I. Chapter 5: Biodiversity, Species Interactions, and Population Control a. 5-1: How Do Species Interact? i. Five basic interactions among species: 1. Interspecific Competition – most common interaction, occurs when members of two or more species interact to use the same limited resources such as food, water, light, and space a. Most interspecific competition involves the ability of one species to become more efficient than another species in obtaining the resources it needs. i. When two species compete with one another for the same resource their niche overlaps. The greater the overlap, the more intense their competition for key resources is. 1. Humans compete with many other species for space, food, and other resources b. Resource Partitioning – populations of some species develop adaptations that allow them to reduce or avoid competition with other species for resources. i. Competing for similar scarce resources evolve specialized traits that allow them to share resources by using parts of them, using them at different times, or using them in different ways. 2. Predation – a member of one species (the predator) feeds directly on all or part of a living organism (the prey) as part of a food web. a. Predator-Prey relationship b. Predator’s have a variety of methods that help them to capture prey. i. Pursuit and Ambush ii. Camouflage iii. Chemical Warfare iv. Prey Species – adaptations include protective shells, thick bark, thorns, spines v. Mimicry – gain protection by looking and acting like other, more dangerous species, a protective device vi. Echolocation – navigate and locate their prey c. Population level – predation plays a role in evolution by natural selection i. Predator and prey can exert intense natural selection pressures on one another d. Coevolution – two different species interact in such a way over a long period of time, changes in the gene pool of one species can lead to changes in the gene pool of the other 3. Parasitism – Occurs when one species (the parasite) feeds on another organisms (the host), usually by living on or inside the host a. Host point of view, parasites are harmful b. Population perspective, parasites can promote biodiversity by helping to keep the populations of their host in check 4. Mutualism – Two species behave in ways that benefit both by providing each with food, shelter, or some other resource a. In reality, the species in a mutualistic interaction benefit one another unintentionally and are in it for themselves 5. Commensalism – an interaction that benefits one species but has little, if any, beneficial or harmful effect on the other. b. 5-2: How do Communities and Ecosystems Respond to Changing Environmental Conditions? i. Ecological Succession – types and numbers of species in biological communities and ecosystems change in response to changing environmental conditions such as fires, volcanic eruptions, climate change, and the clearing of forests to plant crops. 1. Primary Ecological Succession – involves gradual establishment of communities of different species in lifeless areas where there is no soil in a terrestrial ecosystem or no bottom sediment in an aquatic ecosystem. 2. Secondary Ecological Succession – which a series of communities or ecosystems with different species develop in places containing soil or bottom sediment. 3. Ecological Succession is an important ecosystem service that tends to increase the biodiversity of communities and ecosystems by increasing species richness and interactions among species. a. Both primary and secondary ecological succession are example of natural ecological restoration i. Three factors that affect how and at what rate succession occurs: 1. Facilitation – which one set of species makes an area suitable for species with different niche requirements, and often less suitable for itself. 2. Inhibition – which some species hinder the establishment and growth of other species 3. Tolerance – which plants in the late stages of succession succeed because they are not in direct competition with other plants for key resources. b. Traditional view – ecological succession proceeds in an orderly sequence along an expected path until a certain stable type of climax community occupies an area c. Ecological succession reflects the ongoing struggle by different species for enough light, water, nutrients, food, space, and other key resources. ii. Stability or capacity to withstand external stress and disturbance, is maintained only by constant change in response to changing environmental conditions. 1. Two aspects of stability or sustainability in living systems: a. Inertia (Persistence) – the ability of a living system such as a grassland or a forest to survive moderate disturbances b. Resilience – the ability of a living terrestrials system to be restored through secondary ecological succession after a more severe disturbance i. Evidence suggest that some ecosystems have one of these properties but not the other c. 5-3: What Limits the Growth of Populations? i. Population – a group of interbreeding individuals of the same species. 1. Three ways in which the members of a population are typically distributed or dispersed. ii. iii. iv. v. vi. vii. a. Clumps (most populations live like this) i. The resources a species needs vary greatly in availability from place to place ii. Individuals moving in groups have a better chance of encountering patches or clumps of resources iii. Living in groups provides some protection from predators iv. Living in packs gives some predator species a better change of getting a meal 2. Births, deaths, immigration, and emigration govern changes in population size a. Population Change = (Births + Immigration) – (Deaths + Emigration) Age Structure – the distribution of individuals among various age groups 1. Usually described in terms of organisms not mature enough to reproduce (the prereproductive stage), those capable or reproduction (the reproduction stage), and those too old to reproduce (the postreproductive stage). Range of Tolerance – Variations in its physical and chemical environment Limiting Factors – more important than other factors in regular population growth 1. Limiting Factor Principal – too much or too little of any physical or chemical factor can limit or prevent the growth of a population, even if all other factors are at or near the optimal range or tolerance a. On land, precipitation often is the limiting factor b. Aquatic life zones – includes temperature, sunlight, nutrient availability, acidity, and the low levels of oxygen gas in the water i. Salinity – the amounts of various inorganic minerals or salts dissolved in a given volume of water. Population Density – The number of individuals in a population found within a defined area or volume Different reproductive patterns help to ensure the long-term survival of species 1. No population of a species can grow indefinitely because of limitations on resources and competition with populations of other species for those resources a. There are always limits to population growth in nature Environmental Resistance – the combination of all factors that act to limit the growth of a population. 1. Carrying Capacity – the maximum population of a given species that a particular habitat can sustain indefinitely a. A population with few, in any, limitations on its resource supplies can grow exponentially at a fixed rate or percentage per year. b. Plotting the number of individuals against time yields a J-shapes growth curve c. Exponential Growth – occurs when a population has essentially unlimited resources to support its growth i. Eventually converted to logistic growth, which the growth rate decreases as the population becomes larger and faces environmental resistance d. Population size typically stabilizes on or near the carrying capacity which results in a sigmond (S-shaped) population growth curve e. The carrying capacity can vary over time due to changing conditions viii. Population Crash (Dieback)– when a population overshoots or exceeds the carrying capacity of their environment. ix. The carrying capacity of any given area is not fixed 1. It can rise or decline seasonally and from year to year 2. Humans are not exempt from population crashes a. Bubonic Plague b. Global Epidemic of AIDS i. So far, technological, social, and other cultural changes have expanded the earth’s carrying capacity for the human species.