zygote Fallopian tube Egg 1 Ovulation Ovary Uterus Figure 26-18, steps 1–3 Early Development: Blastocyst Reaches Uterus 3 Days 2-4: Cell division takes place. 2 Day 1: Fertilization Zygote Inner cell mass Fallopian tube Egg 1 Ovulation 4 Day 4-5: Blastocyst reaches uterus. Blastocyst Ovary Uterus Figure 26-18, steps 1–4 Early Development: Implantation of Blastocyst 3 Days 2-4: Cell division takes place. 2 Day 1: Fertilization Zygote 4 Day 4-5: Blastocyst reaches uterus. Fallopian tube Egg 1 Ovulation Inner cell mass Blastocyst Ovary Uterus 5 Days 5-9: Blastocyst implants. Figure 26-18, steps 1–5 Placenta and Continuing Development • The chorion is a tissue layer that encloses the developing embryo • The amnion is a tissue layer that secretes amniotic fluid, suspending the embryo • Chorionic villi form a close connection with blood vessels of the endometrium • The placenta continues to grow during pregnancy The Placenta Figure 26-19a The Placenta: Microscopic Anatomy Figure 26-19b The Placenta Makes Numerous Hormones • Human chorionic gonadotropin (hCG) • “Rescue” of corpus luteum • Basis of pregnancy tests • Human placental lactogen (hPL) • Breast development, metabolic effects • Estrogen and progesterone • Critical for maintaining pregnancy, and for normal development • Several additional hormones are produced Parturition: Birth Process • Prior to labor: softening of the cervix • Labor • Rhythmic uterine contractions • Positive feedback cycle of oxytocin release • Cervical dilation • Delivery of the baby • Placental release and expulsion Parturition: Birth Process Figure 26-20a Parturition: Birth Process Figure 26-20b Parturition: Birth Process Figure 26-20c Parturition: Birth Process Figure 26-20d Regulators of Parturition • Labor onset • Possible role of CRH • Stretch of cervix as a stimulus • Role of oxytocin in labor • Stretch stimulates release • Oxytocin stimulates prostaglandin release • Positive feedback (contractions stimulate more oxytocin release) Parturition Is Controlled by a Positive Feedback Loop Figure 26-21 Hormones and Mammary Gland Development • Puberty • Estrogen stimulates grow of ducts and fat deposition • Pregnancy • Estrogen, growth hormone, and cortisol stimulate further gland development • Late pregnancy • Progesterone stimulates conversion of some ducts to secretory epithelium Mammary Gland Anatomy Figure 26-22a Mammary Glands - Microscopic View Figure 26-22b Lactation: Control of Milk Secretion • Before birth • Prolactin-inhibiting hormone (PIH) blocks prolactin secretion • High sex steroid levels suppress milk production • After birth • High prolactin & low estrogen lactation • Breast milk = nutrients + immunity • Suckling • Inhibits PIH, allowing milk production • Oxytocin stimulates “let-down reflex” Hormonal Control of Milk Secretion and Release Higher brain centers Sound of child’s cry Hypothalamus PIH cell Oxytocin neuron Portal system PIH Anterior pituitary Posterior pituitary Prolactin Oxytocin Milk secretion Milk ejected Baby suckling Ascending sensory information Inhibition of prolactin cells is removed Smooth muscle contraction Mechanoreceptors in nipple Figure 26-23 Reproductive Maturation: Puberty • Maturation of reproductive organs, increased hormone levels, and onset of gamete production • Notable characteristics of puberty • Females: menarche, pubic hair, and breasts • Males: pubic hair, beard, and deep voice • Hormone changes at puberty • Increased secretion of GnRH • Increased LH/FSH levels stimulate gonads • Possible role of leptin Aging and Menopause • Causes • Ovaries cease responding to LH/FSH • Low levels of estrogen and progesterone • Cessation of egg development • Symptoms and therapies • Hot flashes and increased osteoporosis risk • Hormone replacement debate Aging and “Andropause” • Controversial (only affects 50% of men over 50 years) • Lower levels of testosterone impact some men’s lives • Many men remain reproductively active, despite low testosterone levels Summary • Sex and genetics • Hormonal regulation of reproduction • Tissues and hormones involved in spermatogenesis and oogenesis • Procreation and contraception • Fertilization and zygote development • Labor and delivery • Puberty, menopause, and andropause