OVERVIEW OF PLANT STRUCTURE BASIC TYPES OF PLANT CELLS ▪ Parenchyma ▪ Collenchyma ▪ Sclerenchyma Parenchyma Cells ▪ The most common type of living differentiated cell. ▪ Greek parenchein, meaning “to pour in beside” ▪ Usually spherical, cubical or elongated. ▪ Parenchyma are alive at maturity with thin primary walls and typically no secondary wall. ▪ Parenchyma cells serve for photosynthesis and storage. ▪ Parenchyma tissue (a). Starch storage parenchyma cells occur in leaves, stems and roots (b). Parenchyma can also serve as water storage cells in roots and stems (c). Chlorenchyma are chloroplast-containing parenchyma cells that are primarily found in leaves • Composed of living parenchyma cells with thin primary cell walls • Functions include photosynthesis, storage, and secretion Collenchyma Cells ▪ ▪ ▪ ▪ Greek kolla, meaning “glue” Alive at maturity and lacking secondary walls Collenchyma cells can attain a variety of shapes and have thicker primary walls than parenchyma cells. Parenchyma and collenchyma cells must be filled with water to become turgid enough for support. Sclerenchyma Cells Collenchyma cells provide flexible support. Collenchyma cells have cell walls that are thickened in some places with additional cellulose, enabling them to provide more support than parenchyma while still allowing living stems to be flexible. ▪ Provide rigid support ▪ Greek scleros, meaning “hard” ▪ Sclerenchyma cells are usually dead at maturity ▪ Have secondary walls, often hardened with lignin (a rigid molecule that strengthens & stiffens cell walls in vascular plants) ▪ They can provide support without being filled with water. Two Main Types of Sclerenchyma 1. Fibers -elongated cells with thick secondary walls reinforced with lignin, making them flexible as well as strong. -enable stems to move in the wind without snapping 2. Sclereids -vary more in shape than fibers but are often cubical or spherical. -make structures such as nutshells and fruit pits typically rock-hard and inflexible. -the gritty sandlike pieces in fleshy part of a pear are also sclereids, commonly called stone cells. • Collenchyma tissue • Composed of collenchyma cells with unevenly thickened primary cell walls • Provides flexible structural support Sclerenchyma cells provide rigid support. (a) This stem has long, tapering sclerenchyma cells called fibers, which provide sturdy support for a plant. (b) Thick, hard cell walls of sclereid cells, also known as stone cells provide the structural support in fruit pits and nutshells, and also give a rough texture to the flesh of a pear. KEY TERMS • PARENCHYMA CELL • • COLLENCHYMA CELL • • • • Sclerenchyma tissue Composed of sclerenchyma cells with both primary and secondary cell walls Sclerenchyma cells are often dead at maturity, but provide structural support Living plant cell with moderately but unevenly thickened primary walls SCLERENCHYMA CELL • • Relatively unspecialized plant cell; thin walled, may contain chlorophyll, loosely packed Plant cell with extremely thick walls; provides strength and support to plant body Tissues of Vascular Plants ▪ A tissue- a group of cells that performs a function. ▪ May be simple (one cell type) or complex (multiple cell types). ▪ Simple tissue- parenchyma, collenchyma and sclerenchyma cells. ▪ Complex tissue- mixture of parenchyma, sclerenchyma and water-conducting cells THE THREE TISSUE SYSTEMS 1. The dermal tissue system 2. The vascular tissue system 3. The ground tissue system The Dermal Tissue Systems ▪ Forms the plant’s outer protective covering ▪ Greek derma, meaning “skin” ▪ A single layer of epidermis is replaced during second year of growth by periderm, (mainly composed of nonliving cells). ▪ Dermal cells can be modified into hairlike trichomes. ▪ The aboveground dermal tissue often forms a waxy protective layer called a cuticle. KEY TERMS • EPIDERMIS • Outermost tissue layer, usually one cell thick • Covers the primary plant body (leaves, young stems and roots) • Epidermis covering aerial parts secretes a wax layer (cuticle) that reduces water loss • Gas is exchanged between interior of shoot system and surrounding atmosphere through stomata • PERIDERM • Outermost layer of cells covering a woody stem or root (the outer bark that replaces epidermis when it is destroyed during secondary growth) KEY TERMS • • • The Vascular Tissue Systems ▪ Conducts water, minerals and food ▪ Two complex tissues of vascular system: 1. xylem- for transporting water and minerals 2. phloem- for transporting food XYLEM A complex vascular tissue that conducts water and dissolved minerals throughout the plant body Actual conducting cells of xylem are tracheids and vessel elements Pits are gaps in the secondary cell wall. Tracheids These are stacked with pits allowing for movement of water and minerals from one tracheid to the other. Vessel elements. A large, water-conducting cell in the xylem of most flowering plants; transports water and minerals more rapidly than tracheid cells. • PHLOEM • • A complex vascular tissue that conducts food (carbohydrate) throughout the plant body Conducting cells of phloem are sieve-tube elements assisted by companion cells Sieve-tube Members. The phloem of flowering plants consists of cells called sieve-tube members (sieve-tube elements). Sieve-tubes. They conduct organic nutrients from the leaves to other parts of the plants. The Ground Tissue System An Overview of Vascular Plant Organs ▪ Also called the fundamental tissue system. ▪ Simple and complex organ tissues form structures known as organs. ▪ Ground tissue fill up space not occupied by vascular or dermal tissue. ▪ An organ consists of several types of tissues adapted as a group to perform particular functions. ▪ Carries out photosynthesis and stores nutrients. ▪ Vascular plants have three types of organs: ▪ Cortex- ground tissue forms between the dermal and vascular tissues. ▪ (1). Stems (2). Leaves (3). Roots ▪ Pith- it also appears to the inside of the vascular tissues, where it is called pith. 3 Tissue Systems in Plant Body STEMS ▪ Position leaves for maximum photosynthesis ▪ A stem is any part of a plant that supports leaves or reproductive structures. ▪ Stems can vary greatly in size, such as slender stalk supporting a small flower or a huge tree trunk dozens of feet in diameter and hundreds of feet high. 2. Transpiration -the loss of water through the pores in leaves, which pulls water and mineral nutrients up from the roots to the leaves. Functions of Stems 1. Stems supports leaves and reproductive structures, 2. Transport water and nutrients and 3. Protect the plant. LEAVES ▪ The earliest land plants were leafless photosynthetic stem systems. ▪ Eventually their flattened stems grew close together, evolving into continuous structures known as leaves. ▪ A leaf is the main photosynthetic organ of modern plants. Cactus stem and leaf adaptations. The stem and leaves of cacti are well adapted for hot, dry climate of the desert. The thick stems serve as water storage, while the small surface area of the thin spines (leaves) loses less water to evaporation. Functions of Leaves 1. Photosynthesis -the process by which the plants use solar energy to make their own food, transforming carbon dioxide and water into sugars that store chemical energy. ROOTS ▪ A root has two main functions: 1. Anchoring the plant in the soil 2. Absorbing water and minerals. Root hairs. Root hairs are found within a centimeter of the tips of roots just above the region of cell elongation. They specialized to take up both water and ions needed by the plant. (a) Here are root hairs on a radish seedling. (b) This micrograph provides a close-up view of root hairs. An Overview of Plant Growth and Development ▪ A typical plant has a: 1. Root system (usually below ground) and a 2. Shoot system (usually above ground) consisting of all stems, leaves and reproductive structures. SHOOT ▪ A shoot is any individual stem and its leaves, as well as any reproductive structures that extend from the stem such as flowers. ▪ A shoot that has leaves but no reproductive structures is called a vegetative shoot. EMBRYOS SEED COAT ▪ Give rise to stems, leaves and roots of adult seed plants. ▪ A seed plant begins as a fertilized egg or zygote that grows into an embryo within a seed. ▪ The typical seed plant embryo includes the following embryonic “organs” that develop into the roots and shoots. PARTS OF A SEED (EMBRYO) ▪ A seed is enclosed with a covering called the seed coat. COTYLEDONS ▪ One or more “seed leaves” called cotyledons, which are usually the largest, most visible parts of the embryo. ▪ Monocot- one cotyledon (corn, rice, palms) ▪ Dicot- two cotyledon (beans, peas, mango) 1. Seed coats 4. Plumule 2. Cotyledons 5. Hypocotyl ▪ Latin radix, meaning “root” 3. Radicle 6. Endosperm ▪ An embryonic “root” which is always prominent. ▪ It is the first root that grows from the seed. RADICLE PLUMULE ▪ Latin plumula, meaning “soft leather” ▪ An embryonic “”shoot” which is usually scarcely developed. ▪ It gives rise to the epicotyl, a portion of the embryonic “stem” located over the cotyledon. HYPOCOTYL ▪ A portion of the embryonic “stem” which is located under the cotyledon and above the radicle and may be either short or long. ▪ It is the first stem that develops from the seed. ENDOSPERM ▪ ▪ The embryo is surrounded by a storage tissue, called endosperm in flowering plants, that breaks down to provide nourishment to the developing embryo. Root Tip Sometimes endosperm also supplies nutrition to the sprouting seed. KEY TERMS PRIMARY GROWTH • An increase in stem and root length due to the activity of apical meristems at the tips of roots and at the buds of stems APICAL MERISTEM • An area of cell division at the tip of a stem or root in a plant; produces primary tissues BUD • A dormant embryonic shoot that eventually develops into an apical meristem. Apical Meristems ▪ Primary growth (growth in length) comes from apical meristems at tips of roots and shoots. ▪ Leaves originate on shoot apical meristems as leaf primordia. ▪ Each mature leaf is attached to a stem by petiole at a point called a node. Stem Tip Stem Tip Development SECONDARY GROWTH • An increase in a plant’s stem and root girth due to the activity of lateral meristems (the vascular cambium and cork cambium) Secondary growth is localized, typically as long cylinders of active growth throughout the lengths of older stems and roots LATERAL MERISTEM • An area of cell division on the side of a vascular plant; the two lateral meristems (vascular cambium and cork cambium) give rise to secondary tissues Lateral Meristems and Secondary Growth