D’YOUVILLE COLLEGE BIOLOGY 108/508 -HUMAN ANATOMY & PHYSIOLOGY II LECTURE # 1 SENSORY ORGANS I Chapter 13 1. Introduction: • homeostasis: maintenance of steady state; external & internal environments monitored by sensory receptors • receptors: environmental inputs to the afferent limb of the reflex arc • single-celled (nerve ending) or complex organs containing receptors + structures designed to facilitate reception of specific type of environmental condition • receptors act as transducers to convert one form of energy (stimulus modality) to another, e.g. light energy to electrical energy of nerve impulses a. Perceptions (sensations): interpretation of inputs of electrical impulses related to the destination in the brain; brain region produces a sensation (smell, sight, touch, etc.) & “projects” perception to sensory field (accounts for “phantom limb” phenomenon) b. Sensory adaptation: reduced response to a sustained stimulus 2. Types of Sensory Organs: (table 13 – 1) According to Stimulus Modality: a. chemoreceptors: chemicals dissolved in body fluids, e.g. olfactory and gustatory organs, osmoreceptors, blood gas receptors, pH receptors b. mechanoreceptors: physical distortion such as stretching, compression, etc., e.g. baroreceptors, spindle organs, touch receptors of skin (Meissner’s corpuscles), pressure receptors of skin (Pacinian corpuscles), organs of hearing and equilibrium c. thermoreceptors: temperature changes, e.g. cold receptors respond to temperatures of 10º - 20º C.; heat receptors are sensitive to temperatures above body temperature (up to 45 º C.) d. photoreceptors: light energy, e.g. rods and cones of the retina. • nociceptors: painful stimuli that may be triggered by extremes of any of the other modalities or by noxious agents (e.g. prostaglandins) released at sites of injury • do not normally exhibit sensory adaptation • neuropeptides (e.g. enkephalins or endorphins) may inhibit or diminish pain; release may be triggered by serotonin & may provide the basis for placebo effects & acupuncture. • visceral pain (associated with internal organs) is experienced at sites removed from the site of the pain stimulus, a phenomenon known as referred pain. Alternative Classification: Bio 108/508 lec. 1 - p. 2 a. exteroceptors: associated with the skin, e.g. Pacinian and Meissner’s corpuscles, temperature receptors b. proprioceptors: associated with muscles, tendons and joints, e.g. spindle organs c. visceroceptors: associated with internal organs, e.g. baroreceptors, osmoreceptors d. special sensory organs: more complex structures; include the senses of taste (gustatory), smell (olfactory), sight (visual), hearing (auditory) and balance (equilibrium) Special Senses Chapter 15 1. Olfactory Organs: smell receptors are sensory hair cells in nasal epithelium of roof of nasal cavity (fig • variety of fragrances recognized including: - floral, musky, pungent, putrid, peppermint, ether, camphor (formerly construed to be the primary odorants); current consensus recognizes many more, based upon identified anosmias and genes identified that code for olfactory receptors • served by cranial nerve I (olfactory nerve): hair cells synapse within olfactory bulb (table 13 – 2) with afferent neurons which transmit impulses, via areas of limbic system, to regions of frontal and temporal lobes 2. Gustatory Organs: taste buds are located mainly on sides of foliate, fungiform and circumvallate papillae on tongue’s superior surface (fig. 15 – 23) • taste buds are barrel-shaped clusters of cells, including hair cells & supporting cells (fig 15 – 23c) • five (formerly four) primary taste qualities are identified: a. sweet: respond to sugars, alcohols, & certain amino acids b. sour: sensitive to acids c. salty: respond to metal ions, especially sodium d. bitter: detect alkaloids (e.g., quinine & caffeine) and aspirin e. umami: responsible ‘beef’ and ‘aged cheese’ tastes • mainly cranial nerves VII, IX, & X (facial, glossopharyngeal, & vagus) (fig. 15 – 24)