The Ground Tissue The ground tissue makes up a large portion of the primary body of the plant. It consists of cortex and pith of the stem and root as well as the mesophyll of the leaves. The cortex lies next to the epidermis while the pith occupies the center of many roots and stems. Three kinds of cells are present in the ground tissue: the parenchyma, sclerenchyma, and collenchyma. Sclerenchyma The chief strengthening tissues of plants are called sclerenchyma. Sclerenchyma consists of elongated cell with very thick walls. There are two kinds of these; the sclereids and the fibers. In some mature sclerenchyma cells, all protoplasm may be replaced by the encroachment of the thick cell walls, which impart mechanical strength to the tissues. It is present in some stem, roots and in some leaves. Parenchyma Parenchyma cells are the most abundant type of cells in the plant. They have thin walls and retain their protoplasm throughout their lifetime. They are the least differentiated among plant cells so they are able to redifferentiate into other type of cells. They also sere as storage cells and may have starch granules contained inside plastids. There are also parenchyma cells which perform specialized functions. Photosynthesis takes place specifically in parenchyma cells which contain chloroplastids. They are called chlorenchyma cells and they make up the ground tissue of the leaves and other photosynthetic parts of the plant. Photosynthetic Tissues The photosynthetic tissues are abundant in the leaves and young stems of the plant. These are the chlorenchyma cells which contain the chloroplasts. The photosynthetic tissues are found in the mesophyll layer of the leaves and young stems. Collenchyma Collenchyma cells are elongated cells with unevenly thickened primary cell walls. Thickenings are usually found at the corners or on certain areas of the cell wall like parenchyma, collenchyma also retain their protoplasm throughout their life but unlike parenchyma, collenchyma cells hae no intercellular spaces at the corners of the cells ecause such spaces are filled with cell wall materials. Collenchyma can perform the various metabolic activities undertaken by parenchyma cells. The main function of collenchyma is to provide mechanical strength to herbaceous and rapidly growing parts of the plants. Mature complex plants are mainly supported by supporting tissues located in roots and stems. Roots anchor the plant to the ground. The roots branch even more profusely than the stem and frequently, the total root surface is greater than the stem with all its branches. The extensively branching stem system of the typical land plant has two primary functions: it supports the leaves in a position which assures balance and to maximize photosynthesis. Permanent Tissues The Vascular or Conducting Tissues Conducting tissues or vascular tissues are also present in the plant. They are concerned with the transport of material and are the equivalent of the circulatory system of the animals. Vascular tissues are made up of the xylem and phloem. Xylem conducts water and mineral salts from the roots to the leaves while the phloem conducts food materials from the leaves to the other parts of the plants. Sieve tubes: The main conducting cells of the phloem Vascular tissues are composed of the sieve tubes and ducts. Sieve tubes are tubular units found in phloem of stems, roots and leaves. Each sieve tube represents a column of elongated cylindrical cells placed end to end. At maturity the end walls of these become perforated, forming sieve plates which give protoplasmic continuity between the successively placed cells in the column. Tracheid and Vessel elements Tracheid and vessel elements are tubular units localized in the xylem regions of roots, stems and leaves. Theyare the two kinds of xylary elements of the xylem. At first, it resembles that of the sieve tube, but later the side walls become much thicker, and the end walls are reabsorbed completely. They form continuous channels from roots to stems and into the leaves. Secondary Tissues and Secondary Growth Vascular tissues which develop from the procambium are referred to as primary phloem and primary xylem. In the dicot, a layer of meristematic tissue becomes the vascular cambium and eventually gives rise to secondary vascular tissues and the cork tissue. Organs and Organ Systems An organ is a group of tissues that perform a certain function. The science of the arrangement of tissues to form organs is called Organology. The chief functional cells of an organ are its parenchyma; the supporting tissues, its stroma. Organ Systems An integrated group of organs form systems. Each system is concerned with one of the basic functions common to all animals.