Irrigation and Plant Physiology The basis of irrigation • • • • Soil Plant Evapotranspiration Plant requirements Photosynthesis • http://www.youtube.com/watch?v=IlmgFYmbAUg • Exchanges of gases and water vapors. • Sugar synthesis Photosynthesis light reaction 4 Whitehead & Gower, 2001 The stomata Http://www.youtube.com/watch?v=IlmgFYmbAUg • The Stomata 6 Stomata structure 7 Opening of the Stomata Controlling factors Lights Plant water potential Environmental conditions: atmosphere humidity Co2 concentration temperature 8 Light • Lights is electromagnetic radiation • It is the energy source for the photosynthesis. • P.A.R. – Photosynthesis Active Radiation is at the range of visible light 400700 nm 9 Why do plants are green • Which light is corresponded to wave length of 550 nm? • Peaks of absorbents 430, 680 nm Light • It is not possible to have photosynthesis without light, therefore when there is no light the stomata is closed 11 Transpiration process • Transpiration is the process of evaporation or vapor transfer from the plant to the atmosphere through the stomata • Transpiration has two stages: 1. The vaporization of liquid water in the stomata 2. Transfer of water vapors from the stomata to the atmosphere • Factors that control the rate of transpiration 1. VPD 2. Stomata conductance 3. Wind speed 12 Vapor transfer What is the driving force for vapor transfer? VPD – Vapor Pressure Deficit If the vapore pressure in the atmosphere is equal to the VP in the stomata transpiration rate is 0. Wind accelerate the rate of vapor transport Under dry and windy conditions we expect high transpiration rate. Knowing the weather forcast can help us plan irrigation. 13 Other factors • CO2 • Temperature • ABA – Abscisic Acid Abscisic acid is also produced in the roots in response to decreased soil water potential and other situations in which the plant may be under stress. ABA then translocates to the leaves, where it rapidly alters the osmotic potential of stomatal guard cells, causing them to shrink and stomata to close. 14 The water potential in the Plant • Turgor pressure • http://www.youtube.com/watch?v=1ZWP65Nbwd4 • Water potential • http://www.youtube.com/watch?v=nDZud2g1RVY Y = Ys + Yp 15 Pressure chamber סכמה גומיה תא לחץ 16 חנקן בלחץ • When pressure is applied on a leaf water rises in the petiole. • The water potential is the pressure reading when water are exiting the petiole. • http://www.youtube.co m/watch?v=fpZIdiygy iE 17 Water potential in Apple tree SWP-H SWP-L LWP-H LWP-L 18 Water potential and the photosynthesis When the water potential is decreasing the stomata close Then the photosynthesis stop Thus the water potential in the plant affect the photosynthesis