Anatomy & Physiology 34B Lecture Chapter 13 & 15 – PNS Sensory Systems I. Overview A. General Properties of Sensory Systems B. Somatic Senses C. Special Senses – Olfactory & Gustatory II. General Properties of Sensory Systems A. _______ organs are extensions of the nervous system that respond to changes in the environment & transmit nerve impulses to the CNS B. In order to ___________ a sensation, the following are necessary: 1. A _____________ (chemical, mechanical, temperature, or light) to initiate a nervous system response 2. A __________ (sensory neuron dendrites or specialized epithelial cell) is a transducer that converts the stimulus to a nerve impulse 3. ______________ of the nerve impulse from the receptor to the brain, via sensory (ascending) nerve tracts in the spinal cord 4. _______________ of the perception in the brain’s cerebral cortex, after passing through the medulla, pons, and thalamus C. ____________ may be general or special senses 1. _______________ (somatic) senses a. Have _____________ throughout the skin, muscles, tendons, joint capsules, and viscera. b. Somatic and visceral __________ _________ transmit impulses from the receptors to the CNS 2. ______________ senses a. Are limited to the head and innervated by ____________ nerves b. Include vision, hearing, equilibrium, taste, and ___________ D. Sense _______________ may be classified several ways 1. By the way they are _______________ a. ________receptors respond to chemicals; found in the tongue, nose, and blood vessels b. __________receptors respond to heat and cold; in skin c. _______ceptors respond to pain and itch; found throughout the body, except in the brain d. __________receptors respond to physical deformation of the plasma membrane caused by touch, pressure, stretch, tension, or vibration; found in skin, viscera, and joints e. __________receptors in the eyes respond to light 2. By the ___________________ of receptors in the body a. ______________ (somatic) senses 1) Have receptors throughout the _______, muscles, tendons, joint capsules, and viscera. 2) Include mechanoreceptors, thermoreceptors, chemoreceptors, and ______________ b. ______________ senses 1) Are limited to the ______ and innervated by cranial nerves 2) Include _________, hearing, equilibrium, taste, and smell 3. By ____________ of the stimuli a. _________ceptors detect stimuli in internal organs and produce feelings of visceral pain, nausea, stretch, and pressure b. ____________ceptors sense the position and movements of body parts; found in muscles, tendons, and joint capsules c. _________ceptors sense external stimuli; include receptors for vision, hearing, taste, smell, touch, and cutaneous pain 4. By the __________ of their electrical impulse firing frequencies a. __________ receptors generate a burst of impulses when first stimulated, adapt quickly, and stop even if stimuli continues (e.g.: touch, pressure, and smell receptors) b. ___________ receptors adapt slowly and generate impulses continually (e.g.: nociceptors, proprioceptors) III. General Senses A. ____________ sense receptors include those with unencapsulated and encapsulated nerve endings 1. __________________ nerve endings are sensory dendrites that are not wrapped in CT a. __________ nerve endings include thermoreceptors and nociceptors; numerous in epithelia and CT b. Tactile (_________) discs are tonic receptors for light touch; found in the epidermis stratum basale c. ___________ receptors monitor movements of hairs; found at the base of hair follicles 2. _______________ nerve endings are dendrites wrapped in glial cells or CT. These include a. Tactile (_____________) corpuscles are phasic receptors for light touch and pressure; found in the dermal papillae b. ___________ end bulbs are similar to tactile corpuscles, but occur in mucous membranes rather than in the skin c. ______________ (lamellated) corpuscles are phasic receptors for deep pressure, stretch, tickle, and vibration; found deep in the dermis and pancreas d. _______________ corpuscles are tonic receptors for heavy touch, pressure, stretch, and joint movements; found in the dermis, hypodermis, ligaments, tendons, and joint capsules E. Each sensory __________ has a receptive field 1. A sensory neuron is activated by stimuli that fall within the neuron’s ____________ _________ 2. Example: a ______-________ discrimination test a. In less sensitive skin areas, _______ primary neurons converge on _____ secondary neuron (a large receptive field), thus two pins 20 mm apart are perceived as _____ pinprick b. More sensitive skin areas often have smaller receptive areas, with a ______-to-_____ relationship between primary and secondary neurons, thus two pins 2 mm apart are perceived as ____________ F. The CNS _______________ sensory information 1. Sensory impulses (except smell) travel to the spinal cord (or brain stem), up projection tracts to the ____________, and are relayed to the cererbral cortex 2. The perceptual _____________ is the level of stimulus intensity necessary for us to be aware of a sensation a. To keep from being overwhelmed by sensory input, the CNS can selectively “turn _____” some stimuli b. _______________ modulation often occurs in secondary or tertiary neurons in the sensory pathway G. The CNS must distinguish the 4 _____________ of a stimulus: modality, location, intensity, and duration 1. Sensory _____________ is indicated by which types of sensory neurons are activated; the brain associates a signal from a particular type of receptor with a specific sensation 2. Stimulus ______________ is coded by different types of sensory neurons sending their impulses through ascending tracts to specific regions of the cerebral cortex 3. Stimulus ____________ is coded by the number of receptors that are activated and by the frequency of their action potentials 4. ______________ of a stimulus is coded by the duration of action potentials in a sensory neuron. Neurons may be either: a. __________ receptors adapt slowly and generate impulses continually (e.g.: nociceptors, proprioceptors) b. _____________ receptors generate a burst of impulses when first stimulated, adapt quickly, and stop even if stimuli continues (e.g.: touch, pressure, and smell receptors) III. ___________ Senses include touch, proprioception, temperature, and nocioception (pain and itch) A. Receptors for somatic senses are found in the _______ and viscera 1. Receptor activation triggers action potentials in associated ____________ sensory neurons, which travel to the spinal cord and synapse with 2. _____________ sensory neurons (interneurons), in ascending tracts, which cross over to the thalamus on the opposite side, where they synapse with 3. ______________ sensory neurons, which project to a particular region of the sematosensory cortex B. Pain that originates in one area, but is felt in another area is called ______________ pain, and occurs because multiple primary sensory neurons converge onto a single ascending tract C. _____________ limb pain in which an amputee feels pain in the amputated limb also occurs because of nerve convergence in the spinal cord IV. Special Senses A. _____________ Sense (sense of smell) 1. Olfactory receptors are dendritic endings of the _____________ nerve (CN I) that respond to chemical stimuli 2. Odor impulses are transmitted it to the olfactory __________ via the following pathway: a. b. Olfactory _____________ cells in nasal epithelium receive the stimulus The impulse is transmitted via olfactory nerves (CN ____) that extend through the cribriform plate of the ethmoid bone to the c. Olfactory _______ on both sides of the crista galli, beneath the frontal lobes. Neurons here convey the impulse to neurons of d. e. Olfactory __________, which transmits the impulse to the Olfactory ___________ within the temporal and frontal lobes, where it is interpreted as odor B. __________________ Sense (sense of taste) 1. Gustatory ____________ are specialized epithelial cells, clustered in taste buds, that respond to chemical stimuli 2. Taste _________ are lemon shaped structures composed of gustatory cells surrounded by supporting cells and basal cells 3. Each gustatory cell has a ________________ taste hairs that extend through a taste pore on the taste bud surface 4. There are 3 major types of tongue ________________: 5. 6. a. ___________ (circumvallate) papillae - largest but least numerous, arranged in an inverted V on the back of tongue b. ______________ papillae - knoblike papilla on the tip & sides of tongue; both fungiform & vallate papillae contain taste buds c. _______________ papillae - short, thick, threadlike; on the anterior 2/3 of tongue; contains no taste buds Five basic ____________ are sensed: a. ___________ b. ___________ c. ___________ d. ___________ e. ___________ – a meaty taste produced by amino acids Taste impulses are transmitted via the __________pharyngeal (CN IX) and __________ (CN VII) nerves to the gustatory cortex in the parietal lobes for interpretation